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Page last updated
Wed 27 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Huang et al. (2019), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Huang, X.-C., J.-H. Su, J.-X. Ouyang, S. Ouyang, C.-H. Zhou & X.-P. Wu. 2019. Towards a global phylogeny of freshwater mussels (Bivalivia: Unionida): Species delimitation of Chinese taxa, mitochondrial phylogenomics, and diversification patterns. Molecular Phylogenetics and Evolution 130: 45-59.

Click here for online access.

Cladogram Figures

(click the figure number to see the cladogram)

fig. 1
fig. 2
fig. 3
fig. 3-M
fig. S1
fig. S3
fig. S4

Attributed Genus Names

Parvasolenaia Huang et al., 2019
Huang & Wu in Huang et al., 2019, MPE 130: 57.
Type species: Mycetopus rivularis Heude, 1877 (orig. desig.) = Parvasolenaia rivularis

Attributed Species Names

Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000, J. Moll. Studies 66: 160.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009, Res. J. BioTech. 4(1): 43.
Whelan et al., 2011, MPE 61: 506.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Wu et al., 2023, ZooKeys 1166: 263, Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
(misidentified reference to Symphynota compressa Lea, 1829)

Pseudodon vondembuschianus Graf, 2002
‘Lea’ of Authors.
Graf, 2002, J. Moll. Studies 68: 66.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Whelan et al., 2011, MPE 61: 506. [in part]
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
(misidentified reference to Unio crassus Retzius, 1788)

Anodontites elongata Whelan et al., 2011
‘Swainson’ of Authors.
Whelan et al., 2011, MPE 61: 506.
Graf et al., 2015, MPE 85: 3.
Bolotov et al., 2016, MPE 103: 108.
Lopes-Lima et al., 2017, MPE 106: 178.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 208.
Combosch et al., 2017, MPE 107: 195.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2018, MPE 120: 302.
Lopes-Lima et al., 2018, MPE 127: 103.
Huang et al., 2019, MPE 130: Table S4.
Bolotov et al., 2019, Sci. Repts. 9: Table S1.
Pfeiffer et al., 2019, MPE 137: 116.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): 3.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Sano et al., 2020, Plankton Benthos Res. 15(2): 101.
Wu et al., 2020, Invert. Syst. 34: 195.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Anodon elongatus Swainson, 1823)

Radiatula bonneaudii Whelan et al., 2011
‘Eydoux’ of Authors.
Whelan et al., 2011, MPE 61: 506.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): [4].
Bolotov et al., 2017, Sci. Repts. 7: 10, Table 3 [as ‘sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: [5].
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Pfeiffer et al., 2018, J. Moll. Studies 84(4): [4]. [in part]
Jeratthitikul et al., 2019, Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Radiatula bonneaudii sp. 1’].
Huang et al., 2019, MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020, MPE 148: Table S1.
(misidentified reference to Unio bonneaudii Eydoux, 1838)

Pseudodon cambodjensis Pfeiffer & Graf, 2013
‘Petit’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016, Sci. Total Environ. 571: 1071.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017, MPE 106: 177.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018, Conserv. Genet. Resour. 11: 459.
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021, Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021, Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023, Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
(misidentified reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.

Pseudodon inoscularis Pfeiffer & Graf, 2013
‘Gould’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. inoscularis’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Huang et al., 2019, MPE 130: Table S4 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
(misidentified reference to Anodon inoscularis Gould, 1844)

Pilsbryoconcha exilis Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016, PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

Ensidens sagittarius Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019, MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Contradens contradens Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Contradens aff. contradens sp. 1’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Contradens aff. contradens sp. 1’].
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘Lens aff. contradens’].
(misidentified reference to Unio contradens Lea, 1838)

Pseudodon cumingii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’].
Huang et al., 2019, MPE 130: Table S4 [as ‘”Bineurus” cumingii’].
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Anodonta cumingii Lea, 1851)
Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.

Pseudodon mouhotii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 3’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 3’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 3’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Pseudodon vondembuschianus Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 2’]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 2’]. [in part]
(reference to Margaritana vondembuschiana Lea, 1840)

Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
(misidentified reference to Unio hainesianus Lea, 1856)

Sinanodonta woodiana Bolotov et al., 2016
‘Lea’ of Authors.
Bolotov et al., 2016, Biochem. Syst. & Ecol. 67: 60, Table 1 [as ‘lineage D’]. [in part]
Vikhrev et al., 2017, Trop. Conserv. Sci. 10: 4, Table 1 [as ‘lineage D’]. [in part]
Bespalaya et al., 2018, Limnologica 69: 97, Table 1 [as ‘lineage D’]. [in part]
Konečný et al., 2018, Evol. Appl. 11: Table S1 [as ‘lineage H’]. [in part]
Huang et al., 2019, MPE 130: Table S1. [in part]
(misidentified reference to Symphynota woodiana Lea, 1834)

Trapezoideus subclathratus Bolotov et al., 2017
‘Martens’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Huang et al., 2019, MPE 130: Table S4.
(misidentified reference to Unio misellus var. subclathratus Martens, 1899)

Lamellidens marginalis Bolotov et al., 2017
‘Lamarck’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 4’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Lamellidens marginalis sp. 4’].
(misidentified reference to Unio marginalis Lamarck, 1819)

Lamellidens marginalis Bolotov et al., 2017
‘Lamarck’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207. [in part]
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Lamellidens marginalis sp. 3’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Lamellidens marginalis sp. 2’ and ‘sp. 3’].
Jeratthitikul et al., 2019, Tropical Nat. Hist. 19(1): 25.
Konopleva et al., 2020, Ecologica Montenegrina 27: 47.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
(misidentified reference to Unio marginalis Lamarck, 1819)

Indonaia khadakvaslaensis Bolotov et al., 2017
‘Ray’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Indonaia khadakvaslaensis sp. 4’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Indonaia khadakvaslaensis sp. 4’].
(misidentified reference to Indonaia khadakvaslaensis Ray, 1966)

Lamellidens generosus Bolotov et al., 2017
‘Gould’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
(misidentified reference to Unio generosus Gould, 1847)

Parreysia corrugata Bolotov et al., 2017
‘Müller’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 3’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1. [in part]
Huang et al., 2019, MPE 130: Table S4 [as ‘Parreysia corrugata sp. 3’].
Imsong & Murali, 2023, Uttar Pradesh J. Zool. 44(20): Table 1.
(misidentified reference to Mya corrugata Müller, 1774)

Radiatula humilis Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula humilis sp. 3].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Radiatula humilis sp. 3].
Huang et al., 2019, MPE 130: Table S4 [as ‘Radiatula humilis sp. 3’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Radiatula humilis sp. 3’].
Ortiz-Sepulveda et al., 2020, MPE 148: Table S1. [in part]
(misidentified reference to Unio humilis Lea, 1856)

Parreysia corrugata Bolotov et al., 2017
‘Müller’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 2’]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1. [in part]
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Parreysia corrygata sp. 2’].
Pfeiffer et al., 2018, J. Moll. Studies 84(4): Table 1. [in part]
Huang et al., 2019, MPE 130: Table S4 [as ‘Parreysia corrugata sp. 2’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 3’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 2’].
Imsong & Murali, 2023, Uttar Pradesh J. Zool. 44(20): Table 1.
(misidentified reference to Mya corrugata Müller, 1774)

Parreysia corrugata Bolotov et al., 2017
‘Müller’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Parreysia corrugata sp. 1’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Parreysia corrugata sp. 1’].
Pfeiffer et al., 2018, J. Moll. Studies 84(4): Table 1. [in part]
Huang et al., 2019, MPE 130: Table S4 [as ‘Parreysia corrugata sp. 1’].
Jeratthitikul et al., 2019, Tropical Nat. Hist. 19(1): 25.
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Parreysia corrugata sp. 1’].
(misidentified reference to Mya corrugata Müller, 1774)

Radiatula khadakvaslaensis Bolotov et al., 2017
‘Ray’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula khadakvaslaensis sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Indonaia khadakvaslaensis sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Indonaia khadakvaslaensis sp. 1’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Indonaia khadakvaslaensis’].
Bolotov et al., 2022, Hindawi 6600359: Table S1 [as ‘Indonaia khadakvaslaensis’].
(reference to Indonaia khadakvaslaensis Ray, 1966)

Radiatula khadakvaslaensis Bolotov et al., 2017
‘Ray’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula khadakvaslaensis sp. 2’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Indonaia khadakvaslaensis sp. 2’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Indonaia khadakvaslaensis sp. 2’].
(misidentified reference to Indonaia khadakvaslaensis Ray, 1966)

Radiatula khadakvaslaensis Bolotov et al., 2017
‘Ray’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula khadakvaslaensis sp. 3’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula khadakvaslaensis sp. 3’].
Pfeiffer et al., 2018, J. Moll. Studies 84(4): [4].
Huang et al., 2019, MPE 130: Table S4 [as ‘Radiatula khadakvaslaensis sp. 3’].
(misidentified reference to Indonaia khadakvaslaensis Ray, 1966)

Ensidens sagittarius Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 2’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 2’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘E. aff. sagittarius sp. 2’].
Muanta et al., 2019, J. Moll. Studies 85(2): 227 [as ‘Clade B4’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 2’].
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1 [as ‘E. aff. sagittarius sp. 2’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 2’].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2242 [as ‘Ensidens aff. sagittarius sp. 2’].
(misidentified reference to Unio sagittarius Lea, 1856)

Pseudodon mouhotii Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. mouhotii sp. 2’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Bineurus aff. mouhotii sp. 2’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Bineurus aff. mouhotii sp. 2’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Bineurus aff. mouhotii sp. 2’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Bineurus aff. mouhotii sp. 2’].
(misidentified reference to Monocondyloea mouhotii Lea, 1863)

Pseudodon vondembuschianus Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
(misidentified reference to Margaritana vondembuschiana Lea, 1840)

Pseudodon inoscularis Bolotov et al., 2017
‘Gould’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. inoscularis sp. 1].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘ “Monodontina” aff. inoscularis sp. 1].
Huang et al., 2019, MPE 130: Table S4 [as ‘ “Monodontina” aff. inoscularis sp. 1].
(misidentified reference to Anodon inoscularis Gould, 1844)

