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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Ellipsaria Rafinesque, 1820: cladomics
     1 valid species

Cladomics Summary

number of species = 1

analyzed in 24 cladograms in 15 publications
maximum number of terminals in an analysis = 1, mean = 1.0


Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 55 47 22 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                                   +-- Cyrtonaias             
  +--------------------------------+                          
  |                                +-- Glebula                
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus 
           |  |              |  |                             
           |  |              +--+----- Lemiox                 
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla              
              |  |              +--+                          
              |  |                 +-- Ellipsaria_lineolata **   
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia                

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria_lineolata **          
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 41 26 6 --
  +----------------------------------- Toxolasma            
  |                                                         
--+  +-------------------------------- Potamilus            
  |  |                                                      
  |  |                             +-- Ellipsaria_lineolata ** 
  +--+  +--------------------------+                        
     |  |                          +-- Truncilla            
     |  |                                                   
     |  |  +-------------------------- Leaunio              
     +--+  |                                                
        |  |     +-------------------- Lampsilis_bracteata  
        |  |     |                                          
        |  |  +--+  +----------------- Lampsilis_fasciola   
        |  |  |  |  |                                       
        +--+  |  +--+  +-------------- Lampsilis_ornata     
           |  |     |  |                                    
           |  |     +--+           +-- Lampsilis_ovata      
           |  |        |  +--------+                        
           |  |        |  |        +-- Lampsilis_cariosa    
           |  |        +--+                                 
           |  |           |  +-------- Lampsilis_cardium__3 
           +--+           +--+                              
              |              |  +----- Lampsilis_cardium__2 
              |              +--+                           
              |                 |  +-- Lampsilis_satura     
              |                 +--+                        
              |                    +-- Lampsilis_cardium__1 
              |                                             
              |                    +-- Lampsilis_higginsii  
              |     +--------------+                        
              |     |              +-- Ortmanniana          
              +-----+                                       
                    |  +-------------- Lampsilis_teres      
                    |  |                                    
                    +--+           +-- Lampsilis_radiata    
                       |  +--------+                        
                       |  |        +-- Lampsilis_siliquoidea
                       +--+                                 
                          |  +-------- Lampsilis_bergmanni  
                          +--+                              
                             |  +----- Lampsilis_hydiana    
                             +--+                           
                                |  +-- Lampsilis_virescens  
                                +--+                        
                                   +-- Lampsilis_straminea  

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 15 9 --
  +----------------- Potamilus            
  |                                       
  |              +-- Ellipsaria_lineolata ** 
--+  +-----------+                        
  |  |           +-- Truncilla            
  |  |                                    
  +--+  +----------- Lampsilis_cardium    
     |  |                                 
     |  |     +----- Ligumia              
     |  +-----+                           
     +--+     |  +-- Obovaria             
        |     +--+                        
        |        +-- Epioblasma           
        |                                 
        |  +-------- Lampsilis_teres      
        +--+                              
           |  +----- Lampsilis_siliquoidea
           +--+                           
              |  +-- Ortmanniana          
              +--+                        
                 +-- Lampsilis_higginsii  

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 8 6 --
  +-------------------- Obliquaria          
--+                                         
  |  +----------------- Medionidus          
  +--+                                      
     |  +-------------- Ptychobranchus      
     +--+                                   
        |  +----------- Potamilus_fragilis  
        +--+                                
           |  +-------- Potamilus_purpuratus
           +--+                             
              |  +----- Ellipsaria_lineolata **
              +--+                          
                 |  +-- Villosa             
                 +--+                       
                    +-- Lampsilis           

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
  +----- Ellipsaria_lineolata **
  |                          
  +----- Potamilus_leptodon  
  |                          
  +----- Medionidus          
  |                          
  +----- Potamilus_alatus    
  |                          
  +----- Ptychobranchus      
--+                          
  +----- Truncilla           
  |                          
  +----- Venustaconcha       
  |                          
  +----- Cambarunio          
  |                          
  +----- Leaunio             
  |                          
  |  +-- Ortmanniana         
  +--+                       
     +-- Lampsilis           

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +----------------- Medionidus          
--+                                      
  |  +-------------- Potamilus_fragilis  
  +--+                                   
     |  +----------- Ptychobranchus      
     +--+                                
        |  +-------- Potamilus_purpuratus
        +--+                             
           |  +----- Lampsilis           
           +--+                          
              |  +-- Villosa             
              +--+                       
                 +-- Ellipsaria_lineolata **

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +-- Villosa             
  |                       
  +-- Ellipsaria_lineolata **
  |                       
--+-- Lampsilis           
  |                       
  +-- Potamilus           
  |                       
  +-- Ptychobranchus      

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
  +-------------- Potamilus_leptodon  
  |                                   
  |  +----------- Potamilus_alatus    
--+  |                                
  |  |  +-------- Venustaconcha       
  |  |  |                             
  +--+  +-------- Leaunio             
     |  |                             
     |  |     +-- Ellipsaria_lineolata **
     |  +-----+                       
     +--+     +-- Truncilla           
        |                             
        |     +-- Medionidus          
        +-----+                       
        |     +-- Ptychobranchus      
        |                             
        |  +----- Cambarunio          
        +--+                          
           |  +-- Ortmanniana         
           +--+                       
              +-- Lampsilis           

