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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win! 
This page documents data for Zhadin (1938), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work. The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=publ&id=24 
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 Publication information 
Zhadin, V.I. 1938. Sem. Unionidae. Faune URSS, Mollusques 4(1). 169 pp. 
Attributed Species Names
  Mytilus cellensis Menke, 1828
    ‘Gmelin’  of Authors.
     Menke, 1828, Syn. Method. Moll.: 63.
     Menke, 1830, Syn. Method. Moll.: 106.
     Küster, 1839, Conchyl. Cab. 9(1): 16.
     Mörch, 1853, Cat. Conch. Yoldi 2: 48.
     Clessin, 1877, Deutsche Excurs. Moll. Fauna: 438.
     Drouët, 1881, Unionidae Russie: 22.
     Bourguignat, 1882, Natural. Sicil. 2 (1): 20 [1].
     Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 211.
     Zhadin, 1938, Faune USSR Mollusques 4(1): 119.
     Modell, 1945, Jena. Z. Med. Naturw. 78: 81.
    (error for Mytilus zellensis Gmelin, 1791)
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  Anodonta tehernaica Westerlund, 1890
    ‘Bourguignat’  of Authors.
     Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 226.
     Zhadin, 1938, Faune USSR Mollusques 4(1): 121.
    (error for Anodonta tchernaica Bourguignat, 1881)
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  Unio (Nodularia) continentalis Zhadin, 1938
     ‘Haas’ Zhadin, 1938, Faune USSR Mollusques 4(1): 100, fig. 31.
     ‘Haas’ Zhadin, 1952, Moll. SSSR (46): 299, fig. 263.
    (misidentified reference to Nodularia continentalis Haas, 1910)
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  Margaritana sachalinensis Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 114-115, fig. 37.
     Zhadin, 1965, Moll. USSR: 238, fig. 252.
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  Unio lindholmi Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 97, text fig. 29.
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  Unio pictorum var. okensis Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 77.
    (intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; new name for Unio pictorum var. okae Kobelt & Haas, 1910)
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  Unio pictorum var. pygmaeus Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 78, text fig. 17.
    (intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; homonym of Unio pygmaeus Lea, 1852)
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  Unio pictorum var. defectivus Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 79.
    (intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)
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  Anodonta (Haasiella) arcaeformis Zhadin, 1938
     ‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 147, fig. 56.
     Zhadin, 1965, Moll. USSR: 260, fig. 279.
    (misidentified reference to Anodon arcaeformis Heude, 1877)
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  Anodonta (Haasiella) euscaphus Zhadin, 1938
     ‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 149, fig. 58.
     Zhadin, 1965, Moll. USSR: 260, fig. 280.
    (misidentified reference to Anodon euscaphys Heude, 1879; error for Anodon euscaphys Heude, 1879)
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  Anodonta piscinalis var. volgensis Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 126, fig. 43.
     Zhadin, 1965, Moll. USSR: 252, fig. 268.
    (intro'd as intra-sp. of Anodonta piscinalis Nilsson, 1822)
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  Anodonta anatina var. petshorica Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 129, fig. 44.
    (intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)
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  Anodonta anatina var. lenae Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 129, fig. 45.
    (intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)
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  Anodonta woodiana var. elliptica Zhadin, 1938
     ‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 144, fig. 54.
     Zhadin, 1965, Moll. USSR: 256, fig. 276.
    (misidentified reference to Anodon elliptica Heude, 1878)
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  Anodonta beringiana var. suifunensis Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 132, fig. 48.
     Zhadin, 1965, Moll. USSR: 255, fig. 272.
    (intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)
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  Anodonta beringiana var. taranetzi Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 133, fig. 49.
     Zhadin, 1965, Moll. USSR: 255, fig. 273.
    (intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)
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  Unio mingrelicus var. stevenianiformis Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 96, text fig. 28.