Pseudodon avae Bolotov et al., 2017
‘Theobald’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. avae’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1 [as ‘Pseudodon cf. avae’].
Huang et al., 2019, MPE 130: Table S4.
Konopleva et al., 2019, Sci. Repts. 9: Table S1.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Dai et al., 2023, Invert. Syst. 37(2): 155, Table 1.
(misidentified reference to Monocondylaea avae Theobald, 1873)

Indonaia lima Bolotov et al., 2017
‘Simpson’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Konopleva et al., 2019, Sci. Repts. 9: Table S1.
Jeratthitikul et al., 2019, Tropical Nat. Hist. 19(1): 25.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Bolotov et al., 2022, Hindawi 6600359: Table S1.
(misidentified reference to Nodularia (sec. Radiatula) lima Simpson, 1900)

Nodularia douglasiae Liu et al., 2017
‘Griffith & Pidgeon’ of Authors.
Liu et al., 2017, PLoS One 12(12 ), e0189737: 2. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57. [in part]
Huang et al., 2019, MPE 130: Tables S1, S4 [as ‘Nodularia cf. douglasiae’]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: Table 1. [in part]
(misidentified reference to Unio douglasiae Griffith & Pidgeon, 1833)

Parreysia burmana Bolotov et al., 2017
‘Blanford’ of Authors.
‘Blanford’ Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Radiatula burmana’].
‘Blanford’ Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Ortiz-Sepulveda et al., 2020, MPE 148: Table S1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio burmanus Blanford, 1869)

Solenaia triangularis Wu et al., 2018
‘Heude’ of Authors.
Wu et al., 2018, PLoS One 13(8), e0200956: 7. [in part]
Huang et al., 2019, MPE 130: Table S4.
(misidentified reference to Mycetopus triangularis Heude, 1885)

Anemina globosula Wu et al., 2018
‘Heude’ of Authors.
Wu et al., 2018, PLoS One 13(8), e0200956: 4.
Huang et al., 2019, MPE 130: Table S1.
(misidentified reference to Anodon globosula Heude, 1878)

Sinanodonta pacifica Huang et al., 2019
Huang et al., 2019, MPE 130: Table S1. [in part]
(misidentified reference to Anodon pacifica Heude, 1878)

Pleurobema sintoxia Huang et al., 2019
‘Rafinesque’ Huang et al., 2019, MPE 130: Table S4. [in part]
(misidentified reference to Obliquaria (Sintoxia) sintoxia Rafinesque, 1820)

Solenaia oleivora Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S4 [as ‘Solenaia oleivora sp. 2’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Solenaia cf. oleivora 3’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 2’].
(misidentified reference to Mycetopus oleivorus Heude, 1877)

Solenaia oleivora Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S4 [as ‘Solenaia oleivora SDCFM189’].
Chen et al., 2020, mtDNA B 5(3): 2800 [as ‘Solenaia oleivora MG’]. [in part]
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Solenaia cf. oleivora 1’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 1’].
Sano et al., 2020, Plankton Benthos Res. 15(2): 101.
Bolotov et al., 2022, Hindawi 6600359: Table S1.
(misidentified reference to Mycetopus oleivorus Heude, 1877)

Sinanodonta elliptica Huang et al., 2019
Huang et al., 2019, MPE 130: Table S1. [in part]
(misidentified reference to Anodon elliptica Heude, 1878)

Sinanodonta pacifica Huang et al., 2019
Huang et al., 2019, MPE 130: Table S1. [in part]
(misidentified reference to Anodon pacifica Heude, 1878)

Sinanodonta joreti Huang et al., 2019
Huang et al., 2019, MPE 130: Table S1.
(misidentified reference to Anodon joreti Heude, 1878)

Anemina euscaphys Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S1. [in part]
Whelan et al., 2023, Zool. J. Linn. Soc. 198(2): Table S1. [in part]
(misidentified reference to Anodon euscaphys Heude, 1879)

Taxonomic Opinions

Actinonaias ligamentina (Lamarck, 1819)

Aculamprotula coreana (Martens, 1886)

Aculamprotula fibrosa (Heude, 1877)

Aculamprotula grandidens (Lea, 1862)

Aculamprotula nodulosa (Wood, 1815)

Aculamprotula polysticta (Heude, 1877)

Aculamprotula scripta (Heude, 1875)

Aculamprotula tientsinensis (Crosse & Debeaux, 1863)

Aculamprotula tortuosa (Lea, 1865)

Aculamprotula zonata (Heude, 1883)

Acuticosta chinensis (Lea, 1868)

Acuticosta ovata

Alasmidonta marginata Say, 1818

Alathyria jacksoni Iredale, 1934

Alathyria pertexta Iredale, 1934

Alathyria profuga (Gould, 1850)

Amblema plicata (Say, 1817)

Anemina arcaeformis (Heude, 1877)

Anemina euscaphys

Anemina globosula

Anemina sp.

Anodonta anatina (Linnaeus, 1758)

Anodonta cygnea (Linnaeus, 1758)

Anodonta nuttalliana Lea, 1838

Anodontites elongata (Swainson, 1823)

Anodontites trapesialis (Lamarck, 1819)

Aspatharia pfeifferiana (Bernardi, 1860)

Bineurus aff. mouhotii sp. 3 (Lea, 1863)

Bineurus cumingii (Lea, 1850)

Bineurus ellipticus (Conrad, 1865)

Castalia ambigua Lamarck, 1819

Chambardia wahlbergi (Krauss, 1848)

Chamberlainia hainesiana (Lea, 1856)

Coelatura aff. aegyptiaca sp. 2 (Cailliaud, 1827)

Coelatura gabonensis (Küster, 1862)

Coelatura kunenensis (Mousson, 1887)

Coelatura mweruensis (Smith, 1908)

Contradens aff. contradens sp. 1 (Lea, 1838)

Contradens eximius (Lea, 1856)

Cristaria plicata (Leach, 1814)

Cuneopsis celtiformis (Heude, 1874)

Cuneopsis heudei (Heude, 1874)

Cuneopsis pisciculus (Heude, 1874)

Cuneopsis rufescens (Heude, 1874)

Diplodon suavidicus (Lea, 1856)

Echyridella menziesii (Gray, 1843)

Ellipsaria lineolata (Rafinesque, 1820)

Elliptio complanata (Lightfoot, 1786)

Elliptio dilatata (Rafinesque, 1820)

Elliptoideus sloatianus (Lea, 1840)

Ensidens aff. sagittarius sp. 3 (Lea, 1856)

Ensidens cf. ingallsianus (Lea, 1852)

Etheria elliptica Lamarck, 1807

Fusconaia flava (Rafinesque, 1820)

Gonidea angulata (Lea, 1838)

Hyridella australis (Lamarck, 1819)

Hyridella depressa (Lamarck, 1819)

Hyridella drapeta (Iredale, 1934)

Hyriopsis bialata Simpson, 1900

Hyriopsis desowitzi Brandt, 1974

Hyriopsis myersiana (Lea, 1856)

Hyriopsis sp. 1

Hyriopsis sp. 2

Indonaia aff. khadakvaslaensis sp. 4 (Ray, 1966)

Indonaia andersoniana (Nevill, 1877)

Indonaia caerulea (Lea, 1831)

Indonaia lima (Simpson, 1900)

Lamellidens aff. marginalis sp. 5 (Lamarck, 1819)

Lamellidens brandti

Lamellidens cf. corrianus (Lea, 1834)

Lamellidens cf. jenkinsianus (Benson, 1862)

Lamellidens generosus (Gould, 1847)

Lamellidens savadiensis (Nevill, 1877)

Lamproscapha ensiformis (Spix & Wagner, 1827)

Lamprotula caveata (Heude, 1877)

Lamprotula cornuumlunae (Heude, 1883)

Lamprotula gottschei

Lamprotula leaii (Griffith & Pidgeon, 1833)

Lamprotula “scripta mitogenome”

Lampsilis cardium Rafinesque, 1820

Lampsilis ornata (Conrad, 1835)

Lanceolaria eucylindrica

Lanceolaria gladiola (Heude, 1877)

Lanceolaria grayana (Lea, 1834)

Lanceolaria grayii (Griffith & Pidgeon, 1833)

Lanceolaria lanceolata (Lea, 1857)
+ Triquetra contorta Lea, 1857
+ Unio contortus var. muticus Heude, 1877
+ Unio contortus var. conjungens Heude, 1877

Lanceolaria triformis (Heude, 1877)

Lasmigona compressa (Lea, 1829)

Leguminaia wheatleyi (Lea, 1862)

Leoparreysia bhamoensis (Theobald, 1873)

Leoparreysia canefrii

Leoparreysia olivacea (Prashad, 1930)

Leoparreysia tavoyensis (Gould, 1843)

Lepidodesma aligera (Heude, 1877)

Lepidodesma languilati (Heude, 1874)

Leptodea leptodon (Rafinesque, 1820)

Lortiella froggatti Iredale, 1934

Margaritifera auricularia (Spengler, 1793)

Margaritifera dahurica (Middendorff, 1850)

Margaritifera falcata (Gould, 1850)

Margaritifera hembeli (Conrad, 1838)

Margaritifera homsensis (Lea, 1865)

Margaritifera laevis (Haas, 1910)

Margaritifera laosensis (Lea, 1863)

Margaritifera margaritifera (Linnaeus, 1758)

Margaritifera marocana (Pallary, 1918)

Margaritifera marrianae Johnson, 1983

Margaritifera middendorffi (Rosén, 1926)

Margaritifera monodonta (Say, 1829)

Margaritifera rochechouartii (Heude, 1875)

Medionidus conradicus (Lea, 1834)

Megalonaias nervosa (Rafinesque, 1820)

Microcondylaea bonellii (A. Férussac, 1827)

Microdontia anodontaeformis (Tapparone Canefri, 1883)

Monodontina aff. inoscularis sp. 2

Monodontina aff. vondembuschiana sp. 2 (Lea, 1840)

Monodontina cambodjensis (Petit de la Saussaye, 1865)

Mutela dubia (Gmelin, 1791)

