|                Page last updated 
                  Sun 07 Sep 2025  | 
              
              
  
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win! 
This page documents data for Sano et al. (2017), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work. The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=publ&id=3799 
Search the database pages:
 
 
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 Publication information 
Sano, I., A. Shirai, T. Kondo & J.-I. Miyazaki. 2017. Phylogenetic relationships of Japanese Unionoida (Mollusca: Bivalvia) based on mitochondrial 16S rDNA sequences. Journal of Water Resources and Protection 9: 493-509. 
Click here for online access. 
Cladogram Figures
(click the figure number to see the cladogram) fig. 1 fig. 2
  
Attributed Species Names
  Pletholophus tenuis Sano et al., 2017
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2.
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1.
     Sano et al., 2022, MPE 175: Table S1.
    (misidentified reference to Unio tenuis Griffith & Pidgeon, 1833)
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  Ptychorhynchus ptisteri Sano et al., 2017
     Sano et al., 2017, J. Wat. Res. Prot. 9: 498.
     Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
    (error for Unio pfisteri Heude, 1874)
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  Sinanodonta japonica Sano et al., 2017
    ‘Clessin’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1. [in part]
    (misidentified reference to Anodonta japonica Clessin, 1874)
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  Sinanodonta japonica Sano et al., 2017
    ‘Clessin’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1. [in part]
    (misidentified reference to Anodonta japonica Clessin, 1874)
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  Sinanodonta calipygos Sano et al., 2017
    ‘Kobelt’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1.
    (misidentified reference to Anodonta calipygos Kobelt, 1879)
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  Sinanodonta lauta Sano et al., 2017
    ‘Martens’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1. [in part]
    (misidentified reference to Anodonta lauta Martens, 1877)
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  Sinanodonta lauta Sano et al., 2017
    ‘Martens’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1. [in part]
    (misidentified reference to Anodonta lauta Martens, 1877)
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  Nodularia douglasiae nipponensis Sano et al., 2017
    ‘Martens’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
    (misidentified reference to Unio nipponensis Martens, 1877)
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  Nodularia douglasiae biwae Sano et al., 2017
    ‘Kobelt’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2.
    (misidentified reference to Unio biwae Kobelt, 1879)
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  Nodularia douglasiae nipponensis Sano et al., 2017
    ‘Martens’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
    (misidentified reference to Unio nipponensis Martens, 1877)
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  Anemina arcaeformis Sano et al., 2017
    ‘Heude’  of Authors.
     Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2.
     Sano et al., 2020, Plankton Benthos Res. 15(2): Table 1. [in part]
    (misidentified reference to Anodon arcaeformis Heude, 1877)
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Taxonomic Opinions
	 Acuticosta ovata 
	 
	Anemina arcaeformis Heude, 1977 [sic]
	 
	Arconaia lanceolata 
	 
	Cristaria plicata Leach, 1815
	 
	Cuneopsis pisciculus 
	 
	Hyriopsis cumingii 
	 
	Hyriopsis schlegeli Martens, 1861
	 
	Inversidens brandti Kobelt, 1879
	 
	Inversiunio jokohamensis Ihering, 1893
	 
	Inversiunio reinianus Kobelt, 1879
	 
	Inversiunio yanagawensis Kondo, 1982
	 
	Lamprotula leai 
	 
	Lanceolaria grayii Gray in Griffith & Pidgeon, 1833
	 
	Lepidodesma languilati 
	 
	Margaritifera laevis Haas, 1910
	 
	Margaritifera togakushiensis Kondo & Kobayashi, 2005
	 
	Neotrigonia lamarckii 
	 
	Neotrigonia margaritacea 
	 
	Nodularia douglasiae Gray in Griffith & Pidgeon, 1833
	 
	Nodularia douglasiae biwae 
	 
	Nodularia douglasiae nipponensis 
	 
	Obovalis omiensis Heimburg, 1884
	 
	Pletholophus tenuis Gray in Griffith & Pidgeon, 1833
	 
	Pronodularia japanensis Lea, 1859
	 
	Ptychorhynchus ptisteri [sic] 
	 
