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Page last updated
Fri 04 Oct 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Zhadin (1938), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Zhadin, V.I. 1938. Sem. Unionidae. Faune URSS, Mollusques 4(1). 169 pp.

Attributed Species Names

Mytilus cellensis Menke, 1828
‘Gmelin’ of Authors.
Menke, 1828, Syn. Method. Moll.: 63.
Menke, 1830, Syn. Method. Moll.: 106.
Küster, 1839, Conchyl. Cab. 9(1): 16.
Mörch, 1853, Cat. Conch. Yoldi 2: 48.
Clessin, 1877, Deutsche Excurs. Moll. Fauna: 438.
Drouët, 1881, Unionidae Russie: 22.
Bourguignat, 1882, Natural. Sicil. 2 (1): 20 [1].
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 211.
Zhadin, 1938, Faune USSR Mollusques 4(1): 119.
Modell, 1945, Jena. Z. Med. Naturw. 78: 81.
(error for Mytilus zellensis Gmelin, 1791)

Anodonta tehernaica Westerlund, 1890
‘Bourguignat’ of Authors.
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 226.
Zhadin, 1938, Faune USSR Mollusques 4(1): 121.
(error for Anodonta tchernaica Bourguignat, 1881)

Unio (Nodularia) continentalis Zhadin, 1938
‘Haas’ Zhadin, 1938, Faune USSR Mollusques 4(1): 100, fig. 31.
‘Haas’ Zhadin, 1952, Moll. SSSR (46): 299, fig. 263.
(misidentified reference to Nodularia continentalis Haas, 1910)

Margaritana sachalinensis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 114-115, fig. 37.
Zhadin, 1965, Moll. USSR: 238, fig. 252.

Unio lindholmi Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 97, text fig. 29.

Unio pictorum var. okensis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 77.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; new name for Unio pictorum var. okae Kobelt & Haas, 1910)

Unio pictorum var. pygmaeus Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 78, text fig. 17.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758; homonym of Unio pygmaeus Lea, 1852)

Unio pictorum var. defectivus Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 79.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)

Anodonta (Haasiella) arcaeformis Zhadin, 1938
‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 147, fig. 56.
Zhadin, 1965, Moll. USSR: 260, fig. 279.
(misidentified reference to Anodon arcaeformis Heude, 1877)

Anodonta (Haasiella) euscaphus Zhadin, 1938
‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 149, fig. 58.
Zhadin, 1965, Moll. USSR: 260, fig. 280.
(misidentified reference to Anodon euscaphys Heude, 1879; error for Anodon euscaphys Heude, 1879)

Anodonta piscinalis var. volgensis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 126, fig. 43.
Zhadin, 1965, Moll. USSR: 252, fig. 268.
(intro'd as intra-sp. of Anodonta piscinalis Nilsson, 1822)

Anodonta anatina var. petshorica Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 129, fig. 44.
(intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)

Anodonta anatina var. lenae Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 129, fig. 45.
(intro'd as intra-sp. of Mytilus anatinus Linnaeus, 1758)

Anodonta woodiana var. elliptica Zhadin, 1938
‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 144, fig. 54.
Zhadin, 1965, Moll. USSR: 256, fig. 276.
(misidentified reference to Anodon elliptica Heude, 1878)

Anodonta beringiana var. suifunensis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 132, fig. 48.
Zhadin, 1965, Moll. USSR: 255, fig. 272.
(intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)

Anodonta beringiana var. taranetzi Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 133, fig. 49.
Zhadin, 1965, Moll. USSR: 255, fig. 273.
(intro'd as intra-sp. of Anodonta cellensis var. beringiana Middendorff, 1851)

Unio mingrelicus var. stevenianiformis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 96, text fig. 28.
(intro'd as intra-sp. of Unio mingrelicus Drouët, 1881)