Mutela hargeri E.A. Smith, 1908

Mutela rostrata (Rang, 1835)

Mycetopoda siliquosa (Spix & Wagner, 1827)

Neotrigonia lamarckii (Gray, 1838)

Neotrigonia margaritacea (Lamarck, 1804)

Nitia teretiuscula (Philippi, 1847)

Nodularia cf. douglasiae (Griffith & Pidgeon, 1833)

Nodularia nuxpersicae Dunker, 1848

Obliquaria reflexa Rafinesque, 1820

Oxynaia pugio (Benson, 1862)

Oxynaia sp. ‘Taungoo’

Oxynaia sp. ‘Tavoy’

Parreysia cf. corrugata sp. 3 (Müller, 1774)

Parvasolenaia neotriangularis (He & Zhuang, 2013)

Parvasolenaia rivularis (Heude, 1877)

Parvasolenaia triangularis (Heude, 1885)

Physunio modelli Brandt, 1974

Physunio superbus (Lea, 1843)

Pilsbryoconcha aff. compressa sp. 1 (Martens, 1860)

Pilsbryoconcha aff. exilis sp. 2 (Lea, 1838)

Plectomerus dombeyanus (Humboldt & Bonpland, 1827)

Pletholophus tenuis (Griffith & Pidgeon, 1833)

Pleurobema sintoxia (Rafinesque, 1820)

Potamilus alatus (Say, 1817)

Potomida littoralis (Cuvier, 1798)

Pronodularia japanensis (Lea, 1859)

Pseudanodonta complanata (Rossmässler, 1835)

Pseudodon avae (Theobald, 1873)

Pseudodon bogani

Pseudodon manueli

Ptychobranchus subtentum (Say, 1825)

Ptychorhynchus pfisteri (Heude, 1874)

Pyganodon grandis (Say, 1829)

Quadrula quadrula (Rafinesque, 1820)

Quadrula verrucosa (Rafinesque, 1820)

Radiatula aff. humilis sp. 3 (Lea, 1856)

Radiatula burmana (Blanford, 1869)

Radiatula cf. bonneaudii sp. 1 (Eydoux, 1838)

Radiatula mouhoti

Rectidens sumatrensis (Dunker, 1852)

Schistodesmus cf. lampreyanus

Schistodesmus sp. ‘Lamprotula gottschei’

Schistodesmus spinosus (Simpson, 1900)

Simpsonaias ambigua (Say, 1825)

Sinanodonta angula (Tchang, Li et Liu, 1965)

Sinanodonta elliptica

Sinanodonta joreti

Sinanodonta lucida (Heude, 1877)

Sinanodonta pacifica

Sinanodonta woodiana
+ Sinanodonta woodiana Bolotov et al., 2016

Sinohyriopsis cumingii (Lea, 1852)

Sinohyriopsis schlegelii (Martens, 1861)

Solenaia carinata (Heude, 1877)

Solenaia oleivora (Heude, 1877)

Strophitus undulatus (Say, 1817)

Toxolasma parvum (Barnes, 1823)

Trapezidens exolescens (Gould, 1843)

Trapezidens obesa fea

Trapezidens obesa obesa (Hanley & Theobald, 1876)

Trapezoideus nesemanni

Trapezoideus panhai

Trapezoideus subclathratus (Martens, 1899)

Triplodon corrugatus (Lamarck, 1819)

Truncilla truncata Rafinesque, 1820

Unio crassus Philipsson, 1788

Unio delphinus Spengler, 1793

Unio pictorum (Linnaeus, 1758)

Unio tumidus Philipsson, 1788

Uniomerus carolinianus (Bosc, 1801)

Uniomerus tetralasmus

Utterbackia imbecillis (Say, 1829)

Utterbackia peninsularis Bogan & Hoeh, 1995

Velesunio ambiguus (Philippi, 1847)

Venustaconcha ellipsiformis (Conrad, 1836)

Villosa iris (Lea, 1829)

Villosa lienosa (Conrad, 1834)

