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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Obovaria Rafinesque, 1819: cladomics
     7 valid species

Cladomics Summary

number of species = 7

analyzed in 27 cladograms in 15 publications
maximum number of terminals in an analysis = 5, mean = 2.15

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included terminals = 5, mean = 2.72
proportion of cladograms that support basic monophyly = 0.56


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 12 5 5 0.0

*** Top Monophyly Cladogram ***

  +----------------- Lampsilis             
  |                                        
--+  +-------------- Leaunio               
  |  |                                     
  |  |           +-- Villosa               
  +--+-----------+                         
     |           +-- Cambarunio            
     |                                     
     |  +----------- Obovaria_olivaria **     
     +--+                                  
        |  +-------- Obovaria_arkansasensis **
        +--+                               
           |  +----- Obovaria_retusa **       
           +--+                            
              |  +-- Obovaria_unicolor **     
              +--+                         
                 +-- Obovaria_subrotunda **   

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
4 10 6 4 53 44 22 9.0

*** Top Sister-Group Cladogram ***

                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria **      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor **      
                                +--+                             
                                   |  +-- Obovaria_subrotunda **    
                                   +--+                          
                                      +-- Obovaria_arkansasensis ** 

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 52 44 22 3.0
                                         +-- Cyrtonaias             
  +--------------------------------------+                          
  |                                      +-- Glebula                
  |                                                                 
--+  +-------------------------------------- Paetulunio             
  |  |                                                              
  |  |                                   +-- Obliquaria             
  +--+  +--------------------------------+                          
     |  |                                +-- Toxolasma              
     |  |                                                           
     |  |  +-------------------------------- Sagittunio             
     +--+  |                                                        
        |  |                    +----------- Potamilus              
        |  |                    |                                   
        |  |  +-----------------+        +-- Medionidus_conradicus  
        |  |  |                 |  +-----+                          
        +--+  |                 |  |     +-- Ptychobranchus         
           |  |                 +--+                                
           |  |                    |  +----- Medionidus_acutissimus 
           |  |                    |  |                             
           |  |                    +--+----- Lemiox                 
           |  |                       |                             
           |  |                       |  +-- Dromus                 
           +--+                       +--+                          
              |                          +-- Cyprogenia             
              |                                                     
              |                       +----- Lampsilis_teres        
              |  +--------------------+                             
              |  |                    |  +-- Truncilla              
              |  |                    +--+                          
              |  |                       +-- Ellipsaria             
              |  |                                                  
              +--+  +----------------------- Leaunio                
                 |  |                                               
                 |  |           +----------- Hamiota                
                 |  |           |                                   
                 |  |  +--------+        +-- Lampsilis_fasciola     
                 +--+  |        |  +-----+                          
                    |  |        |  |     +-- Ortmanniana_pectorosa  
                    |  |        +--+                                
                    |  |           |  +----- Lampsilis_ornata       
                    |  |           +--+                             
                    |  |              |  +-- Lampsilis_ovata        
                    +--+              +--+                          
                       |                 +-- Lampsilis_cardium      
                       |                                            
                       |           +-------- Lampsilis_siliquoidea  
                       |  +--------+                                
                       |  |        |  +----- Ortmanniana_ligamentina
                       |  |        +--+                             
                       |  |           |  +-- Cambarunio             
                       +--+           +--+                          
                          |              +-- Ligumia                
                          |                                         
                          |  +-------------- Venustaconcha          
                          +--+                                      
                             |  +----------- Epioblasma             
                             +--+                                   
                                |  +-------- Obovaria_olivaria **      
                                +--+                                
                                   |  +----- Obovaria_unicolor **      
                                   +--+                             
                                      |  +-- Obovaria_subrotunda **    
                                      +--+                          
                                         +-- Obovaria_arkansasensis ** 

