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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Lemiox Rafinesque, 1831: cladomics
     2 valid species

Cladomics Summary

number of species = 2

analyzed in 18 cladograms in 12 publications
maximum number of terminals in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = 0, mean = 0
proportion of cladograms that support basic monophyly = 0


Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 55 47 22 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                                   +-- Cyrtonaias             
  +--------------------------------+                          
  |                                +-- Glebula                
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus 
           |  |              |  |                             
           |  |              +--+----- Lemiox_rimosus **         
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla              
              |  |              +--+                          
              |  |                 +-- Ellipsaria             
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia                

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 35 19 --
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Medionidus             
  |                 +--+                          
  |                 |  +-- Lemiox_rimosus **         
--+  +--------------+                             
  |  |              |  +-- Cyprogenia             
  |  |              +--+                          
  |  |                 +-- Ptychobranchus         
  |  |                                            
  +--+           +-------- Potamilus              
     |  +--------+                                
     |  |        |  +----- Atlanticoncha          
     |  |        +--+                             
     |  |           |  +-- Ellipsaria             
     |  |           +--+                          
     +--+              +-- Truncilla              
        |                                         
        |              +-- Hamiota                
        |  +-----------+                          
        |  |           +-- Villosa                
        |  |                                      
        +--+  +----------- Ligumia                
           |  |                                   
           |  |     +----- Venustaconcha          
           +--+  +--+                             
              |  |  |  +-- Obovaria               
              |  |  +--+                          
              +--+     +-- Epioblasma             
                 |                                
                 |  +----- Ortmanniana_pectorosa  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina
                    +--+                          
                       +-- Lampsilis              

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 41 4 3 --
  +----- Lemiox_rimosus **
--+                    
  |  +-- Lampsilis     
  +--+                 
     +-- Obovaria      

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 42 4 3 --
  +-------- Obovaria_unicolor
--+                          
  |  +----- Obovaria_olivaria
  +--+                       
     |  +-- Lampsilis        
     +--+                    
        +-- Lemiox_rimosus **   

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox_rimosus **                
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 56 47 22 --
                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox_rimosus **         
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis 

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 11 7 --
  +----------------- Toxolasma     
  |                                
--+  +-------------- Venustaconcha 
  |  |                             
  +--+           +-- Lemiox_rimosus **
     |  +--------+                 
     |  |        +-- Lampsilis     
     +--+                          
        |  +-------- Potamilus     
        +--+                       
           |  +----- Cambarunio    
           +--+                    
              |  +-- Ellipsaria    
              +--+                 
                 +-- Truncilla     

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma          
  |                                              
--+  +----------------------- Obliquaria         
  |  |                                           
  |  |                    +-- Cyrtonaias         
  +--+  +-----------------+                      
     |  |                 +-- Glebula            
     |  |                                        
     +--+                 +-- Ptychobranchus     
        |  +--------------+                      
        |  |              +-- Lemiox_rimosus **     
        |  |                                     
        +--+              +-- Venustaconcha      
           |  +-----------+                      
           |  |           +-- Obovaria           
           +--+                                  
              |  +----------- Hamiota            
              +--+                               
                 |  +-------- Lampsilis_ovata    
                 +--+                            
                    |  +----- Ortmanniana        
                    +--+                         
                       |  +-- Lampsilis_straminea
                       +--+                      
                          +-- Lampsilis_hydiana  

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 72 14 10 --
  +-------------- Potamilus_alatus      
  |                                     
  +-------------- Potamilus_capax       
  |                                     
  +-------------- Glebula               
  |                                     
  |           +-- Toxolasma             
  +-----------+                         
--+           +-- Paetulunio            
  |                                     
  |           +-- Dromus                
  +-----------+                         
  |           +-- Lemiox_rimosus **        
  |                                     
  |           +-- Lampsilis_ovata       
  |  +--------+                         
  |  |        +-- Obliquaria            
  +--+                                  
     |  +-------- Lampsilis_virescens   
     |  |                               
     +--+     +-- Epioblasma            
        |  +--+                         
        |  |  +-- Obovaria_arkansasensis
        +--+                            
           |  +-- Obovaria_olivaria     
           +--+                         
              +-- Venustaconcha         

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox_rimosus **                
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
1 1 0 1 103 10 7 --
-- Lemiox_rimosus