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 13 11 --
  +-------------------- Potamilus_leptodon  
  |                                         
--+  +----------------- Potamilus_alatus    
  |  |                                      
  |  |              +-- Medionidus          
  +--+  +-----------+                       
     |  |           +-- Ptychobranchus      
     |  |                                   
     +--+           +-- Ellipsaria_lineolata **
        |  +--------+                       
        |  |        +-- Truncilla           
        +--+                                
           |  +-------- Leaunio             
           |  |                             
           +--+     +-- Venustaconcha       
              |  +--+                       
              |  |  +-- Cambarunio          
              +--+                          
                 |  +-- Ortmanniana         
                 +--+                       
                    +-- Lampsilis           

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 40 25 6 --
  +----------------------------------- Toxolasma            
  |                                                         
--+  +-------------------------------- Ellipsaria_lineolata ** 
  |  |                                                      
  +--+  +----------------------------- Truncilla            
     |  |                                                   
     |  |  +-------------------------- Potamilus            
     +--+  |                                                
        |  |  +----------------------- Leaunio              
        |  |  |                                             
        +--+  |     +----------------- Lampsilis_bracteata  
           |  |     |                                       
           |  |  +--+              +-- Lampsilis_ornata     
           |  |  |  |  +-----------+                        
           +--+  |  |  |           +-- Lampsilis_fasciola   
              |  |  +--+                                    
              |  |     |  +----------- Lampsilis_ovata      
              |  |     +--+                                 
              |  |        |  +-------- Lampsilis_cardium__3 
              |  |        +--+                              
              +--+           |  +----- Lampsilis_cardium__2 
                 |           +--+                           
                 |              |  +-- Lampsilis_satura     
                 |              +--+                        
                 |                 +-- Lampsilis_cardium__1 
                 |                                          
                 |     +-------------- Lampsilis_teres      
                 |     |                                    
                 |     |           +-- Lampsilis_higginsii  
                 +-----+  +--------+                        
                       |  |        +-- Ortmanniana          
                       |  |                                 
                       +--+        +-- Lampsilis_radiata    
                          |  +-----+                        
                          |  |     +-- Lampsilis_siliquoidea
                          |  |                              
                          +--+     +-- Lampsilis_hydiana    
                             |  +--+                        
                             |  |  +-- Lampsilis_bergmanni  
                             +--+                           
                                |  +-- Lampsilis_virescens  
                                +--+                        
                                   +-- Lampsilis_straminea  

Szumowski et al., 2012 [Publ 3569], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 14 9 --
  +----------------------- Ellipsaria_lineolata ** 
  |                                             
--+  +-------------------- Truncilla            
  |  |                                          
  +--+  +----------------- Obliquaria           
     |  |                                       
     |  |  +-------------- Potamilus            
     +--+  |                                    
        |  |        +----- Lampsilis_ornata     
        |  |  +-----+                           
        |  |  |     |  +-- Lampsilis_cardium    
        +--+  |     +--+                        
           |  |        +-- Lampsilis_ovata      
           |  |                                 
           |  |  +-------- Lampsilis_teres      
           |  +--+                              
           +--+  |  +----- Lampsilis_siliquoidea
              |  +--+                           
              |     |  +-- Ortmanniana          
              |     +--+                        
              |        +-- Lampsilis_higginsii  
              |                                 
              |     +----- Hamiota              
              |     |                           
              +-----+  +-- Epioblasma           
                    |  |                        
                    +--+-- Cambarunio           
                       |                        
                       +-- Ligumia              

Smith et al., 2020 [Publ 4129], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 27 19 7 --
  +-------------- Obliquaria          
  |                                   
--+  +----------- Cyrtonaias          
  |  |                                
  |  |        +-- Pachynaias          
  +--+  +-----+                       
     |  |     +-- Lampsilis           
     |  |                             
     +--+     +-- Ellipsaria_lineolata **
        |  +--+                       
        |  |  +-- Truncilla           
        +--+                          
           |  +-- Atlanticoncha       
           +--+                       
              +-- Potamilus           

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 35 19 --
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Medionidus             
  |                 +--+                          
  |                 |  +-- Lemiox                 
--+  +--------------+                             
  |  |              |  +-- Cyprogenia             
  |  |              +--+                          
  |  |                 +-- Ptychobranchus         
  |  |                                            
  +--+           +-------- Potamilus              
     |  +--------+                                
     |  |        |  +----- Atlanticoncha          
     |  |        +--+                             
     |  |           |  +-- Ellipsaria_lineolata **   
     |  |           +--+                          
     +--+              +-- Truncilla              
        |                                         
        |              +-- Hamiota                
        |  +-----------+                          
        |  |           +-- Villosa                
        |  |                                      
        +--+  +----------- Ligumia                
           |  |                                   
           |  |     +----- Venustaconcha          
           +--+  +--+                             
              |  |  |  +-- Obovaria               
              |  |  +--+                          
              +--+     +-- Epioblasma             
                 |                                
                 |  +----- Ortmanniana_pectorosa  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina
                    +--+                          
                       +-- Lampsilis              