Unio tumidus var. ilikensis Zhadin, 1938
‘Kobelt’ of Authors.
Zhadin, 1938, Faune USSR Mollusques 4(1): 83.
Kantor & Sysoev, 2005, Cat. Moll. Russia: 327.
(error for Unio tumidus ilekensis Kobelt, 1912)

Unio crassus var. chlebnikovi Zhadin, 1938
‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques 4(1): 89.
(new name for Unio kungurensis chlebnikowi Kobelt, 1912)

Unio crassus var. stepanovi Zhadin, 1938
‘Drouët’ Zhadin, 1938, Faune USSR Mollusques 4(1): 90.
(error for Unio stepanoffi Drouët, 1881)

Unio crassus consentancus Zhadin, 1938
‘Ziegler’ Zhadin, 1938, Faune USSR Mollusques 4(1): 91.
(error for Unio consentaneus Rossmässler, 1836)

Unio raddei var. kutaisiana Zhadin, 1938
‘Kobelt’ of Authors.
Zhadin, 1938, Faune USSR Mollusques 4(1): 94.
Zatravkin, 1987, Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 49.
(error for Unio raddei var. koutaisiana Kobelt, 1886)

Unio tumidus var. bashkiricus Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 81, text fig. 19.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788)

Unio tumidus var. moltshanovi Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 85, text fig. 20.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788)

Unio tumidus var. fridmani Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 86, text fig. 21.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788)

Unio tumidus var. kobeltianus Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 82.
(intro'd as intra-sp. of Unio tumidus Retzius, 1788; new name for Unio tumidus var. okae Kobelt & Haas, 1910; homonym of Unio batavus kobeltianus Haas, 1913)

Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938
‘Westerlund’ Zhadin, 1938, Faune USSR Mollusques 4(1): 99.
(error for Unio schrencki Westerlund, 1897)

Psilunio rothi var. komarovi Zhadin, 1938
‘Boettg.’ Zhadin, 1938, Faune USSR Mollusques 4(1): 102.
(error for Unio rothi var. komarowi O. Boettger, 1880)

Anodonta cellensis var. sorensis Zhadin, 1938
‘Dybowski’ Zhadin, 1938, Faune USSR Mollusques 4(1): 120.
(error for Anodonta complanata var. sorensiana Dybowski, 1913)

Anodonta sedakovi Zhadin, 1938
‘Siemaschko’ Zhadin, 1938, Faune USSR Mollusques 4(1): 130.
Zhadin, 1965, Moll. USSR: 252.
Kantor & Sysoev, 2005, Cat. Moll. Russia: 334 [as ‘Colletopterum ponderosum sedakovi’].
Bogatov et al., 2005, Zool. Zhur. 84(9): 1059.
Vinarski & Kantor, 2016, Analyt. Cat. Fr. Brack. Moll. Russia: 38.
Klishko et al., 2018, PLoS One 13(4), e0194944: 3.
(error for Anodonta sedakowi Siemaschko, 1848)

Anodonta cyrea var. samarkadensis Zhadin, 1938
‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques 4(1): 140.
(error for Anodonta samarkandensis Kobelt, 1913)

Unio crassus var. ishmensis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 160.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Sinanodonta (Anemina) shadini Moskvicheva, 1973
‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 147, fig. 56 [as ‘Anodonta arcaeformis’].
Moskvicheva, 1973, Zool. Zhur. 52(6): 828.

Sinanodonta (Anemina) fuscoviridis Moskvicheva, 1973
‘Heude’ Zhadin, 1938, Faune USSR Mollusques 4(1): 149, fig. 58 [as ‘Anodonta (Haasiella) euscaphus’].
Moskvicheva, 1973, Zool. Zhur. 52(6): 830.

Middendorffinaia (Pseudopotomida) shadini Moskvicheva & Starobogatov, 1973
‘Haas’ Zhadin, 1938, Faune USSR Mollusques 4(1): 100, fig. 31 [as ‘Unio continentalis’].
‘Haas’ Zhadin, 1952, Moll. SSSR (46): 299, fig. 263 [as ‘Unio continentalis’].
Moskvicheva & Starobogatov, 1973, Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 31, figs. 5а-в.