    (intro'd as intra-sp. of Unio mingrelicus Drouët, 1881)
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  Unio tumidus var. ilikensis Zhadin, 1938
    ‘Kobelt’  of Authors.
     Zhadin, 1938, Faune USSR Mollusques 4(1): 83.
     Kantor & Sysoev, 2005, Cat. Moll. Russia: 327.
    (error for Unio tumidus ilekensis Kobelt, 1912)
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  Unio crassus var. chlebnikovi Zhadin, 1938
     ‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques 4(1): 89.
    (new name for Unio kungurensis chlebnikowi Kobelt, 1912)
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  Unio crassus var. stepanovi Zhadin, 1938
     ‘Drouët’ Zhadin, 1938, Faune USSR Mollusques 4(1): 90.
    (error for Unio stepanoffi Drouët, 1881)
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  Unio crassus consentancus Zhadin, 1938
     ‘Ziegler’ Zhadin, 1938, Faune USSR Mollusques 4(1): 91.
    (error for Unio consentaneus Rossmässler, 1836)
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  Unio raddei var. kutaisiana Zhadin, 1938
    ‘Kobelt’  of Authors.
     Zhadin, 1938, Faune USSR Mollusques 4(1): 94.
     Zatravkin, 1987, Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 49.
    (error for Unio raddei var. koutaisiana Kobelt, 1886)
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  Unio tumidus var. bashkiricus Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 81, text fig. 19.
    (intro'd as intra-sp. of Unio tumidus Retzius, 1788)
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  Unio tumidus var. moltshanovi Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 85, text fig. 20.
    (intro'd as intra-sp. of Unio tumidus Retzius, 1788)
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  Unio tumidus var. fridmani Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 86, text fig. 21.
    (intro'd as intra-sp. of Unio tumidus Retzius, 1788)
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  Unio tumidus var. kobeltianus Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 82.
    (intro'd as intra-sp. of Unio tumidus Retzius, 1788; new name for Unio tumidus var. okae Kobelt & Haas, 1910; homonym of Unio batavus kobeltianus Haas, 1913)
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  Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938
     ‘Westerlund’ Zhadin, 1938, Faune USSR Mollusques 4(1): 99.
    (error for Unio schrencki Westerlund, 1897)
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  Psilunio rothi var. komarovi Zhadin, 1938
     ‘Boettg.’ Zhadin, 1938, Faune USSR Mollusques 4(1): 102.
    (error for Unio rothi var. komarowi O. Boettger, 1880)
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  Anodonta cellensis var. sorensis Zhadin, 1938
     ‘Dybowski’ Zhadin, 1938, Faune USSR Mollusques 4(1): 120.
    (error for Anodonta complanata var. sorensiana Dybowski, 1913)
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  Anodonta sedakovi Zhadin, 1938
     ‘Siemaschko’ Zhadin, 1938, Faune USSR Mollusques 4(1): 130.
     Zhadin, 1965, Moll. USSR: 252.
     Kantor & Sysoev, 2005, Cat. Moll. Russia: 334 [as ‘Colletopterum ponderosum sedakovi’].
     Bogatov et al., 2005, Zool. Zhur. 84(9): 1059.
     Vinarski & Kantor, 2016, Analyt. Cat. Fr. Brack. Moll. Russia: 38.
     Klishko et al., 2018, PLoS One 13(4), e0194944: 3.
    (error for Anodonta sedakowi Siemaschko, 1848)
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  Anodonta cyrea var. samarkadensis Zhadin, 1938
     ‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques 4(1): 140.
    (error for Anodonta samarkandensis Kobelt, 1913)
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  Unio crassus var. ishmensis Zhadin, 1938
     Zhadin, 1938, Faune USSR Mollusques 4(1): 160.
    (intro'd as intra-sp. of Unio crassus Retzius, 1788)
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  Sinanodonta (Anemina) shadini Moskvicheva, 1973
     ‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 147, fig. 56 [as ‘Anodonta arcaeformis’].
     Moskvicheva, 1973, Zool. Zhur. 52(6): 828.
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  Sinanodonta (Anemina) fuscoviridis Moskvicheva, 1973
     ‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 149, fig. 58 [as ‘Anodonta (Haasiella) euscaphus’].
     Moskvicheva, 1973, Zool. Zhur. 52(6): 830.
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  Middendorffinaia (Pseudopotomida) shadini Moskvicheva & Starobogatov, 1973
     ‘Haas’ Zhadin, 1938, Faune USSR Mollusques 4(1): 100, fig. 31 [as ‘Unio continentalis’].
     ‘Haas’ Zhadin, 1952, Moll. SSSR (46): 299, fig. 263 [as ‘Unio continentalis’].
     Moskvicheva & Starobogatov, 1973, Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 31, figs. 5а-в.
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Taxonomic Opinions
	 Anodonta (Haasiella) arcaeformis (Heude, 1877)
	 
