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                  Sun 07 Sep 2025  | 
              
              
  
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win! 
This page documents data for Inoue et al. (2020), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work. The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=publ&id=4128 
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 Publication information 
Inoue, K., K.S. Cummings, J.S. Tiemann, T.D. Miller, N.A. Johnson et al. 2020. A new species of freshwater mussel in the genus Popenaias Frierson, 1927, from the Gulf coast rivers of central Mexico (Bivalvia: Unionida: Unionidae) with comments on the genus. Zootaxa 4816(4): 457-490. 
Click here for online access. 
Cladogram Figures
(click the figure number to see the cladogram) fig. 2 fig. 3
  
Attributed Species Names
  Popenaias berezai Inoue et al., 2020
     Inoue et al., 2020, Zootaxa 4816(4): 464, figs. 5-7, 8A, 9A.
    Type(s): HOLOTYPE (ORIG. DESIG.) CNMO_8037, Rio Valles (= Rio Micos), Estacion Micos, San Luis Potosi, Mexico. 22.11417, -99.16166.
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Taxonomic Opinions
	 Actinonaias ligamentina 
	 + Unio callosus Lea, 1841
	 
	Amblema elliottii 
	 
	Amblema neislerii 
	 
	Amblema plicata 
	 
	Cyrtonaias explicata 
	 
	Cyrtonaias tampicoensis 
	 
	Epioblasma triquetra 
	 
	Glebula rotundata 
	 
	Lampsilis cardium 
	 
	Ligumia recta 
	 
	Nephronaias ravistella (Morelet, 1849)
	 + Unio veraepacis Tristram, 1863
	 
	Nephronaias rowellii (Lea, 1859)
	 
	Pachynaias spheniopsis 
	 
	Popenaias berezai Inoue et al. n. sp.
	 + Unio popeii Fischer & Crosse, 1894
	 
	Popenaias ganina (Pilsbry, 1910)
	 + Lampsilis semirasa Pilsbry, 1910
	 + Elliptio (Popenaias) popei var. regalis Haas, 1929
	 
	Popenaias metallica (Say, 1831)
	 
	Popenaias popeii (Lea, 1857)
	 
	Popenaias tehuantepecensis (Crosse & Fischer, 1893)
	 
	Potamilus alatus 
	 
	Psoronaias semigranosa 
	 
	Psorula guatemalensis 
	 
	Psorula percompressa 
	 
	Psorula profunda 
	 
	Psorula rudis 
	 
	Psorula usumasintae 
	 
	Ptychobranchus fasciolaris 
	 
	Quadrula quadrula 
	 
	Reginaia ebenus 
	 
	Reginaia rotulata 
	 
	Sphenonaias microdon 
	 
	Truncilla truncata 
	 
	Villosa iris 
	 
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Cladogram Figure 2
  +-------------------------- Quadrula_quadrula          
  |                                                      
  |                       +-- Reginaia_ebenus            
  |  +--------------------+                              
  |  |                    +-- Reginaia_rotulata          
--+  |                                                   
  |  |                 +----- Amblema_plicata            
  |  |  +--------------+                                 
  |  |  |              |  +-- Amblema_elliottii          
  |  |  |              +--+                              
  +--+  |                 +-- Amblema_neislerii          
     |  |                                                
     |  |              +----- Popenaias_popeii           
     |  |           +--+                                 
     |  |           |  |  +-- Popenaias_berezai          
     |  |           |  +--+                              
     |  |           |     +-- Popenaias_semirasa         
     +--+           |                                    
        |  +--------+     +-- Psoronaias_profunda        
        |  |        |     |                              
        |  |        |     +-- Psoronaias_semigranosa__1  
        |  |        |     |                              
        |  |        |     +-- Psoronaias_semigranosa__2  
        |  |        |     |                              
        |  |        +-----+-- Psoronaias_guatemalensis__1
        |  |              |                              
        |  |              +-- Psoronaias_guatemalensis__2
        +--+              |                              
           |              +-- Psoronaias_percompressa    
           |              |                              
           |              +-- Psoronaias_semigranosa__3  
           |                                             
           |              +-- Glebula_rotundata          
           |  +-----------+                              
           |  |           +-- Cyrtonaias_tampicoensis    
           |  |                                          
           +--+  +----------- Potamilus_alatus           
              |  |                                       
              |  |        +-- Ptychobranchus_fasciolaris 
              +--+  +-----+                              
                 |  |     +-- Truncilla_truncata         
                 |  |                                    
                 |  |  +----- Ortmanniana_ligamentina    
                 +--+  |                                 
                    |  +----- Epioblasma_triquetra       
                    |  |                                 
                    |  +----- Lampsilis_cardium          
                    |  |                                 
                    +--+----- Ligumia_recta              
                       |                                 
                       +----- Cambarunio_iris            
                       |                                 
                       +----- Pachynaias_spheniopsis     
                       |                                 
                       |  +-- Cyrtonaias_explicata       
                       +--+                              
                          +-- Sphenonaias_microdon       
 
Cladogram Figure 3
  +----------------------------------------- Quadrula_quadrula          
  |                                                                     
  |                                      +-- Reginaia_ebenus            
  |  +-----------------------------------+                              
--+  |                                   +-- Reginaia_rotulata          
  |  |                                                                  
  |  |                                +----- Amblema_neislerii          
  |  |  +-----------------------------+                                 
  |  |  |                             |  +-- Amblema_plicata            
  +--+  |                             +--+                              
     |  |                                +-- Amblema_elliottii          
     |  |                                                               
     |  |                             +----- Popenaias_popeii           
     |  |                    +--------+                                 
     |  |                    |        |  +-- Popenaias_berezai          
     |  |                    |        +--+                              
     +--+                    |           +-- Popenaias_semirasa         
        |  +-----------------+                                          
        |  |                 |  +----------- Psoronaias_profunda        
        |  |                 |  |                                       
        |  |                 |  |     +----- Psoronaias_guatemalensis__1
        |  |                 +--+  +--+                                 
        |  |                    |  |  |  +-- Psoronaias_semigranosa__1  
        |  |                    |  |  +--+                              
        |  |                    +--+     +-- Psoronaias_percompressa    
        +--+                       |                                    
           |                       |  +----- Psoronaias_semigranosa__3  
           |                       +--+                                 
           |                          |  +-- Psoronaias_guatemalensis__2
           |                          +--+                              
           |                             +-- Psoronaias_semigranosa__2  
           |                                                            
           |  +----------------------------- Cyrtonaias_tampicoensis    
           |  |                                                         
           +--+  +-------------------------- Ptychobranchus_fasciolaris 
              |  |                                                      
              +--+  +----------------------- Ligumia_recta              
                 |  |                                                   
                 +--+  +-------------------- Potamilus_alatus           
                    |  |                                                
                    +--+  +----------------- Pachynaias_spheniopsis     
                       |  |                                             
                       |  |              +-- Epioblasma_triquetra       
                       +--+  +-----------+                              
                          |  |           +-- Truncilla_truncata         
                          |  |                                          
                          +--+           +-- Ortmanniana_ligamentina    
                             |  +--------+                              
                             |  |        +-- Lampsilis_cardium          
                             +--+                                       
                                |  +-------- Cambarunio_iris            
                                +--+                                    
                                   |  +----- Sphenonaias_microdon__2    
                                   +--+                                 
                                      |  +-- Cyrtonaias_explicata       
                                      +--+                              
                                         +-- Sphenonaias_microdon__1    
 
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  Page updated: 08:31:48 Sun 07 Sep 2025  |  
 
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