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
4 9 6 4 98 39 22 9.0
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria **             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor **             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis **        
     |     +--+                                 
     |        +-- Obovaria_subrotunda **           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
4 9 8 4 98 39 22 9.0
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria **             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor **             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis **        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda **           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 15 14 5 4.0
  +----------------------- Obliquaria            
  |                                              
  |                    +-- Cambarunio            
  |  +-----------------+                         
  |  |                 +-- Leaunio               
  |  |                                           
--+  |        +----------- Lampsilis_teres       
  |  |        |                                  
  |  +--------+  +-------- Ortmanniana           
  |  |        |  |                               
  |  |        +--+     +-- Lampsilis_hydiana     
  |  |           |  +--+                         
  +--+           |  |  +-- Lampsilis_virescens   
     |           +--+                            
     |              |  +-- Lampsilis_siliquoidea 
     |              +--+                         
     |                 +-- Lampsilis_radiata     
     |                                           
     |              +----- Obovaria_arkansasensis **
     |  +-----------+                            
     |  |           |  +-- Obovaria_olivaria **     
     |  |           +--+                         
     +--+              +-- Obovaria_subrotunda **   
        |                                        
        |  +-------------- Lampsilis_bracteata   
        |  |                                     
        +--+  +----------- Lampsilis_ornata      
           |  |                                  
           +--+  +-------- Lampsilis_ovata       
              |  |                               
              +--+  +----- Lampsilis_cardium__3  
                 |  |                            
                 +--+  +-- Lampsilis_satura      
                    |  |                         
                    +--+-- Lampsilis_cardium__1  
                       |                         
                       +-- Lampsilis_cardium__2  

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 15 14 5 4.0
  +----------------- Lampsilis_teres       
  |                                        
  |              +-- Leaunio               
  +--------------+                         
  |              +-- Cambarunio            
  |                                        
  |           +----- Obovaria_arkansasensis **
  +-----------+                            
  |           |  +-- Obovaria_olivaria **     
  |           +--+                         
  |              +-- Obovaria_subrotunda **   
  |                                        
--+        +-------- Ortmanniana           
  |        |                               
  +--------+     +-- Lampsilis_hydiana     
  |        |  +--+                         
  |        |  |  +-- Lampsilis_virescens   
  |        +--+                            
  |           |  +-- Lampsilis_siliquoidea 
  |           +--+                         
  |              +-- Lampsilis_radiata     
  |                                        
  |  +-------------- Lampsilis_bracteata   
  |  |                                     
  +--+  +----------- Lampsilis_ornata      
     |  |                                  
     +--+  +-------- Lampsilis_ovata       
        |  |                               
        +--+  +----- Lampsilis_cardium__3  
           |  |                            
           +--+  +-- Lampsilis_satura      
              |  |                         
              +--+-- Lampsilis_cardium__1  
                 |                         
                 +-- Lampsilis_cardium__2  

Lydeard et al., 2000 [Publ 2654], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 12 1 1 --
  +----- Lampsilis        
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Lydeard et al., 2000 [Publ 2654], fig. 2c

iT tT bi riS oT roT roS iD
2 3 0 2 12 1 1 4.0
  +----- Obovaria_unicolor **
--+                       
  |  +-- Obovaria_olivaria **
  +--+                    
     +-- Lampsilis        

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 40 3 3 --
  +-------- Lemiox           
--+                          
  |  +----- Lampsilis        
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 40 3 3 --
  +-------- Lemiox           
--+                          
  |  +----- Lampsilis        
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Lydeard et al., 2000 [Publ 2654], fig. 2b

iT tT bi riS oT roT roS iD
2 2 0 2 12 1 1 --
  +----- Lampsilis        
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
2 4 0 2 41 3 3 5.0
  +-------- Obovaria_unicolor **
--+                          
  |  +----- Obovaria_olivaria **
  +--+                       
     |  +-- Lampsilis        
     +--+                    
        +-- Lemiox           

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
2 12 0 2 102 9 7 13.0
  +-- Dromus             
  |                      
  +-- Epioblasma         
  |                      
  +-- Glebula            
  |                      
  +-- Lampsilis_ovata    
  |                      
  +-- Lampsilis_virescens
  |                      
  +-- Lemiox             
--+                      
  +-- Obliquaria         
  |                      
  +-- Obovaria_olivaria **  
  |                      
  +-- Obovaria_unicolor **  
  |                      
  +-- Plectomerus        
  |                      
  +-- Potamilus_alatus   
  |                      
  +-- Potamilus_capax    