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 56 47 22 --
                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria_lineolata **   
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis 

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 11 7 --
  +----------------- Toxolasma           
  |                                      
--+  +-------------- Ellipsaria_lineolata **
  |  |                                   
  +--+  +----------- Truncilla           
     |  |                                
     +--+        +-- Lemiox              
        |  +-----+                       
        |  |     +-- Potamilus           
        +--+                             
           |  +----- Lampsilis           
           +--+                          
              |  +-- Cambarunio          
              +--+                       
                 +-- Venustaconcha       

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 15 9 --
  +----------- Toxolasma            
  |                                 
  |  +-------- Obliquaria           
--+  |                              
  |  |  +----- Obovaria             
  |  |  |                           
  |  |  +----- Lampsilis_cardium    
  +--+  |                           
     |  +----- Potamilus            
     |  |                           
     |  |  +-- Epioblasma           
     |  +--+                        
     |  |  +-- Lampsilis_siliquoidea
     +--+                           
        |  +-- Ortmanniana          
        +--+                        
        |  +-- Lampsilis_higginsii  
        |                           
        |  +-- Ligumia              
        +--+                        
        |  +-- Lampsilis_teres      
        |                           
        |  +-- Ellipsaria_lineolata ** 
        +--+                        
           +-- Truncilla            

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 8 6 --
  +-------------- Potamilus_fragilis  
  |                                   
--+  +----------- Ptychobranchus      
  |  |                                
  +--+  +-------- Medionidus          
     |  |                             
     +--+  +----- Potamilus_purpuratus
        |  |                          
        +--+----- Lampsilis           
           |                          
           |  +-- Ellipsaria_lineolata **
           +--+                       
              +-- Villosa             

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 32 10 7 --
           +----- Ptychobranchus       
  +--------+                           
  |        |  +-- Potamilus_fragilis   
  |        +--+                        
  |           +-- Medionidus           
--+                                    
  |           +-- Villosa              
  |  +--------+                        
  |  |        +-- Ellipsaria_lineolata ** 
  +--+                                 
     |  +-------- Lampsilis_teres      
     +--+                              
        |  +----- Lampsilis_siliquoidea
        +--+                           
           |  +-- Potamilus_purpuratus 
           +--+                        
              +-- Potamilus_alatus     

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
  +----------------- Toxolasma            
  |                                       
  |  +-------------- Potamilus            
  |  |                                    
  |  |           +-- Ellipsaria_lineolata ** 
  |  +-----------+                        
--+  |           +-- Truncilla            
  |  |                                    
  |  |        +----- Medionidus           
  |  +--------+                           
  |  |        |  +-- Lemiox               
  |  |        +--+                        
  +--+           +-- Dromus               
     |                                    
     |  +----------- Cambarunio           
     |  |                                 
     |  +----------- Leaunio              
     |  |                                 
     |  +----------- Villosa              
     |  |                                 
     |  |        +-- Sagittunio           
     |  +--------+                        
     +--+        +-- Lampsilis_ovata      
        |                                 
        |     +----- Venustaconcha        
        +-----+                           
        |     |  +-- Epioblasma           
        |     +--+                        
        |        +-- Obovaria             
        |                                 
        |  +-------- Lampsilis_teres      
        +--+                              
           |  +----- Lampsilis_siliquoidea
           +--+                           
              |  +-- Lampsilis_hydiana    
              +--+                        
                 +-- Lampsilis_straminea  

Saxon et al., 2023 [Publ 4338], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 4 4 1 --
  +-- Ellipsaria_lineolata **
--+                       
  +-- Truncilla           

Smith et al., 2020 [Publ 4129], fig. 2-3

iT tT bi riS oT roT roS iD
1 1 0 1 27 19 7 --
  +-------------- Obliquaria          
  |                                   
--+  +----------- Cyrtonaias          
  |  |                                
  |  |        +-- Pachynaias          
  +--+  +-----+                       
     |  |     +-- Lampsilis           
     |  |                             
     +--+     +-- Ellipsaria_lineolata **
        |  +--+                       
        |  |  +-- Truncilla           
        +--+                          
           |  +-- Atlanticoncha       
           +--+                       
              +-- Potamilus           

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria_lineolata **          
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 11 7 --
  +----------------- Toxolasma           
  |                                      
--+  +-------------- Venustaconcha       
  |  |                                   
  +--+           +-- Lemiox              
     |  +--------+                       
     |  |        +-- Lampsilis           
     +--+                                
        |  +-------- Potamilus           
        +--+                             
           |  +----- Cambarunio          
           +--+                          
              |  +-- Ellipsaria_lineolata **
              +--+                       
                 +-- Truncilla           

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