Westralunio carteri Iredale, 1934

Cladogram Figure 1
                                         +-- Margaritifera_dahurica     
  +--------------------------------------+                              
  |                                      +-- Gibbosula_rochechouartii   
  |                                                                     
  |                                   +----- Lamprotula_leaii           
  |                                   |                                 
  |                       +-----------+  +-- Lamprotula_caveata__1      
  |                       |           |  |                              
  |                       |           +--+-- Lamprotula_cornuumlunae    
  |                       |              |                              
--+  +--------------------+              +-- Lamprotula_caveata__2      
  |  |                    |                                             
  |  |                    |              +-- Parvasolenaia_rivularis    
  |  |                    |  +-----------+                              
  |  |                    |  |           +-- Parvasolenaia_triangularis 
  |  |                    +--+                                          
  |  |                       |  +----------- Ptychorhynchus_pfisteri    
  |  |                       +--+                                       
  |  |                          |  +-------- Sinosolenaia_carinata      
  |  |                          +--+                                    
  +--+                             |  +----- Sinosolenaia_recognita     
     |                             +--+                                 
     |                                |  +-- Sinosolenaia_oleivora      
     |                                +--+                              
     |                                   +-- Sinosolenaia_iridinea      
     |                                                                  
     |                                +----- Sinohyriopsis_cumingii__1  
     |  +-----------------------------+                                 
     |  |                             |  +-- Sinohyriopsis_cumingii__2  
     |  |                             +--+                              
     |  |                                +-- Sinohyriopsis_schlegelii   
     +--+                                                               
        |  +-------------------------------- Anemina_arcaeformis        
        |  |                                                            
        |  |  +----------------------------- Acuticosta_chinensis       
        |  |  |                                                         
        +--+  |                          +-- Sinanodonta_angula         
           |  |                 +--------+                              
           |  |                 |        +-- Sinanodonta_lucida         
           |  |  +--------------+                                       
           |  |  |              |  +-------- Sinanodonta_jourdyi        
           +--+  |              +--+                                    
              |  |                 |  +----- Sinanodonta_woodiana       
              |  |                 +--+                                 
              |  |                    |  +-- Sinanodonta_elliptica      
              |  |                    +--+                              
              |  |                       +-- Sinanodonta_pacifica       
              |  |                                                      
              +--+              +----------- Aculamprotula_tortuosa     
                 |  +-----------+                                       
                 |  |           |  +-------- Aculamprotula_scripta      
                 |  |           +--+                                    
                 |  |              |  +----- Aculamprotula_coreana      
                 |  |              +--+                                 
                 |  |                 |  +-- Aculamprotula_fibrosa      
                 |  |                 +--+                              
                 |  |                    +-- Aculamprotula_tientsinensis
                 +--+                                                   
                    |           +----------- Cristaria_plicata          
                    |  +--------+                                       
                    |  |        |  +-------- Lanceolaria_gladiola       
                    |  |        +--+                                    
                    |  |           |  +----- Lanceolaria_grayii         
                    |  |           +--+                                 
                    |  |              |  +-- Lanceolaria_lanceolata     
                    +--+              +--+                              
                       |                 +-- Lanceolaria_triformis      
                       |                                                
                       |  +----------------- Lepidodesma_languilati     
                       |  |                                             
                       |  |              +-- Schistodesmus_lampreyanus  
                       +--+  +-----------+                              
                          |  |           +-- Schistodesmus_spinosus     
                          |  |                                          
                          +--+        +----- Nodularia_douglasiae       
                             |  +-----+                                 
                             |  |     |  +-- Nodularia_huana            
                             |  |     +--+                              
                             +--+        +-- Nodularia_nuxpersicae      
                                |                                       
                                |  +-------- Cuneopsis_rufescens        
                                +--+                                    
                                   |  +----- Tchangsinaia_piscicula     
                                   +--+                                 
                                      |  +-- Cuneopsis_heudei           
                                      +--+                              
                                         +-- Cuneopsis_celtiformis      
Cladogram Figure 2
                                                     +-- Neotrigonia_margaritacea   
  +--------------------------------------------------+                              
  |                                                  +-- Neotrigonia_lamarckii      
  |                                                                                 
  |                                               +----- Alathyria_jacksoni         
  |                                            +--+                                 
  |                                            |  |  +-- Alathyria_profuga          
  |                                            |  +--+                              
  |                                   +--------+     +-- Velesunio_ambiguus         
  |                                   |        |                                    
  |                                   |        |  +----- Alathyria_pertexta         
  |                                   |        +--+                                 
  |                                   |           |  +-- Lortiella_froggatti        
  |  +--------------------------------+           +--+                              
  |  |                                |              +-- Microdontia_anodontaeformis
  |  |                                |                                             
  |  |                                |  +-------------- Westralunio_carteri        
  |  |                                |  |                                          
  |  |                                +--+  +----------- Echyridella_menziesii      
  |  |                                   |  |                                       
  |  |                                   |  |     +----- Diplodon_suavidicus        
--+  |                                   +--+  +--+                                 
  |  |                                      |  |  |  +-- Castalia_ambigua           
  |  |                                      |  |  +--+                              
  |  |                                      +--+     +-- Triplodon_corrugatus       
  |  |                                         |                                    
  |  |                                         |  +----- Hyridella_australis        
  |  |                                         +--+                                 
  |  |                                            |  +-- Hyridella_depressa         
  |  |                                            +--+                              
  |  |                                               +-- Hyridella_drapeta          
  |  |                                                                              
  |  |                                         +-------- Anodontites_trapesialis    
  |  |                                      +--+                                    
  |  |                                      |  |  +----- Mutela_rostrata            
  |  |                                      |  +--+                                 
  |  |                                      |     |  +-- Mutela_dubia               
  |  |                                      |     +--+                              
  |  |  +-----------------------------------+        +-- Mutela_hargeri             
  |  |  |                                   |                                       
  |  |  |                                   |     +----- Mycetopoda_siliquosa       
  |  |  |                                   |  +--+                                 
  +--+  |                                   |  |  |  +-- Tamsiella_schroeteriana    
     |  |                                   |  |  +--+                              
     |  |                                   +--+     +-- Lamproscapha_ensiformis    
     |  |                                      |                                    
     |  |                                      |  +----- Etheria_elliptica          
     |  |                                      +--+                                 
     |  |                                         |  +-- Aspatharia_pfeifferiana    
     |  |                                         +--+                              
     |  |                                            +-- Chambardia_wahlbergi       
     |  |                                                                           
     |  |                             +----------------- Gibbosula_rochechouartii   
     |  |                             |                                             
     |  |                             |  +-------------- Gibbosula_laosensis        
     |  |  +--------------------------+  |                                          
     |  |  |                          |  |  +----------- Cumberlandia_monodonta     
     |  |  |                          |  |  |                                       
     |  |  |                          +--+  |     +----- Pseudunio_marocanus        
     |  |  |                             |  +-----+                                 
     |  |  |                             |  |     |  +-- Pseudunio_auricularius     
     |  |  |                             |  |     +--+                              
     |  |  |                             +--+        +-- Pseudunio_homsensis        
     |  |  |                                |                                       
     |  |  |                                |        +-- Margaritifera_dahurica     
     |  |  |                                |  +-----+                              
     |  |  |                                |  |     +-- Margaritifera_margaritifera
     |  |  |                                |  |                                    
     |  |  |                                +--+  +----- Margaritifera_laevis       
     |  |  |                                   |  |                                 
     |  |  |                                   |  |  +-- Margaritifera_falcata      
     |  |  |                                   +--+--+                              
     +--+  |                                      |  +-- Margaritifera_middendorffi 
        |  |                                      |                                 
        |  |                                      |  +-- Margaritifera_hembeli      
        |  |                                      +--+                              
        |  |                                         +-- Margaritifera_marrianae    
        |  |                                                                        
        |  |                                      +----- Trapezidens_exolescens     
        |  |                    +-----------------+                                 
        |  |                    |                 |  +-- Trapezidens_dolichorhynchus
        |  |                    |                 +--+                              
        |  |                 +--+                    +-- Trapezidens_angustior      
        |  |                 |  |                                                   
        |  |                 |  |  +-------------------- Lamellidens_corrianus      
        |  |                 |  |  |                                                
        |  |                 |  +--+  +----------------- Lamellidens_savadiensis    
        |  |                 |     |  |                                             
        |  |                 |     +--+              +-- Lamellidens_lamellatus     
        |  |                 |        |  +-----------+                              
        |  |                 |        |  |           +-- Lamellidens_jenkinsianus   
        |  |                 |        +--+                                          
        |  |                 |           |  +----------- Lamellidens_generosus      
        |  |                 |           +--+                                       
        |  |                 |              |  +-------- Lamellidens_mainwaringi__2 
        |  |                 |              +--+                                    
        |  |  +--------------+                 |  +----- Lamellidens_mainwaringi__1 
        |  |  |              |                 +--+                                 
        |  |  |              |                    |  +-- Lamellidens_ferrugineus    
        |  |  |              |                    +--+                              
        |  |  |              |                       +-- Lamellidens_marginalis     
        |  |  |              |                                                      
        |  |  |              |                    +----- Parreysia_keralaensis      
        |  |  |              |  +-----------------+                                 
        |  |  |              |  |                 |  +-- Parreysia_nagpoorensis     
        |  |  |              |  |                 +--+                              
        |  |  |              |  |                    +-- Parreysia_plagiosoma       
        +--+  |              |  |                                                   
           |  |              |  |                    +-- Leoparreysia_tavoyensis    
           |  |              |  |                    |                              
           |  |              +--+  +-----------------+-- Leoparreysia_canefrii      
           |  |                 |  |                 |                              
           |  |                 |  |                 +-- Leoparreysia_olivacea      
           |  |                 |  |                                                
           |  |                 |  |           +-------- Coelatura_aegyptiaca__2    
           |  |                 |  |           |                                    
           |  |                 |  |  +--------+  +----- Coelatura_aegyptiaca__1    
           |  |                 +--+  |        |  |                                 
           |  |                    |  |        |  +----- Nitia_teretiuscula         
           |  |                    |  |        +--+                                 
           |  |                    |  |           |  +-- Coelatura_gabonensis       
           |  |                    |  |           |  |                              
           |  |                    |  |           +--+-- Coelatura_kunenensis       
           |  |                    |  |              |                              
           |  |                    +--+              +-- Prisodontopsis_mweruensis  
           |  |                       |                                             
           |  |                       |        +-------- Indonaia_shurtleffiana     
           |  |                       |        |                                    
           |  |                       |  +-----+  +----- Indonaia_bonneaudii        
           |  |                       |  |     |  |                                 
           |  |                       |  |     +--+----- Indonaia_rugosa            
           |  |                       |  |        |                                 
           |  |                       |  |        |  +-- Indonaia_andersoniana      
           |  |                       +--+        +--+                              
           |  |                          |           +-- Indonaia_caerulea          
           |  |                          |                                          
           |  |                          |  +----------- Indochinella_pugio         
           |  |                          |  |                                       
           |  |                          |  |     +----- Scabies_humilis            
           |  |                          +--+  +--+                                 
           |  |                             |  |  |  +-- Scabies_phaselus           
           |  |                             |  |  +--+                              
           |  |                             +--+     +-- Scabies_mandarinus         
           |  |                                |                                    
           |  |                                |  +----- Radiatula_myitkyinae       
           |  |                                +--+                                 
           |  |                                   |  +-- Radiatula_chaudhurii       
           |  |                                   +--+                              
           |  |                                      +-- Radiatula_mouhoti          
           |  |                                                                     
           |  |        +-------------------------------- Acuticosta_chinensis       
           |  |        |                                                            
           |  |        |  +----------------------------- Lepidodesma_languilati     
           |  |        |  |                                                         
           |  |        |  |                    +-------- Aculamprotula_tortuosa     
           |  |        |  |                    |                                    
           |  |        |  |        +-----------+  +----- Aculamprotula_coreana      