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 40 3 3 --
  +-------- Lemiox           
--+                          
  |  +----- Lampsilis        
  +--+                       
     |  +-- Obovaria_unicolor **
     +--+                    
        +-- Obovaria_olivaria **

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
2 5 0 2 71 13 10 6.0
  +-------------- Potamilus_alatus      
  |                                     
  +-------------- Potamilus_capax       
  |                                     
  +-------------- Glebula               
  |                                     
  |           +-- Toxolasma             
  +-----------+                         
--+           +-- Paetulunio            
  |                                     
  |           +-- Dromus                
  +-----------+                         
  |           +-- Lemiox                
  |                                     
  |           +-- Lampsilis_ovata       
  |  +--------+                         
  |  |        +-- Obliquaria            
  +--+                                  
     |  +-------- Lampsilis_virescens   
     |  |                               
     +--+     +-- Epioblasma            
        |  +--+                         
        |  |  +-- Obovaria_arkansasensis **
        +--+                            
           |  +-- Obovaria_olivaria **     
           +--+                         
              +-- Venustaconcha         

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
2 5 0 2 71 13 10 6.0
  +----------------- Obliquaria            
  |                                        
  |           +----- Potamilus             
--+  +--------+                            
  |  |        |  +-- Dromus                
  |  |        +--+                         
  +--+           +-- Lemiox                
     |                                     
     |  +----------- Lampsilis_ovata       
     |  |                                  
     +--+  +-------- Lampsilis_virescens   
        |  |                               
        +--+     +-- Epioblasma            
           |  +--+                         
           |  |  +-- Obovaria_arkansasensis **
           +--+                            
              |  +-- Obovaria_olivaria **     
              +--+                         
                 +-- Venustaconcha         

Lydeard et al., 2000 [Publ 2654], fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 12 1 1 --
  +----- Lampsilis        
--+                       
  |  +-- Obovaria_unicolor **
  +--+                    
     +-- Obovaria_olivaria **

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 35 19 --
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Medionidus             
  |                 +--+                          
  |                 |  +-- Lemiox                 
--+  +--------------+                             
  |  |              |  +-- Cyprogenia             
  |  |              +--+                          
  |  |                 +-- Ptychobranchus         
  |  |                                            
  +--+           +-------- Potamilus              
     |  +--------+                                
     |  |        |  +----- Atlanticoncha          
     |  |        +--+                             
     |  |           |  +-- Ellipsaria             
     |  |           +--+                          
     +--+              +-- Truncilla              
        |                                         
        |              +-- Hamiota                
        |  +-----------+                          
        |  |           +-- Villosa                
        |  |                                      
        +--+  +----------- Ligumia                
           |  |                                   
           |  |     +----- Venustaconcha          
           +--+  +--+                             
              |  |  |  +-- Obovaria_subrotunda **    
              |  |  +--+                          
              +--+     +-- Epioblasma             
                 |                                
                 |  +----- Ortmanniana_pectorosa  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina
                    +--+                          
                       +-- Lampsilis              

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 15 9 --
  +----------------- Potamilus            
  |                                       
  |              +-- Ellipsaria           
--+  +-----------+                        
  |  |           +-- Truncilla            
  |  |                                    
  +--+  +----------- Lampsilis_cardium    
     |  |                                 
     |  |     +----- Ligumia              
     |  +-----+                           
     +--+     |  +-- Obovaria_olivaria **    
        |     +--+                        
        |        +-- Epioblasma           
        |                                 
        |  +-------- Lampsilis_teres      
        +--+                              
           |  +----- Lampsilis_siliquoidea
           +--+                           
              |  +-- Ortmanniana          
              +--+                        
                 +-- Lampsilis_higginsii  

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 0 0 --
-- Obovaria_arkansasensis