Taxonomic Opinions

Anodonta (Haasiella) arcaeformis (Heude, 1877)

Anodonta (Haasiella) arcaeformis var. suifunica Lindholm, 1925

Anodonta (Haasiella) euscaphys (Heude, 1879)

Anodonta (Pseudanodonta) complanata ‘Ziegl.’ Rossmässler, 1835

Anodonta (Pseudanodonta) complanata var. borealis Kobelt, 1891

Anodonta (Pseudanodonta) complanata var. middendorffi Siemaschko, 1848

Anodonta (Pseudanodonta) complanata var. nordenskiöldi Boettger, 1880 [sic]

Anodonta (Pseudanodonta) rossmaessleri Bourguignat, 1881

Anodonta (Pteranodon) woodiana (Lea, 1834)

Anodonta (Pteranodon) woodiana var. elliptica (Heude, 1878)

Anodonta (s.s.) anatina (Linnaeus, 1758)

Anodonta (s.s.) anatina var. convexa Drouet, 1888

Anodonta (s.s.) anatina var. lenae Zhadin, 1938

Anodonta (s.s.) anatina var. petshorica Zhadin, 1938

Anodonta (s.s.) anatina var. viridiflava Drouet, 1893

Anodonta (s.s.) beringiana Middendorff, 1851

Anodonta (s.s.) beringiana var. suifunensis Zhadin, 1938

Anodonta (s.s.) beringiana var. taranetzi Zhadin, 1938

Anodonta (s.s.) cellensis [sic] (Gmelin, 1791)

Anodonta (s.s.) cellensis [sic] var. cariosa Küster, 1876 [sic]

Anodonta (s.s.) cellensis [sic] var. maganica Servain, 1882

Anodonta (s.s.) cellensis [sic] var. selengensis Dybowski, 1913

Anodonta (s.s.) cellensis [sic] var. sorensis [sic] Dybowski, W., 1913

Anodonta (s.s.) cellensis [sic] var. tehernaica [sic] Bourguignat, 1881
+ Anodonta tchernaica Bourguignat, 1881

Anodonta (s.s.) cygnea (Linnaeus, 1758)

Anodonta (s.s.) cyrea Drouet, 1881

Anodonta (s.s.) cyrea var. armenica Ovtschinnikov, 1935

Anodonta (s.s.) cyrea var. bactriana Rolle, 1897

Anodonta (s.s.) cyrea var. georgiana Drouet, 1881

Anodonta (s.s.) cyrea var. lenkoranensis Drouet, 1881

Anodonta (s.s.) cyrea var. samarkadensis [sic] Kobelt, 1913

Anodonta (s.s.) cyrea var. sogdiana Kobelt, 1896

Anodonta (s.s.) piscinalis Nilsson, 1823

Anodonta (s.s.) piscinalis var. crimeana Bourguignat, 1881

Anodonta (s.s.) piscinalis var. letourneuxi Bourguignat, 1870

Anodonta (s.s.) piscinalis var. ostiaria Drouet, 1881

Anodonta (s.s.) piscinalis var. ponderosa Pfeiffer, C., 1825

Anodonta (s.s.) piscinalis var. seisanensis Kobelt, 1912

Anodonta (s.s.) piscinalis var. sorica Dybowski, 1913

Anodonta (s.s.) piscinalis var. volgensis Zhadin, 1938

Anodonta (s.s.) sedakovi [sic] Siemaschko, 1848

Anodonta (s.s.) sedakowi var. nova Dybowski, 1913

Cristaria plicata (Leach, 1815)

Cristaria plicata var. herculea Middendorff, 1848

Lanceolaria cylindrica Simpson, 1900

Lanceolaria grayana (Lea, 1834)

Margaritana dahurica (Middendorff, 1850)