           |  |        |  |        |           |  |                                 
           |  |        |  |        |           +--+----- Aculamprotula_scripta      
           |  |        |  |        |              |                                 
           +--+        |  |        |              |  +-- Aculamprotula_fibrosa      
              |        |  |        |              +--+                              
              |        |  |        |                 +-- Aculamprotula_tientsinensis
              |  +-----+  +--------+                                                
              |  |     |  |        |           +-------- Unio_tumidus               
              |  |     |  |        |  +--------+                                    
              |  |     |  |        |  |        |  +----- Unio_crassus               
              |  |     |  |        |  |        +--+                                 
              |  |     |  |        |  |           |  +-- Unio_delphinus             
              |  |     |  |        |  |           +--+                              
              |  |     |  |        +--+              +-- Unio_pictorum              
              |  |     |  |           |                                             
              |  |     |  |           |              +-- Schistodesmus_lampreyanus  
              |  |     |  |           |  +-----------+                              
              |  |     |  |           |  |           +-- Schistodesmus_spinosus     
              |  |     |  |           |  |                                          
              |  |     |  |           +--+        +----- Nodularia_douglasiae       
              |  |     +--+              |  +-----+                                 
              |  |        |              |  |     |  +-- Nodularia_huana            
              |  |        |              |  |     +--+                              
              |  |        |              +--+        +-- Nodularia_nuxpersicae      
              |  |        |                 |                                       
              |  |        |                 |  +-------- Tchangsinaia_piscicula     
              |  |        |                 +--+                                    
              |  |        |                    |  +----- Cuneopsis_rufescens        
              |  |        |                    +--+                                 
              |  |        |                       |  +-- Cuneopsis_heudei           
              |  |        |                       +--+                              
              |  |        |                          +-- Cuneopsis_celtiformis      
              |  |        |                                                         
              |  |        |                    +-------- Lanceolaria_gladiola       
              |  |        |                    |                                    
              |  |        |  +-----------------+  +----- Lanceolaria_grayii__1      
              |  |        |  |                 |  |                                 
              |  |        |  |                 +--+----- Lanceolaria_grayii__2      
              |  |        |  |                    |                                 
              |  |        |  |                    |  +-- Lanceolaria_lanceolata     
              |  |        |  |                    +--+                              
              |  |        |  |                       +-- Lanceolaria_triformis      
              |  |        |  |                                                      
              |  |        |  |     +-------------------- Anemina_arcaeformis        
              |  |        |  |     |                                                
              |  |        |  |  +--+  +----------------- Cristaria_plicata          
              |  |        +--+  |  |  |                                             
              |  |           |  |  +--+  +-------------- Pletholophus_tenuis        
              |  |           |  |     |  |                                          
              |  |           |  |     +--+           +-- Sinanodonta_angula         
              |  |           |  |        |  +--------+                              
              |  |           |  |        |  |        +-- Sinanodonta_lucida         
              |  |           |  |        +--+                                       
              |  |           |  |           |  +-------- Sinanodonta_woodiana       
              |  |           |  |           +--+                                    
              |  |           |  |              |  +----- Sinanodonta_jourdyi        
              |  |           |  |              +--+                                 
              |  |           +--+                 |  +-- Sinanodonta_elliptica      
              |  |              |                 +--+                              
              |  |              |                    +-- Sinanodonta_pacifica       
              |  |              |                                                   
              |  |              |              +-------- Alasmidonta_marginata      
              |  |              |              |                                    
              |  |              |              +-------- Lasmigona_costata          
              |  |              |              |                                    
              |  |              |              +-------- Pseudanodonta_complanata   
              |  |              |              |                                    
              |  |              |              +-------- Simpsonaias_ambigua        
              +--+              |              |                                    
                 |              +--------------+-------- Strophitus_undulatus       
                 |                             |                                    
                 |                             |  +----- Anodonta_anatina           
                 |                             +--+                                 
                 |                             |  |  +-- Anodonta_cygnea            
                 |                             |  +--+                              
                 |                             |     +-- Anodonta_nuttalliana       
                 |                             |                                    
                 |                             |  +----- Pyganodon_grandis          
                 |                             +--+                                 
                 |                                |  +-- Utterbackia_imbecillis     
                 |                                +--+                              
                 |                                   +-- Utterbackia_peninsularis   
                 |                                                                  
                 |                          +----------- Hyriopsis_desowitzi        
                 |                          |                                       
                 |                          +----------- Rectidens_sumatrensis      
                 |                          |                                       
                 |                          |        +-- Hyriopsis_bialata          
                 |                          |        |                              
                 |                          |        +-- Hyriopsis_myersiana        
                 |                    +-----+--------+                              
                 |                    |     |        +-- Hyriopsis_panhai           
                 |                    |     |        |                              
                 |                    |     |        +-- Hyriopsis_khoratensis      
                 |                    |     |                                       
                 |                    |     |        +-- Ensidens_ingallsianus      
                 |                    |     |  +-----+                              
                 |                    |     |  |     +-- Ensidens_telus             
                 |                    |     +--+                                    
                 |                    |        |  +----- Ensidens_sagittarius       
                 |  +-----------------+        +--+                                 
                 |  |                 |           |  +-- Ensidens_spiculus          
                 |  |                 |           +--+                              
                 |  |                 |              +-- Ensidens_jaculus           
                 |  |                 |                                             
                 |  |                 |           +----- Physunio                   
                 |  |                 |  +--------+                                 
                 |  |                 |  |        |  +-- Physunio_modelli           
                 |  |                 |  |        +--+                              
                 |  |                 |  |           +-- Physunio_superbus          
                 |  |                 |  |                                          
                 |  |                 +--+     +-------- Yaukthwa_inlenensis        
                 |  |                    |  +--+                                    
                 |  |                    |  |  |  +----- Yaukthwa_panhai            
                 |  |                    |  |  +--+                                 
                 |  |                    |  |     |  +-- Yaukthwa_nesemanni         
                 |  |                    |  |     +--+                              
                 |  |                    +--+        +-- Yaukthwa_rectangularis     
                 |  |                       |                                       
                 |  |                       |        +-- Lens_contradens            
                 |  |                       |  +-----+                              
                 |  |                       |  |     +-- Lens_micropterus           
                 |  |                       +--+                                    
                 |  |                          |  +----- Lens_eximius               
                 |  |                          +--+                                 
                 |  |                             |  +-- Lens_comptus               
                 |  |                             +--+                              
                 |  |                                +-- Lens_novoselovi            
                 |  |                                                               
                 |  |                                +-- Megalonaias_nervosa        
                 |  |                             +--+                              
                 +--+                             |  +-- Uniomerus_carolinianus     
                    |     +-----------------------+                                 
                    |     |                       |  +-- Quadrula_quadrula          
                    |     |                       +--+                              
                    |     |                          +-- Quadrula_verrucosa         
                    |     |                                                         
                    |     |                    +-------- Elliptoideus_sloatianus    
                    |  +--+                    |                                    
                    |  |  |  +-----------------+     +-- Elliptio_complanata        
                    |  |  |  |                 |  +--+                              
                    |  |  |  |                 |  |  +-- Eurynia_dilatata           
                    |  |  |  |                 +--+                                 
                    |  |  |  |                    |  +-- Fusconaia_flava            
                    |  |  +--+                    +--+                              
                    |  |     |                       +-- Pleurobema_sintoxia        
                    |  |     |                                                      
                    |  |     |  +----------------------- Amblema_plicata            
                    |  |     |  |                                                   
                    |  |     |  |                 +----- Toxolasma_parvum           
                    |  |     +--+  +--------------+                                 
                    |  |        |  |              |  +-- Obliquaria_reflexa         
                    |  |        |  |              +--+                              
                    |  |        |  |                 +-- Plectomerus_dombeyanus     
                    |  |        +--+                                                
                    |  |           |  +----------------- Potamilus_leptodon         
                    |  |           |  |                                             
                    |  |           |  |  +-------------- Potamilus_alatus           
                    |  |           +--+  |                                          
                    |  |              |  |  +----------- Venustaconcha_ellipsiformis
                    |  |              |  |  |                                       
                    |  |              +--+  +----------- Leaunio_lienosus           
                    |  |                 |  |                                       
                    +--+                 |  |        +-- Ellipsaria_lineolata       
                       |                 |  +--------+                              
                       |                 +--+        +-- Truncilla_truncata         
                       |                    |                                       
                       |                    |        +-- Medionidus_conradicus      
                       |                    +--------+                              
                       |                    |        +-- Ptychobranchus_subtentus   
                       |                    |                                       
                       |                    |  +-------- Cambarunio_iris            
                       |                    +--+                                    
                       |                       |  +----- Ortmanniana_ligamentina    
                       |                       +--+                                 
                       |                          |  +-- Lampsilis_cardium          
                       |                          +--+                              
                       |                             +-- Lampsilis_ornata           
                       |                                                            
                       |                          +----- Chamberlainia_somsakpanhai 
                       |  +-----------------------+                                 
                       |  |                       |  +-- Sinohyriopsis_cumingii     
                       |  |                       +--+                              
                       |  |                          +-- Sinohyriopsis_schlegelii   
                       |  |                                                         
                       |  |                          +-- Potomida_littoralis        
                       |  |                    +-----+                              
                       |  |                    |     +-- Pronodularia_japanensis    
                       |  |  +-----------------+                                    
                       +--+  |                 |  +----- Lamprotula_leaii           
                          |  |                 +--+                                 
                          |  |                    |  +-- Lamprotula_caveata         
                          |  |                    +--+                              
                          |  |                       +-- Lamprotula_cornuumlunae    
                          |  |                                                      
                          |  |                 +-------- Sinosolenaia_carinata      
                          |  |              +--+                                    
                          |  |              |  |  +----- Sinosolenaia_recognita     
                          |  |              |  +--+                                 
                          +--+              |     |  +-- Sinosolenaia_oleivora      
                             |              |     +--+                              
                             |  +-----------+        +-- Sinosolenaia_iridinea      
                             |  |           |                                       
                             |  |           |     +----- Gonidea_angulata           
                             |  |           |  +--+                                 
                             |  |           |  |  |  +-- Leguminaia_wheatleyi       
                             |  |           |  |  +--+                              
                             |  |           +--+     +-- Microcondylaea_bonellii    
                             |  |              |                                    
                             |  |              |     +-- Ptychorhynchus_pfisteri    
                             |  |              |     |                              
                             +--+              +-----+-- Parvasolenaia_rivularis    
                                |                    |                              
                                |                    +-- Koreosolenaia_sitgyensis   
                                |                                                   
                                |                 +----- Indopseudodon_salwenianus  
                                |  +--------------+                                 
                                |  |              |  +-- Indopseudodon_indawgyiensis
                                |  |              +--+                              
                                |  |                 +-- Indopseudodon_bogani       
                                |  |                                                
                                |  |           +-------- Pilsbryoconcha_exilis__3   
                                +--+  +--------+                                    
                                   |  |        |  +----- Pilsbryoconcha_exilis__2   
                                   |  |        +--+                                 
                                   |  |           |  +-- Pilsbryoconcha_exilis__1   
                                   |  |           +--+                              
                                   |  |              +-- Pilsbryoconcha_carinifera  
                                   |  |                                             
                                   +--+     +----------- Nyeinchanconcha_nyeinchani 
                                      |  +--+                                       
                                      |  |  |  +-------- Thaiconcha_callifera       
                                      |  |  +--+                                    