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 15 9 --
  +----------- Toxolasma            
  |                                 
  |  +-------- Obliquaria           
--+  |                              
  |  |  +----- Obovaria_olivaria **    
  |  |  |                           
  |  |  +----- Lampsilis_cardium    
  +--+  |                           
     |  +----- Potamilus            
     |  |                           
     |  |  +-- Epioblasma           
     |  +--+                        
     |  |  +-- Lampsilis_siliquoidea
     +--+                           
        |  +-- Ortmanniana          
        +--+                        
        |  +-- Lampsilis_higginsii  
        |                           
        |  +-- Ligumia              
        +--+                        
        |  +-- Lampsilis_teres      
        |                           
        |  +-- Ellipsaria           
        +--+                        
           +-- Truncilla            

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma          
  |                                              
--+  +----------------------- Obliquaria         
  |  |                                           
  |  |                    +-- Cyrtonaias         
  +--+  +-----------------+                      
     |  |                 +-- Glebula            
     |  |                                        
     +--+                 +-- Ptychobranchus     
        |  +--------------+                      
        |  |              +-- Lemiox             
        |  |                                     
        +--+              +-- Venustaconcha      
           |  +-----------+                      
           |  |           +-- Obovaria_olivaria **  
           +--+                                  
              |  +----------- Hamiota            
              +--+                               
                 |  +-------- Lampsilis_ovata    
                 +--+                            
                    |  +----- Ortmanniana        
                    +--+                         
                       |  +-- Lampsilis_straminea
                       +--+                      
                          +-- Lampsilis_hydiana  

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 6 2 --
  +----- Obovaria_olivaria **
--+                       
  |  +-- Ortmanniana      
  +--+                    
     +-- Lampsilis        

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 8 3 --
           +-- Obovaria_olivaria **    
  +--------+                        
  |        +-- Ortmanniana          
  |                                 
  |        +-- Obliquaria           
  +--------+                        
--+        +-- Lampsilis_ornata     
  |                                 
  |     +----- Lampsilis_radiata__2 
  |  +--+                           
  |  |  |  +-- Lampsilis_siliquoidea
  |  |  +--+                        
  +--+     +-- Lampsilis_radiata__1 
     |                              
     |  +----- Lampsilis_virescens  
     +--+                           
        |  +-- Lampsilis_hydiana    
        +--+                        
           +-- Lampsilis_straminea  

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma          
  |                                              
--+  +----------------------- Obliquaria         
  |  |                                           
  |  |                    +-- Cyrtonaias         
  +--+  +-----------------+                      
     |  |                 +-- Glebula            
     |  |                                        
     +--+                 +-- Ptychobranchus     
        |  +--------------+                      
        |  |              +-- Lemiox             
        |  |                                     
        +--+              +-- Venustaconcha      
           |  +-----------+                      
           |  |           +-- Obovaria_olivaria **  
           +--+                                  
              |  +----------- Hamiota            
              +--+                               
                 |  +-------- Lampsilis_ovata    
                 +--+                            
                    |  +----- Ortmanniana        
                    +--+                         
                       |  +-- Lampsilis_straminea
                       +--+                      
                          +-- Lampsilis_hydiana  

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
  +----------------- Toxolasma            
  |                                       
  |  +-------------- Potamilus            
  |  |                                    
  |  |           +-- Ellipsaria           
  |  +-----------+                        
--+  |           +-- Truncilla            
  |  |                                    
  |  |        +----- Medionidus           
  |  +--------+                           
  |  |        |  +-- Lemiox               
  |  |        +--+                        
  +--+           +-- Dromus               
     |                                    
     |  +----------- Cambarunio           
     |  |                                 
     |  +----------- Leaunio              
     |  |                                 
     |  +----------- Villosa              
     |  |                                 
     |  |        +-- Sagittunio           
     |  +--------+                        
     +--+        +-- Lampsilis_ovata      
        |                                 
        |     +----- Venustaconcha        
        +-----+                           
        |     |  +-- Epioblasma           
        |     +--+                        
        |        +-- Obovaria_unicolor **    
        |                                 
        |  +-------- Lampsilis_teres      
        +--+                              
           |  +----- Lampsilis_siliquoidea
           +--+                           
              |  +-- Lampsilis_hydiana    
              +--+                        
                 +-- Lampsilis_straminea  

Page updated: 14:56:27 Wed 04 Dec 2024

 
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