MUSSELpdb | publication Freeman et al., 2025

	Home \| About \| Database \| Mussel of the Month \| Publications & Presentations


MUSSELpdb
	Introduction Browse d'basing


Taxonomy
	all genera Unionidae Margaritiferidae Hyriidae Etheriidae Mycetopodidae Iridinidae Species & Genus Richness by Taxonomy


Geography
	Summary North America Central America South America Afrotropics North Eurasia East Asia Australasia Species Richness & Endemism by Geography

Page last updated
Tue 03 Jun 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Freeman et al. (2025), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

Search the database pages:

Publication information

Freeman, E.-J.G., W.G. Bauer, R.H. Easy, R.Q. Céspedes, S.A. Valverde & D.T. Stewart. 2025. Phylogenetic analysis of the freshwater mussel Nephronaias tempisquensis (Bivalvia: Unioinidae: Popenaiadini) from Costa Rica. Malacologia 67(1/2): 241-255.

Click here for online access.

Cladogram Figures

(click the figure number to see the cladogram)

fig. 2
fig. 3
fig. 4
fig. 5

Taxonomic Opinions

Amblema plicata (Say, 1817)

Cyclonaias pustulosa (I. Lea, 1831)

Cyrtonaias explicata (Morelet, 1849)

Cyrtonaias tampicoensis (I. Lea, 1857) [sic]

Fusconaia flava (Rafinesque, 1820)

Glebula rotundata (Lamarck, 1819)

Nephronaias tempisquensis Pilsbry, 1920

Obliquaria reflexa Rafinesque, 1820

Pachynaias spheniopsis (Morelet, 1849)

Plectomerus dombeyanus (Valenciennes, 1827)

Pleurobema sintoxia (Rafinesque, 1820)

Popenaias popeii (I. Lea, 1857)

Psoronaias semigranosa (Busch in Philippi, 1843)

Psorula guatemalensis (Simpson, 1900)

Psorula percompressus (Martens, 1887)

Psorula profunda (Simpson, 1914)

Psorula rudis (Simpson, 1900)

Psorula usumasintae (Crosse & Fischer, 1893)

Quadrula quadrula (Rafinesque, 1820)

Sphenonaias microdon (Martens, 1887)

Toxolasma lividum Rafinesque, 1831

Cladogram Figure 2

           +----- Pachynaias_spheniopsis   
  +--------+                               
  |        |  +-- Sphenonaias_microdon     
  |        +--+                            
--+           +-- Cyrtonaias_explicata     
  |                                        
  |  +----------- Popenaias_popeii         
  |  |                                     
  +--+  +-------- Nephronaias_tempisquensis
     |  |                                  
     +--+  +----- Psoronaias_guatemalensis 
        |  |                               
        +--+  +-- Psoronaias_usumasintae   
           |  |                            
           |  +-- Psoronaias_semigranosa   
           +--+                            
              +-- Psoronaias_profunda      
              |                            
              +-- Psoronaias_percompressa

Cladogram Figure 3

              +-- Nephronaias_tempisquensis
     +--------+                            
     |        +-- Popenaias_popeii         
  +--+                                     
  |  |  +-------- Toxolasma_lividum        
  |  +--+                                  
  |     |  +----- Obliquaria_reflexa       
  |     +--+                               
  |        |  +-- Glebula_rotundata        
--+        +--+                            
  |           +-- Cyrtonaias_tampicoensis  
  |                                        
  |           +-- Amblema_plicata          
  |     +-----+                            
  |     |     +-- Plectomerus_dombeyanus   
  |     |                                  
  +-----+     +-- Pleurobema_sintoxia      
        |  +--+                            
        |  |  +-- Fusconaia_flava          
        +--+                               
           |  +-- Quadrula_quadrula        
           +--+                            
              +-- Pustulosa_pustulosa

Cladogram Figure 4

        +----- Obliquaria_reflexa       
        |                               
        +----- None                     
        |                               
  +-----+  +-- Glebula_rotundata        
  |     +--+                            
  |     |  +-- Cyrtonaias_tampicoensis  
  |     |                               
  |     |  +-- Nephronaias_tempisquensis
  |     +--+                            
--+        +-- Popenaias_popeii         
  |                                     
  |        +-- Amblema_plicata          
  |  +-----+                            
  |  |     +-- Plectomerus_dombeyanus   
  |  |                                  
  +--+     +-- Pleurobema_sintoxia      
     |  +--+                            
     |  |  +-- Fusconaia_flava          
     +--+                               
        |  +-- Quadrula_quadrula        
        +--+                            
           +-- Pustulosa_pustulosa

Cladogram Figure 5

           +-- Nephronaias_tempisquensis
     +-----+                            
     |     +-- Popenaias_popeii         
  +--+                                  
  |  |  +----- Obliquaria_reflexa       
  |  +--+                               
  |     |  +-- Glebula_rotundata        
  |     +--+                            
--+        +-- Cyrtonaias_tampicoensis  
  |                                     
  |        +-- Amblema_plicata          
  |  +-----+                            
  |  |     +-- Plectomerus_dombeyanus   
  |  |                                  
  +--+     +-- Pleurobema_sintoxia      
     |  +--+                            
     |  |  +-- Fusconaia_flava          
     +--+                               
        |  +-- Quadrula_quadrula        
        +--+                            
           +-- Pustulosa_pustulosa

Page updated: 11:46:58 Tue 03 Jun 2025

"Making the world a better place, one mollusk at a time."
	The MUSSEL Project — Home Page http://www.mussel-project.net/. Site developed and maintained by Dan Graf & Kevin Cummings. Hosted by The University of Wisconsin-Stevens Point. Funded by The National Science Foundation.