	Anodonta (Haasiella) arcaeformis var. suifunica Lindholm, 1925
	 
	Anodonta (Haasiella) euscaphys (Heude, 1879)
	 
	Anodonta (Pseudanodonta) complanata ‘Ziegl.’ Rossmässler, 1835
	 
	Anodonta (Pseudanodonta) complanata var. borealis Kobelt, 1891
	 
	Anodonta (Pseudanodonta) complanata var. middendorffi Siemaschko, 1848
	 
	Anodonta (Pseudanodonta) complanata var. nordenskiöldi Boettger, 1880 [sic]
	 
	Anodonta (Pseudanodonta) rossmaessleri Bourguignat, 1881
	 
	Anodonta (Pteranodon) woodiana (Lea, 1834)
	 
	Anodonta (Pteranodon) woodiana var. elliptica (Heude, 1878)
	 
	Anodonta (s.s.) anatina (Linnaeus, 1758)
	 
	Anodonta (s.s.) anatina var. convexa Drouet, 1888
	 
	Anodonta (s.s.) anatina var. lenae Zhadin, 1938
	 
	Anodonta (s.s.) anatina var. petshorica Zhadin, 1938
	 
	Anodonta (s.s.) anatina var. viridiflava Drouet, 1893
	 
	Anodonta (s.s.) beringiana Middendorff, 1851
	 
	Anodonta (s.s.) beringiana var. suifunensis Zhadin, 1938
	 
	Anodonta (s.s.) beringiana var. taranetzi Zhadin, 1938
	 
	Anodonta (s.s.) cellensis [sic] (Gmelin, 1791)
	 
	Anodonta (s.s.) cellensis [sic] var. cariosa Küster, 1876 [sic]
	 
	Anodonta (s.s.) cellensis [sic] var. maganica Servain, 1882
	 
	Anodonta (s.s.) cellensis [sic] var. selengensis Dybowski, 1913
	 
	Anodonta (s.s.) cellensis [sic] var. sorensis [sic] Dybowski, W., 1913
	 
	Anodonta (s.s.) cellensis [sic] var. tehernaica [sic] Bourguignat, 1881
	 + Anodonta tchernaica Bourguignat, 1881
	 
	Anodonta (s.s.) cygnea (Linnaeus, 1758)
	 
	Anodonta (s.s.) cyrea Drouet, 1881
	 
	Anodonta (s.s.) cyrea var. armenica Ovtschinnikov, 1935
	 
	Anodonta (s.s.) cyrea var. bactriana Rolle, 1897
	 
	Anodonta (s.s.) cyrea var. georgiana Drouet, 1881
	 
	Anodonta (s.s.) cyrea var. lenkoranensis Drouet, 1881
	 
	Anodonta (s.s.) cyrea var. samarkadensis [sic] Kobelt, 1913
	 
	Anodonta (s.s.) cyrea var. sogdiana Kobelt, 1896
	 
	Anodonta (s.s.) piscinalis Nilsson, 1823
	 
	Anodonta (s.s.) piscinalis var. crimeana Bourguignat, 1881
	 
	Anodonta (s.s.) piscinalis var. letourneuxi Bourguignat, 1870
	 
	Anodonta (s.s.) piscinalis var. ostiaria Drouet, 1881
	 
	Anodonta (s.s.) piscinalis var. ponderosa Pfeiffer, C., 1825
	 
	Anodonta (s.s.) piscinalis var. seisanensis Kobelt, 1912
	 
	Anodonta (s.s.) piscinalis var. sorica Dybowski, 1913
	 
	Anodonta (s.s.) piscinalis var. volgensis Zhadin, 1938
	 
	Anodonta (s.s.) sedakovi [sic] Siemaschko, 1848
	 
	Anodonta (s.s.) sedakowi var. nova Dybowski, 1913
	 
	Cristaria plicata (Leach, 1815)
	 
	Cristaria plicata var. herculea Middendorff, 1848
	 
	Lanceolaria cylindrica Simpson, 1900
	 
	Lanceolaria grayana (Lea, 1834)
	 
	Margaritana dahurica (Middendorff, 1850)
	 
	Margaritana margaritifera (Linnaeus, 1758)
	 
	Margaritana middendorffi Rosén, 1926
	 + Unio (Alasm.) complanatus Middendorff, 1851
	 