Margaritana margaritifera (Linnaeus, 1758)

Margaritana middendorffi Rosén, 1926
+ Unio (Alasm.) complanatus Middendorff, 1851

Margaritana mongolica (Middendorff, 1851)

Margaritana sachalinensis Zhadin, 1938

Psilunio rothi (Bourguignat, 1863)

Psilunio rothi var. armeniacus (Kobelt, 1912)

Psilunio rothi var. komarovi [sic] (Boettger, O., 1880)

Unio (Nodularia) continentalis (Haas, 1910)

Unio (Nodularia) douglasiae Gray in Griffith & Pidgeon, 1834

Unio (Nodularia) douglasiae var. schrenki [sic] Westerlund, 1897
+ Unio abbreviatus Westerlund, 1897

Unio (s.s.) ater Nilsson, 1822

Unio (s.s.) crassus Retzius, 1788

Unio (s.s.) crassus consentancus [sic] ‘Ziegler’ Rossmässler, 1836

Unio (s.s.) crassus kungurensis Kobelt, 1912

Unio (s.s.) crassus var. chlebnikovi [sic] Kobelt, 1912
+ Unio kungurensis sylwensis Kobelt, 1912
+ Unio kungurensis irenjensis Kobelt, 1912

Unio (s.s.) crassus var. ishmensis Zhadin, 1938

Unio (s.s.) crassus var. okae Kobelt, 1911

Unio (s.s.) crassus var. pseudolitoralis Clessin, 1875

Unio (s.s.) crassus var. roseni Kobelt, 1912

Unio (s.s.) crassus var. stepanovi [sic] Drouet, 1881

Unio (s.s.) hueti Bourguignat, 1855

Unio (s.s.) lindholmi Zhadin, 1938

Unio (s.s.) mingrelicus Drouet, 1881

Unio (s.s.) mingrelicus var. gregorii Kobelt, 1912

Unio (s.s.) mingrelicus var. stevenianiformis Zhadin, 1938

Unio (s.s.) pictorum (Linnaeus, 1758)

Unio (s.s.) pictorum defectivus Zhadin, 1938

Unio (s.s.) pictorum gentilis Haas in Hesse, 1911

Unio (s.s.) pictorum var. annulatus Kobelt, 1912

Unio (s.s.) pictorum var. okensis Zhadin, 1938
+ Unio pictorum var. okae Kobelt & Haas, 1910

Unio (s.s.) pictorum var. platyrhynchus Rossmässler, 1835

Unio (s.s.) pictorum var. ponderosus Spitzi in Rossmässler, 1838

Unio (s.s.) pictorum var. pygmaeus Zhadin, 1938

Unio (s.s.) pictorum var. schrenkianus Clessin, 1880

Unio (s.s.) sieversi Drouet, 1881

Unio (s.s.) sieversi var. araxenus Drouet, 1881

Unio (s.s.) sieversi var. colchicus Drouet, 1881

Unio (s.s.) sieversi var. kutaisiana [sic] Kobelt, 1886

Unio (s.s.) sieversi var. raddei Drouet, 1881

Unio (s.s.) stevenianus Drouet, 1881

Unio (s.s.) tumidus Retzius, 1788

Unio (s.s.) tumidus limicola Mörch, 1864

Unio (s.s.) tumidus var. bashkiricus Zhadin, 1938

Unio (s.s.) tumidus var. borysthenicus Kobelt, 1879

Unio (s.s.) tumidus var. conus Spengler, 1793

Unio (s.s.) tumidus var. fridmani Zhadin, 1938

Unio (s.s.) tumidus var. gerstfeldtianus Clessin, 1880

Unio (s.s.) tumidus var. ilikensis [sic] Kobelt, 1912

Unio (s.s.) tumidus var. kobeltianus Zhadin, 1938
+ Unio tumidus var. okae Kobelt & Haas, 1910

Unio (s.s.) tumidus var. moltshanovi Zhadin, 1938

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