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma          
  |                                              
--+  +----------------------- Obliquaria         
  |  |                                           
  |  |                    +-- Cyrtonaias         
  +--+  +-----------------+                      
     |  |                 +-- Glebula            
     |  |                                        
     +--+                 +-- Ptychobranchus     
        |  +--------------+                      
        |  |              +-- Lemiox_rimosus **     
        |  |                                     
        +--+              +-- Venustaconcha      
           |  +-----------+                      
           |  |           +-- Obovaria           
           +--+                                  
              |  +----------- Hamiota            
              +--+                               
                 |  +-------- Lampsilis_ovata    
                 +--+                            
                    |  +----- Ortmanniana        
                    +--+                         
                       |  +-- Lampsilis_straminea
                       +--+                      
                          +-- Lampsilis_hydiana  

Grobler et al., 2011 [Publ 3359], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 16 15 9 --
        +-------- Lemiox_rimosus **       
  +-----+                              
  |     |  +----- Medionidus           
  |     +--+                           
  |        |  +-- Dromus               
  |        +--+                        
--+           +-- Cyprogenia           
  |                                    
  |  +----------- Potamilus            
  |  |                                 
  |  |     +----- Lampsilis_siliquoidea
  |  |  +--+                           
  +--+  |  |  +-- Lampsilis_higginsii  
     |  |  +--+                        
     |  |     +-- Ortmanniana          
     |  |                              
     |  |  +----- Lampsilis_fasciola   
     +--+--+                           
        |  |  +-- Lampsilis_cardium    
        |  +--+                        
        |     +-- Lampsilis_ovata      
        |                              
        |     +-- Cambarunio           
        |     |                        
        +-----+-- Ligumia              
              |                        
              +-- Epioblasma           

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
  +----------------- Toxolasma            
  |                                       
  |  +-------------- Potamilus            
  |  |                                    
  |  |           +-- Ellipsaria           
  |  +-----------+                        
--+  |           +-- Truncilla            
  |  |                                    
  |  |        +----- Medionidus           
  |  +--------+                           
  |  |        |  +-- Lemiox_rimosus **       
  |  |        +--+                        
  +--+           +-- Dromus               
     |                                    
     |  +----------- Cambarunio           
     |  |                                 
     |  +----------- Leaunio              
     |  |                                 
     |  +----------- Villosa              
     |  |                                 
     |  |        +-- Sagittunio           
     |  +--------+                        
     +--+        +-- Lampsilis_ovata      
        |                                 
        |     +----- Venustaconcha        
        +-----+                           
        |     |  +-- Epioblasma           
        |     +--+                        
        |        +-- Obovaria             
        |                                 
        |  +-------- Lampsilis_teres      
        +--+                              
           |  +----- Lampsilis_siliquoidea
           +--+                           
              |  +-- Lampsilis_hydiana    
              +--+                        
                 +-- Lampsilis_straminea  

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 72 14 10 --
  +----------------- Obliquaria            
  |                                        
  |           +----- Potamilus             
--+  +--------+                            
  |  |        |  +-- Dromus                
  |  |        +--+                         
  +--+           +-- Lemiox_rimosus **        
     |                                     
     |  +----------- Lampsilis_ovata       
     |  |                                  
     +--+  +-------- Lampsilis_virescens   
        |  |                               
        +--+     +-- Epioblasma            
           |  +--+                         
           |  |  +-- Obovaria_arkansasensis
           +--+                            
              |  +-- Obovaria_olivaria     
              +--+                         
                 +-- Venustaconcha         

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 11 7 --
  +----------------- Toxolasma     
  |                                
--+  +-------------- Ellipsaria    
  |  |                             
  +--+  +----------- Truncilla     
     |  |                          
     +--+        +-- Lemiox_rimosus **
        |  +-----+                 
        |  |     +-- Potamilus     
        +--+                       
           |  +----- Lampsilis     
           +--+                    
              |  +-- Cambarunio    
              +--+                 
                 +-- Venustaconcha 

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 41 4 3 --
  +----- Lemiox_rimosus **
--+                    
  |  +-- Lampsilis     
  +--+                 
     +-- Obovaria      

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 41 4 3 --
  +----- Lemiox_rimosus **
--+                    
  |  +-- Lampsilis     
  +--+                 
     +-- Obovaria      

Page updated: 14:56:24 Wed 04 Dec 2024

 
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