	Margaritana mongolica (Middendorff, 1851)
	 
	Margaritana sachalinensis Zhadin, 1938
	 
	Psilunio rothi (Bourguignat, 1863)
	 
	Psilunio rothi var. armeniacus (Kobelt, 1912)
	 
	Psilunio rothi var. komarovi [sic] (Boettger, O., 1880)
	 
	Unio (Nodularia) continentalis (Haas, 1910)
	 
	Unio (Nodularia) douglasiae Gray in Griffith & Pidgeon, 1834
	 
	Unio (Nodularia) douglasiae var. schrenki [sic] Westerlund, 1897
	 + Unio abbreviatus Westerlund, 1897
	 
	Unio (s.s.) ater Nilsson, 1822
	 
	Unio (s.s.) crassus Retzius, 1788
	 
	Unio (s.s.) crassus consentancus [sic] ‘Ziegler’ Rossmässler, 1836
	 
	Unio (s.s.) crassus kungurensis Kobelt, 1912
	 
	Unio (s.s.) crassus var. chlebnikovi [sic] Kobelt, 1912
	 + Unio kungurensis sylwensis Kobelt, 1912
	 + Unio kungurensis irenjensis Kobelt, 1912
	 
	Unio (s.s.) crassus var. ishmensis Zhadin, 1938
	 
	Unio (s.s.) crassus var. okae Kobelt, 1911
	 
	Unio (s.s.) crassus var. pseudolitoralis Clessin, 1875
	 
	Unio (s.s.) crassus var. roseni Kobelt, 1912
	 
	Unio (s.s.) crassus var. stepanovi [sic] Drouet, 1881
	 
	Unio (s.s.) hueti Bourguignat, 1855
	 
	Unio (s.s.) lindholmi Zhadin, 1938
	 
	Unio (s.s.) mingrelicus Drouet, 1881
	 
	Unio (s.s.) mingrelicus var. gregorii Kobelt, 1912
	 
	Unio (s.s.) mingrelicus var. stevenianiformis Zhadin, 1938
	 
	Unio (s.s.) pictorum (Linnaeus, 1758)
	 
	Unio (s.s.) pictorum defectivus Zhadin, 1938
	 
	Unio (s.s.) pictorum gentilis Haas in Hesse, 1911
	 
	Unio (s.s.) pictorum var. annulatus Kobelt, 1912
	 
	Unio (s.s.) pictorum var. okensis Zhadin, 1938
	 + Unio pictorum var. okae Kobelt & Haas, 1910
	 
	Unio (s.s.) pictorum var. platyrhynchus Rossmässler, 1835
	 
	Unio (s.s.) pictorum var. ponderosus Spitzi in Rossmässler, 1838
	 
	Unio (s.s.) pictorum var. pygmaeus Zhadin, 1938
	 
	Unio (s.s.) pictorum var. schrenkianus Clessin, 1880
	 
	Unio (s.s.) sieversi Drouet, 1881
	 
	Unio (s.s.) sieversi var. araxenus Drouet, 1881
	 
	Unio (s.s.) sieversi var. colchicus Drouet, 1881
	 
	Unio (s.s.) sieversi var. kutaisiana [sic] Kobelt, 1886
	 
	Unio (s.s.) sieversi var. raddei Drouet, 1881
	 
	Unio (s.s.) stevenianus Drouet, 1881
	 
	Unio (s.s.) tumidus Retzius, 1788
	 
	Unio (s.s.) tumidus limicola Mörch, 1864
	 
	Unio (s.s.) tumidus var. bashkiricus Zhadin, 1938
	 
	Unio (s.s.) tumidus var. borysthenicus Kobelt, 1879
	 
	Unio (s.s.) tumidus var. conus Spengler, 1793
	 
	Unio (s.s.) tumidus var. fridmani Zhadin, 1938
	 
	Unio (s.s.) tumidus var. gerstfeldtianus Clessin, 1880
	 
	Unio (s.s.) tumidus var. ilikensis [sic] Kobelt, 1912
	 
	Unio (s.s.) tumidus var. kobeltianus Zhadin, 1938
	 + Unio tumidus var. okae Kobelt & Haas, 1910
	 
	Unio (s.s.) tumidus var. moltshanovi Zhadin, 1938
	 
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