                                      |  |     |  +----- Bineurus_loeiensis         
                                      |  |     +--+                                 
                                      |  |        |  +-- Bineurus_mouhotii          
                                      +--+        +--+                              
                                         |           +-- Bineurus_exilis            
                                         |                                          
                                         |        +----- Pseudodon_vagulus          
                                         |     +--+                                 
                                         |     |  |  +-- Pseudodon_mekongi          
                                         |     |  +--+                              
                                         +-----+     +-- Pseudodon_vondembuschianus 
                                               |                                    
                                               |     +-- Sundadontina_cumingii      
                                               |     |                              
                                               +-----+-- Sundadontina_tumida__1     
                                                     |                              
                                                     +-- Sundadontina_tumida__2     
Cladogram Figure 3
  +-------------------------------------------- Neotrigonia_margaritacea   
  |                                                                        
  |                                         +-- Anodontites_trapesialis    
  |  +--------------------------------------+                              
  |  |                                      +-- Mutela_dubia               
--+  |                                                                     
  |  |  +-------------------------------------- Echyridella_menziesii      
  |  |  |                                                                  
  |  |  |                       +-------------- Gibbosula_rochechouartii   
  |  |  |  +--------------------+                                          
  +--+  |  |                    |  +----------- Cumberlandia_monodonta     
     |  |  |                    +--+                                       
     |  |  |                       |  +-------- Pseudunio_marocanus        
     |  |  |                       +--+                                    
     |  |  |                          |  +----- Margaritifera_falcata      
     |  |  |                          +--+                                 
     |  |  |                             |  +-- Margaritifera_margaritifera
     |  |  |                             +--+                              
     +--+  |                                +-- Margaritifera_dahurica     
        |  |                                                               
        |  |                                +-- Sinohyriopsis_cumingii     
        |  |                    +-----------+                              
        |  |                    |           +-- Sinohyriopsis_schlegelii   
        |  |                    |                                          
        |  |                    |     +-------- Potomida_littoralis        
        |  |           +--------+  +--+                                    
        |  |           |        |  |  |  +----- Pronodularia_japanensis    
        |  |           |        |  |  +--+                                 
        |  |           |        |  |     |  +-- Lamprotula_caveata         
        |  |           |        |  |     +--+                              
        +--+           |        +--+        +-- Lamprotula_leaii           
           |           |           |                                       
           |           |           |        +-- Ptychorhynchus_pfisteri    
           |           |           |     +--+                              
           |  +--------+           |     |  +-- Parvasolenaia_rivularis    
           |  |        |           +-----+                                 
           |  |        |                 |  +-- Sinosolenaia_oleivora      
           |  |        |                 +--+                              
           |  |        |                    +-- Sinosolenaia_carinata      
           |  |        |                                                   
           |  |        |                    +-- Quadrula_quadrula          
           |  |        |  +-----------------+                              
           |  |        |  |                 +-- Uniomerus_tetralasmus      
           |  |        +--+                                                
           |  |           |  +----------------- Elliptio_complanata        
           |  |           |  |                                             
           |  |           +--+  +-------------- Toxolasma_parvum           
           |  |              |  |                                          
           |  |              +--+           +-- Potamilus_leptodon         
           |  |                 |  +--------+                              
           |  |                 |  |        +-- Potamilus_alatus           
           +--+                 +--+                                       
              |                    |  +-------- Leaunio_lienosus           
              |                    +--+                                    
              |                       |  +----- Venustaconcha_ellipsiformis
              |                       +--+                                 
              |                          |  +-- Lampsilis_cardium          
              |                          +--+                              
              |                             +-- Lampsilis_ornata           
              |                                                            
              |  +----------------------------- Lepidodesma_languilati     
              |  |                                                         
              |  |                    +-------- Aculamprotula_tortuosa     
              |  |              +-----+                                    
              |  |              |     |  +----- Aculamprotula_scripta      
              |  |              |     +--+                                 
              |  |              |        |  +-- Aculamprotula_tientsinensis
              |  |              |        +--+                              
              |  |  +-----------+           +-- Aculamprotula_coreana      
              +--+  |           |                                          
                 |  |           |        +----- Nodularia_douglasiae       
                 |  |           |  +-----+                                 
                 |  |           |  |     |  +-- Tchangsinaia_piscicula     
                 |  |           |  |     +--+                              
                 |  |           +--+        +-- Schistodesmus_lampreyanus  
                 |  |              |                                       
                 |  |              |  +-------- Unio_tumidus               
                 |  |              +--+                                    
                 +--+                 |  +----- Unio_nanus                 
                    |                 +--+                                 
                    |                    |  +-- Unio_delphinus             
                    |                    +--+                              
                    |                       +-- Unio_pictorum              
                    |                                                      
                    |                    +----- Lanceolaria_gladiola       
                    |  +-----------------+                                 
                    |  |                 |  +-- Lanceolaria_lanceolata     
                    |  |                 +--+                              
                    |  |                    +-- Lanceolaria_grayii         
                    +--+                                                   
                       |  +-------------------- Acuticosta_chinensis       
                       |  |                                                
                       |  |           +-------- Cristaria_plicata          
                       |  |           |                                    
                       +--+  +--------+     +-- Sinanodonta_lucida         
                          |  |        |  +--+                              
                          |  |        |  |  +-- Sinanodonta_woodiana       
                          |  |        +--+                                 
                          +--+           |  +-- Anemina_arcaeformis        
                             |           +--+                              
                             |              +-- Anemina_euscaphys          
                             |                                             
                             |  +-------------- Anodonta_cygnea            
                             +--+                                          
                                |  +----------- Anodonta_anatina           
                                +--+                                       
                                   |  +-------- Lasmigona_costata          
                                   +--+                                    
                                      |  +----- Pyganodon_grandis          
                                      +--+                                 
                                         |  +-- Utterbackia_imbecillis     
                                         +--+                              
                                            +-- Utterbackia_peninsularis   
Cladogram Figure 3-M
  +----------------------------- Echyridella_menziesii      
  |                                                         
  |                          +-- Pseudunio_marocanus        
  |  +-----------------------+                              
--+  |                       +-- Margaritifera_margaritifera
  |  |                                                      
  |  |                    +----- Quadrula_quadrula          
  |  |  +-----------------+                                 
  |  |  |                 |  +-- Venustaconcha_ellipsiformis
  +--+  |                 +--+                              
     |  |                    +-- Potamilus_alatus           
     |  |                                                   
     |  |           +----------- Sinohyriopsis_cumingii     
     |  |           |                                       
     |  |           |     +----- Potomida_littoralis        
     +--+  +--------+  +--+                                 
        |  |        |  |  |  +-- Lamprotula_leaii           
        |  |        |  |  +--+                              
        |  |        +--+     +-- Pronodularia_japanensis    
        |  |           |                                    
        |  |           |  +----- Parvasolenaia_rivularis    
        |  |           +--+                                 
        |  |              |  +-- Sinosolenaia_oleivora      
        +--+              +--+                              
           |                 +-- Sinosolenaia_carinata      
           |                                                
           |           +-------- Sinanodonta_woodiana       
           |  +--------+                                    
           |  |        |  +----- Anodonta_anatina           
           |  |        +--+                                 
           |  |           |  +-- Pyganodon_grandis          
           +--+           +--+                              
              |              +-- Utterbackia_peninsularis   
              |                                             
              |  +-------------- Lanceolaria_lanceolata     
              +--+                                          
                 |  +----------- Schistodesmus_lampreyanus  
                 +--+                                       
                    |  +-------- Aculamprotula_tortuosa     
                    +--+                                    
                       |  +----- Unio_tumidus               
                       +--+                                 
                          |  +-- Unio_nanus                 
                          +--+                              
                             +-- Unio_delphinus             
Cladogram Figure S1
                                      +-- Gibbosula_rochechouartii   
  +-----------------------------------+                              
  |                                   +-- Margaritifera_dahurica     
  |                                                                  
  |                                +----- Sinohyriopsis_cumingii__2  
  |                 +--------------+                                 
  |                 |              |  +-- Sinohyriopsis_cumingii__1  
  |                 |              +--+                              
  |                 |                 +-- Sinohyriopsis_schlegelii   
  |                 |                                                
  |                 |              +----- Parvasolenaia_rivularis    
  |  +--------------+  +-----------+                                 
  |  |              |  |           |  +-- Parvasolenaia_triangularis 
  |  |              |  |           +--+                              
  |  |              |  |              +-- Ptychorhynchus_pfisteri    
  |  |              |  |                                             
  |  |              |  |        +-------- Sinosolenaia_recognita     
  |  |              +--+        |                                    
--+  |                 |  +-----+  +----- Sinosolenaia_oleivora__2   
  |  |                 |  |     |  |                                 
  |  |                 |  |     +--+  +-- Sinosolenaia_oleivora__1   
  |  |                 |  |        |  |                              
  |  |                 |  |        +--+-- Sinosolenaia_iridinea      
  |  |                 +--+           |                              
  |  |                    |           +-- Sinosolenaia_oleivora__3   
  |  |                    |                                          
  |  |                    |  +----------- Sinosolenaia_carinata      
  |  |                    |  |                                       
  |  |                    +--+  +-------- Lamprotula_leaii           
  |  |                       |  |                                    
  |  |                       +--+  +----- Lamprotula_cornuumlunae__1 
  |  |                          |  |                                 
  |  |                          |  +----- Lamprotula_cornuumlunae__3 
  |  |                          +--+                                 
  |  |                             +----- Lamprotula_cornuumlunae__4 
  +--+                             |                                 
     |                             |  +-- Lamprotula_cornuumlunae__2 
     |                             +--+                              
     |                                +-- Lamprotula_caveata         
     |                                                               
     |                    +-------------- Anemina_arcaeformis__2     
     |                    |                                          
     |  +-----------------+  +----------- Anemina_arcaeformis__5     
     |  |                 |  |                                       
     |  |                 |  |  +-------- Anemina_arcaeformis__3     
     |  |                 +--+  |                                    
     |  |                    |  +-------- Anemina_arcaeformis__7     
     |  |                    |  |                                    
     |  |                    +--+  +----- Anemina_arcaeformis__4     
     |  |                       |  |                                 
     |  |                       |  +----- Anemina_arcaeformis__6     
     |  |                       |  |                                 
     |  |                       +--+----- Anemina_arcaeformis__8     
     |  |                          |                                 
     |  |                          +----- Anemina_arcaeformis__9     
     |  |                          |                                 
     |  |                          |  +-- Anemina_arcaeformis__1     
     |  |                          +--+                              
     |  |                             +-- Anemina_euscaphys          
     +--+                                                            
        |                             +-- Sinanodonta_angula         
        |                 +-----------+                              
        |                 |           +-- Sinanodonta_lucida         
        |  +--------------+                                          
        |  |              |  +----------- Sinanodonta_woodiana__2    
        |  |              +--+                                       
        |  |                 |  +-------- Sinanodonta_jourdyi        
        |  |                 +--+                                    
        |  |                    |  +----- Sinanodonta_elliptica      
        |  |                    +--+                                 
        |  |                       |  +-- Sinanodonta_woodiana__1    
        |  |                       +--+                              
        |  |                          +-- Sinanodonta_lauta          
        |  |                                                         
        |  |              +-------------- Acuticosta_ovata__1        
        |  |              |                                          
        |  |              |  +----------- Acuticosta_chinensis__3    
        +--+  +-----------+  |                                       
           |  |           |  +----------- Acuticosta_chinensis__4    
           |  |           |  |                                       
           |  |           +--+----------- Acuticosta_chinensis__6    
           |  |              |                                       
           |  |              |        +-- Acuticosta_chinensis__1    
           |  |              |  +-----+                              
           |  |              |  |     +-- Acuticosta_ovata__2        
           |  |              +--+                                    
           |  |                 |  +----- Acuticosta_chinensis__5    
           |  |                 |  |                                 
           |  |                 +--+----- Acuticosta_chinensis__7    
           |  |                    |                                 
           |  |                    |  +-- Acuticosta_chinensis__2    
           +--+                    +--+                              
              |                       +-- Acuticosta_ovata__3        
              |                                                      
              |                       +-- Lepidodesma_languilati     
              |  +--------------------+                              
              |  |                    +-- Cristaria_plicata          
              |  |                                                   
              |  |           +----------- Lanceolaria_gladiola       
              |  |           |                                       