	Schistodesmus lampreyanus 
	 
	Sinanodonta calipygos Kobelt, 1879
	 
	Sinanodonta japonica Clessin, 1874
	 
	Sinanodonta lauta Martens, 1877
	 
	Sinanodonta ogurae Kuroda & Habe, 1987
	 
	Sinanodonta woodiana 
	 
	Solenaia oleivora 
	 
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Cladogram Figure 1
                                +-- Neotrigonia_margaritacea   
  +-----------------------------+                              
  |                             +-- Neotrigonia_lamarckii      
  |                                                            
  |                       +-------- Margaritifera_laevis__2    
  |  +--------------------+                                    
  |  |                    |  +----- Margaritifera_laevis__1    
--+  |                    +--+                                 
  |  |                       |  +-- Margaritifera_middendorffi 
  |  |                       +--+                              
  |  |                          +-- Margaritifera_laevis__3    
  |  |                                                         
  |  |                    +-------- Obovalis_omiensis          
  +--+  +-----------------+                                    
     |  |                 |  +----- Inversidens_brandtii       
     |  |                 +--+                                 
     |  |                    |  +-- Pronodularia_japanensis    
     |  |                    +--+                              
     |  |                       +-- Sinohyriopsis_schlegelii   
     |  |                                                      
     |  |     +-------------------- Lanceolaria_grayii         
     +--+  +--+                                                
        |  |  |  +----------------- Nodularia_nipponensis      
        |  |  +--+                                             
        |  |     |  +-------------- Inversiunio_jokohamensis   
        |  |     +--+                                          
        |  |        |  +----------- Inversiunio_yanagawensis__1
        |  |        +--+                                       
        |  |           |  +-------- Inversiunio_yanagawensis__2
        |  |           +--+                                    
        +--+              |  +----- Inversiunio_yanagawensis__3
           |              +--+                                 
           |                 |  +-- Inversiunio_yanagawensis__4
           |                 +--+                              
           |                    +-- Inversiunio_reinianus      
           |                                                   
           |                    +-- Cristaria_plicata          
           |        +-----------+                              
           |        |           +-- Buldowskia_iwakawai        
           +--------+                                          
                    |  +----------- Pletholophus_reinianus     
                    |  |                                       
                    +--+        +-- Sinanodonta_tumens         
                       |  +-----+                              
                       |  |     +-- Sinanodonta_lauta          
                       +--+                                    
                          |  +----- Beringiana_fukuharai       
                          +--+                                 
                             |  +-- Beringiana_japonica        
                             +--+                              
                                +-- Beringiana_beringiana      
 
Cladogram Figure 2
                             +-- Neotrigonia_lamarckii      
  +--------------------------+                              
  |                          +-- Neotrigonia_margaritacea   
  |                                                         
  |                          +-- Margaritifera_laevis       
  |  +-----------------------+                              
  |  |                       +-- Margaritifera_middendorffi 
  |  |                                                      
--+  |                       +-- Obovalis_omiensis          
  |  |              +--------+                              
  |  |              |        +-- Ptychorhynchus_pfisteri    
  |  |              |                                       
  |  |  +-----------+        +-- Inversidens_brandtii       
  |  |  |           |  +-----+                              
  |  |  |           |  |     +-- Sinosolenaia_oleivora      
  +--+  |           |  |                                    
     |  |           +--+     +-- Pronodularia_japanensis    
     |  |              |  +--+                              
     |  |              |  |  +-- Lamprotula_leaii           
     |  |              +--+                                 
     |  |                 |  +-- Sinohyriopsis_schlegelii   
     |  |                 +--+                              
     |  |                    +-- Sinohyriopsis_cumingii     
     |  |                                                   
     |  |                    +-- Cristaria_plicata          
     +--+        +-----------+                              
        |        |           +-- Buldowskia_iwakawai        
        |        |                                          
        |  +-----+        +----- Beringiana_fukuharai       
        |  |     |  +-----+                                 
        |  |     |  |     |  +-- Beringiana_japonica        
        |  |     |  |     +--+                              
        |  |     +--+        +-- Beringiana_beringiana      
        |  |        |                                       
        |  |        |  +-------- Pletholophus_reinianus     
        |  |        +--+                                    
        |  |           |  +----- Sinanodonta_woodiana       
        |  |           +--+                                 
        |  |              |  +-- Sinanodonta_lauta          
        +--+              +--+                              
           |                 +-- Sinanodonta_tumens         
           |                                                
           |        +----------- Tchangsinaia_piscicula     
           |  +-----+                                       
           |  |     |  +-------- Nodularia_nipponensis__1   
           |  |     +--+                                    
           |  |        |  +----- Nodularia_breviconcha      
           |  |        +--+                                 
           |  |           |  +-- Nodularia_nipponensis__2   
           |  |           +--+                              
           +--+              +-- Nodularia_douglasiae       
              |                                             
              |              +-- Schistodesmus_lampreyanus  
              |  +-----------+                              
              |  |           +-- Acuticosta_ovata           
              |  |                                          
              |  |     +-------- Lepidodesma_languilati     
              +--+  +--+                                    
                 |  |  |  +----- Lanceolaria_grayii__1      
                 |  |  +--+                                 
                 |  |     |  +-- Lanceolaria_lanceolata     
                 +--+     +--+                              
                    |        +-- Lanceolaria_grayii__2      
                    |                                       
                    |  +-------- Inversiunio_jokohamensis   
                    |  |                                    
                    +--+  +----- Inversiunio_yanagawensis__3
                       |  |                                 
                       +--+  +-- Inversiunio_yanagawensis__1
                          |  |                              
                          +--+-- Inversiunio_yanagawensis__2
                             |                              
                             +-- Inversiunio_reinianus      
 
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  Page updated: 08:32:46 Sun 07 Sep 2025  |  
 
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