              |  |           |  +-------- Lanceolaria_grayii__5      
              |  |  +--------+  |                                    
              |  |  |        |  |     +-- Lanceolaria_lanceolata     
              |  |  |        |  +-----+                              
              |  |  |        |  |     +-- Lanceolaria_triformis      
              +--+  |        +--+                                    
                 |  |           |  +----- Lanceolaria_grayii__4      
                 |  |           |  |                                 
                 |  |           |  |  +-- Lanceolaria_eucylindrica__1
                 |  |           |  |  |                              
                 |  |           +--+  +-- Lanceolaria_eucylindrica__2
                 |  |              |  |                              
                 |  |              |  +-- Lanceolaria_eucylindrica__3
                 |  |              |  |                              
                 |  |              +--+-- Lanceolaria_eucylindrica__4
                 +--+                 |                              
                    |                 +-- Lanceolaria_grayii__1      
                    |                 |                              
                    |                 +-- Lanceolaria_grayii__2      
                    |                 |                              
                    |                 +-- Lanceolaria_grayii__3      
                    |                                                
                    |        +----------- Aculamprotula_tortuosa     
                    |  +-----+                                       
                    |  |     |  +-------- Aculamprotula_tientsinensis
                    |  |     +--+                                    
                    |  |        |  +----- Aculamprotula_fibrosa      
                    |  |        +--+                                 
                    |  |           |  +-- Aculamprotula_coreana      
                    +--+           +--+                              
                       |              +-- Aculamprotula_scripta      
                       |                                             
                       |           +----- Nodularia_huana            
                       |  +--------+                                 
                       |  |        |  +-- Nodularia_nuxpersicae      
                       |  |        +--+                              
                       +--+           +-- Nodularia_douglasiae       
                          |                                          
                          |  +----------- Tchangsinaia_piscicula     
                          |  |                                       
                          +--+        +-- Schistodesmus_spinosus     
                             |  +-----+                              
                             |  |     +-- Schistodesmus_lampreyanus  
                             +--+                                    
                                |  +----- Cuneopsis_rufescens        
                                +--+                                 
                                   |  +-- Cuneopsis_heudei           
                                   +--+                              
                                      +-- Cuneopsis_celtiformis      
Cladogram Figure S3
                                                                    +-- Neotrigonia_margaritacea     
  +-----------------------------------------------------------------+                                
  |                                                                 +-- Neotrigonia_lamarckii        
  |                                                                                                  
  |                                                              +----- Velesunio_ambiguus           
  |                                                           +--+                                   
  |                                                           |  |  +-- Alathyria_jacksoni           
  |                                                           |  +--+                                
  |                                                  +--------+     +-- Alathyria_profuga            
  |                                                  |        |                                      
  |                                                  |        |  +----- Alathyria_pertexta           
  |                                                  |        +--+                                   
  |                                                  |           |  +-- Lortiella_froggatti          
  |  +-----------------------------------------------+           +--+                                
  |  |                                               |              +-- Microdontia_anodontaeformis  
  |  |                                               |                                               
  |  |                                               |  +-------------- Westralunio_carteri          
  |  |                                               |  |                                            
  |  |                                               +--+  +----------- Echyridella_menziesii        
  |  |                                                  |  |                                         
  |  |                                                  |  |     +----- Diplodon_suavidicus          
--+  |                                                  +--+  +--+                                   
  |  |                                                     |  |  |  +-- Castalia_ambigua             
  |  |                                                     |  |  +--+                                
  |  |                                                     +--+     +-- Triplodon_corrugatus         
  |  |                                                        |                                      
  |  |                                                        |  +----- Hyridella_australis          
  |  |                                                        +--+                                   
  |  |                                                           |  +-- Hyridella_depressa           
  |  |                                                           +--+                                
  |  |                                                              +-- Hyridella_drapeta            
  |  |                                                                                               
  |  |                                                        +-------- Anodontites_trapesialis      
  |  |                                                     +--+                                      
  |  |                                                     |  |  +----- Mutela_rostrata              
  |  |                                                     |  +--+                                   
  |  |                                                     |     |  +-- Mutela_dubia                 
  |  |                                                     |     +--+                                
  |  |  +--------------------------------------------------+        +-- Mutela_hargeri               
  |  |  |                                                  |                                         
  |  |  |                                                  |     +----- Mycetopoda_siliquosa         
  +--+  |                                                  |  +--+                                   
     |  |                                                  |  |  |  +-- Tamsiella_schroeteriana      
     |  |                                                  |  |  +--+                                
     |  |                                                  +--+     +-- Lamproscapha_ensiformis      
     |  |                                                     |                                      
     |  |                                                     |  +----- Etheria_elliptica            
     |  |                                                     +--+                                   
     |  |                                                        |  +-- Aspatharia_pfeifferiana      
     |  |                                                        +--+                                
     |  |                                                           +-- Chambardia_wahlbergi         
     |  |                                                                                            
     |  |                                         +-------------------- Gibbosula_rochechouartii     
     |  |                                         |                                                  
     |  |                                         |  +----------------- Gibbosula_laosensis          
     |  |  +--------------------------------------+  |                                               
     |  |  |                                      |  |        +-------- Cumberlandia_monodonta       
     |  |  |                                      |  |  +-----+                                      
     |  |  |                                      +--+  |     |  +----- Pseudunio_marocanus          
     |  |  |                                         |  |     +--+                                   
     |  |  |                                         |  |        |  +-- Pseudunio_auricularius       
     |  |  |                                         |  |        +--+                                
     |  |  |                                         +--+           +-- Pseudunio_homsensis          
     |  |  |                                            |                                            
     |  |  |                                            |           +-- Margaritifera_dahurica       
     |  |  |                                            |  +--------+                                
     |  |  |                                            |  |        +-- Margaritifera_margaritifera  
     |  |  |                                            +--+                                         
     |  |  |                                               |  +-------- Margaritifera_falcata        
     +--+  |                                               |  |                                      
        |  |                                               +--+     +-- Margaritifera_hembeli        
        |  |                                                  |  +--+                                
        |  |                                                  |  |  +-- Margaritifera_marrianae      
        |  |                                                  +--+                                   
        |  |                                                     |  +-- Margaritifera_laevis         
        |  |                                                     +--+                                
        |  |                                                        +-- Margaritifera_middendorffi   
        |  |                                                                                         
        |  |                                                     +----- Trapezidens_exolescens       
        |  |                                               +-----+                                   
        |  |                                               |     |  +-- Trapezidens_angustior        
        |  |                                               |     +--+                                
        |  |                                               |        +-- Trapezidens_dolichorhynchus  
        |  |                                               |                                         
        |  |                                +--------------+        +-- Lamellidens_corrianus        
        |  |                                |              |  +-----+                                
        |  |                                |              |  |     +-- Lamellidens_savadiensis      
        |  |                                |              |  |                                      
        |  |                                |              |  |  +----- Lamellidens_lamellatus       
        |  |                                |              +--+  |                                   
        |  |                                |                 |  +----- Lamellidens_jenkinsianus     
        |  |                                |                 |  |                                   
        |  |                                |                 |  |  +-- Lamellidens_marginalis       
        |  |                                |                 +--+--+                                
        |  |                                |                    |  +-- Lamellidens_ferrugineus      
        |  |                                |                    |                                   
        |  |  +-----------------------------+                    |  +-- Lamellidens_mainwaringi__1   
        |  |  |                             |                    |  |                                
        |  |  |                             |                    +--+-- Lamellidens_mainwaringi__2   
        |  |  |                             |                       |                                
        |  |  |                             |                       +-- Lamellidens_generosus        
        +--+  |                             |                                                        
           |  |                             |                    +----- Parreysia_keralaensis        
           |  |                             |  +-----------------+                                   
           |  |                             |  |                 |  +-- Parreysia_nagpoorensis       
           |  |                             |  |                 +--+                                
           |  |                             |  |                    +-- Parreysia_plagiosoma         
           |  |                             |  |                                                     
           |  |                             |  |                 +----- Leoparreysia_canefrii        
           |  |                             +--+  +--------------+                                   
           |  |                                |  |              |  +-- Leoparreysia_tavoyensis      
           |  |                                |  |              +--+                                
           |  |                                |  |                 +-- Leoparreysia_olivacea        
           |  |                                |  |                                                  
           |  |                                |  |        +----------- Coelatura_aegyptiaca__2      
           |  |                                |  |        |                                         
           |  |                                +--+  +-----+        +-- Coelatura_aegyptiaca__1      
           |  |                                   |  |     |  +-----+                                
           |  |                                   |  |     |  |     +-- Nitia_teretiuscula           
           |  |                                   |  |     +--+                                      
           |  |                                   |  |        |  +----- Coelatura_kunenensis         
           |  |                                   |  |        +--+                                   
           |  |                                   |  |           |  +-- Coelatura_gabonensis         
           |  |                                   |  |           +--+                                
           |  |                                   +--+              +-- Prisodontopsis_mweruensis    
           |  |                                      |                                               
           |  |                                      |        +-------- Indonaia_shurtleffiana       
           |  |                                      |        |                                      
           |  |                                      |  +-----+  +----- Indonaia_bonneaudii          
           |  |                                      |  |     |  |                                   
           |  |                                      |  |     +--+----- Indonaia_rugosa              
           |  |                                      |  |        |                                   
           |  |                                      |  |        |  +-- Indonaia_andersoniana        
           |  |                                      +--+        +--+                                
           |  |                                         |           +-- Indonaia_caerulea            
           |  |                                         |                                            
           |  |                                         |  +----------- Indochinella_pugio           
           |  |                                         |  |                                         
           |  |                                         |  |     +----- Scabies_phaselus             
           |  |                                         +--+  +--+                                   
           |  |                                            |  |  |  +-- Scabies_humilis              
           |  |                                            |  |  +--+                                
           |  |                                            +--+     +-- Scabies_mandarinus           
           |  |                                               |                                      
           |  |                                               |  +----- Radiatula_mouhoti            
           |  |                                               +--+                                   
           |  |                                                  |  +-- Radiatula_myitkyinae         
           |  |                                                  +--+                                
           +--+                                                     +-- Radiatula_chaudhurii         
              |                                                                                      
              |                                                  +----- Indopseudodon_indawgyiensis  
              |                                   +--------------+                                   
              |                                   |              |  +-- Indopseudodon_bogani         
              |                                   |              +--+                                
              |                                   |                 +-- Indopseudodon_salwenianus    
              |                                   |                                                  
              |                                   |           +-------- Pilsbryoconcha_exilis__3     
              |  +--------------------------------+  +--------+                                      
              |  |                                |  |        |  +----- Pilsbryoconcha_exilis__2     
              |  |                                |  |        +--+                                   
              |  |                                |  |           |  +-- Pilsbryoconcha_carinifera    
              |  |                                |  |           +--+                                
              |  |                                |  |              +-- Pilsbryoconcha_exilis__1     
              |  |                                |  |                                               
              |  |                                +--+     +----------- Sundadontina_tumida          
              |  |                                   |  +--+                                         
              |  |                                   |  |  |  +-------- Pseudodon_vagulus            
              |  |                                   |  |  +--+                                      
              |  |                                   |  |     |  +----- Pseudodon_vondembuschianus__1
              |  |                                   |  |     +--+                                   
              |  |                                   |  |        |  +-- Pseudodon_mekongi            
              |  |                                   +--+        +--+                                
              |  |                                      |           +-- Pseudodon_vondembuschianus__2
              |  |                                      |                                            
              |  |                                      |           +-- Nyeinchanconcha_nyeinchani   
              |  |                                      |  +--------+                                
              |  |                                      |  |        +-- Sundadontina_cumingii        
              |  |                                      +--+                                         
              |  |                                         |  +-------- Thaiconcha_callifera         
              |  |                                         +--+                                      
              |  |                                            |  +----- Bineurus_loeiensis           
              |  |                                            +--+                                   
              |  |                                               |  +-- Bineurus_mouhotii            
              |  |                                               +--+                                
              |  |                                                  +-- Bineurus_exilis              
              |  |                                                                                   
              |  |                                                  +-- Ensidens_telus               
              |  |                                            +-----+                                
              |  |                                            |     +-- Ensidens_ingallsianus        
              |  |                                      +-----+                                      
              |  |                                      |     |  +----- Ensidens_sagittarius         
              |  |                                      |     +--+                                   
              |  |                                      |        |  +-- Ensidens_spiculus            
              |  |                                   +--+        +--+                                
              |  |                                   |  |           +-- Ensidens_jaculus             
              |  |                                   |  |                                            
              |  |                                   |  |  +----------- Hyriopsis_desowitzi          
              |  |                                   |  |  |                                         
              |  |                                   |  +--+  +-------- Rectidens_sumatrensis        
              |  |                                   |     |  |                                      
              |  |                                   |     +--+     +-- Hyriopsis_panhai             
              +--+                                   |        |  +--+                                
                 |                                   |        |  |  +-- Hyriopsis_khoratensis        
                 |  +--------------------------------+        +--+                                   
                 |  |                                |           |  +-- Hyriopsis_bialata            
                 |  |                                |           +--+                                
                 |  |                                |              +-- Hyriopsis_myersiana          
                 |  |                                |                                               
                 |  |                                |           +----- Physunio                     
                 |  |                                |  +--------+                                   
                 |  |                                |  |        |  +-- Physunio_superbus            
                 |  |                                |  |        +--+                                
                 |  |                                |  |           +-- Physunio_modelli             
                 |  |                                |  |                                            
                 |  |                                +--+     +-------- Yaukthwa_inlenensis          
                 |  |                                   |  +--+                                      
                 |  |                                   |  |  |  +----- Yaukthwa_panhai              
                 |  |                                   |  |  +--+                                   
                 |  |                                   |  |     |  +-- Yaukthwa_nesemanni           
                 |  |                                   |  |     +--+                                
                 |  |                                   +--+        +-- Yaukthwa_rectangularis       
                 |  |                                      |                                         
                 |  |                                      |        +-- Lens_contradens              
                 |  |                                      |  +-----+                                
                 |  |                                      |  |     +-- Lens_micropterus             
                 |  |                                      +--+                                      
                 |  |                                         |  +----- Lens_eximius                 
                 |  |                                         +--+                                   
                 |  |                                            |  +-- Lens_novoselovi              
                 |  |                                            +--+                                
                 |  |                                               +-- Lens_comptus                 
                 |  |                                                                                
                 |  |                                            +----- Chamberlainia_somsakpanhai   
                 |  |     +--------------------------------------+                                   
                 |  |     |                                      |  +-- Sinohyriopsis_cumingii       
                 |  |     |                                      +--+                                
                 |  |     |                                         +-- Sinohyriopsis_schlegelii     
                 |  |     |                                                                          
                 |  |     |                                         +-- Potomida_littoralis          
                 |  |     |                                   +-----+                                
                 |  |     |                                   |     +-- Pronodularia_japanensis      
                 |  |     |                             +-----+                                      
                 |  |     |                             |     |  +----- Lamprotula_leaii             
                 |  |     |                             |     +--+                                   
                 |  |     |                             |        |  +-- Lamprotula_caveata           
                 +--+     |                             |        +--+                                
                    |     |                             |           +-- Lamprotula_cornuumlunae      
                    |  +--+  +--------------------------+                                            
                    |  |  |  |                          |        +----- Sinosolenaia_recognita       
                    |  |  |  |                          |  +-----+                                   
                    |  |  |  |                          |  |     |  +-- Sinosolenaia_oleivora        
                    |  |  |  |                          |  |     +--+                                
                    |  |  |  |                          |  |        +-- Sinosolenaia_iridinea        
                    |  |  |  |                          +--+                                         
                    |  |  |  |                             |     +----- Gonidea_angulata             
                    |  |  |  |                             |  +--+                                   
                    |  |  |  |                             |  |  |  +-- Leguminaia_wheatleyi         
                    |  |  |  |                             |  |  +--+                                
                    |  |  |  |                             +--+     +-- Microcondylaea_bonellii      
                    |  |  |  |                                |                                      
                    |  |  |  |                                |  +----- Ptychorhynchus_pfisteri      
                    |  |  +--+                                +--+                                   
                    |  |     |                                   |  +-- Parvasolenaia_rivularis      
                    |  |     |                                   +--+                                
                    |  |     |                                      +-- Koreosolenaia_sitgyensis     
                    |  |     |                                                                       
                    |  |     |                                      +-- Megalonaias_nervosa          
                    |  |     |                                   +--+                                
                    |  |     |                                   |  +-- Uniomerus_carolinianus       
                    |  |     |  +--------------------------------+                                   
                    |  |     |  |                                |  +-- Quadrula_quadrula            
                    |  |     |  |                                +--+                                
                    |  |     |  |                                   +-- Quadrula_verrucosa           
                    |  |     |  |                                                                    
                    |  |     |  |                             +-------- Elliptoideus_sloatianus      
                    |  |     +--+                             |                                      
                    |  |        |  +--------------------------+     +-- Elliptio_complanata          
                    |  |        |  |                          |  +--+                                
                    |  |        |  |                          |  |  +-- Eurynia_dilatata             
                    |  |        |  |                          +--+                                   
                    |  |        |  |                             |  +-- Fusconaia_flava              
                    |  |        +--+                             +--+                                
                    |  |           |                                +-- Pleurobema_sintoxia          
                    |  |           |                                                                 
                    |  |           |  +-------------------------------- Amblema_plicata              
                    |  |           |  |                                                              
                    |  |           +--+  +----------------------------- Toxolasma_parvum             
                    |  |              |  |                                                           
                    +--+              +--+                          +-- Obliquaria_reflexa           
                       |                 |  +-----------------------+                                
                       |                 |  |                       +-- Plectomerus_dombeyanus       
                       |                 +--+                                                        
                       |                    |  +----------------------- Potamilus_leptodon           
                       |                    |  |                                                     
                       |                    +--+  +-------------------- Potamilus_alatus             
                       |                       |  |                                                  
                       |                       |  |                 +-- Medionidus_conradicus        
                       |                       +--+  +--------------+                                
                       |                          |  |              +-- Ptychobranchus_subtentus     
                       |                          |  |                                               
                       |                          +--+              +-- Ellipsaria_lineolata         
                       |                             |  +-----------+                                
                       |                             |  |           +-- Truncilla_truncata           
                       |                             +--+                                            
                       |                                |  +----------- Leaunio_lienosus             
                       |                                |  |                                         
                       |                                +--+        +-- Venustaconcha_ellipsiformis  
                       |                                   |  +-----+                                
                       |                                   |  |     +-- Cambarunio_iris              
                       |                                   +--+                                      
                       |                                      |  +----- Ortmanniana_ligamentina      
                       |                                      +--+                                   
                       |                                         |  +-- Lampsilis_cardium            
                       |                                         +--+                                
                       |                                            +-- Lampsilis_ornata             
                       |                                                                             
                       |           +----------------------------------- Acuticosta_chinensis         
                       |           |                                                                 
                       |           |                          +-------- Aculamprotula_tortuosa       
                       |           |                          |                                      
                       |           |              +-----------+     +-- Aculamprotula_coreana        
                       |           |              |           |  +--+                                
                       |           |              |           |  |  +-- Aculamprotula_scripta        
                       |           |              |           +--+                                   
                       |           |              |              |  +-- Aculamprotula_fibrosa        
                       |           |              |              +--+                                
                       |           |              |                 +-- Aculamprotula_tientsinensis  
                       |           |  +-----------+                                                  
                       |           |  |           |           +-------- Unio_tumidus                 
                       +-----------+  |           |  +--------+                                      
                                   |  |           |  |        |  +----- Unio_crassus                 
                                   |  |           |  |        +--+                                   
                                   |  |           |  |           |  +-- Unio_delphinus               
                                   |  |           |  |           +--+                                
                                   |  |           +--+              +-- Unio_pictorum                
                                   |  |              |                                               
                                   |  |              |           +----- Nodularia_douglasiae         
                                   |  |              |  +--------+                                   
                                   |  |              |  |        |  +-- Nodularia_huana              
                                   |  |              |  |        +--+                                
                                   |  |              +--+           +-- Nodularia_nuxpersicae        
                                   |  |                 |                                            
                                   +--+                 |  +----------- Schistodesmus_lampreyanus    
                                      |                 +--+                                         
                                      |                    |  +-------- Tchangsinaia_piscicula       
                                      |                    +--+                                      
                                      |                       |  +----- Cuneopsis_rufescens          
                                      |                       +--+                                   
                                      |                          |  +-- Cuneopsis_heudei             
                                      |                          +--+                                
                                      |                             +-- Cuneopsis_celtiformis        
                                      |                                                              
                                      |  +----------------------------- Lepidodesma_languilati       
                                      |  |                                                           
                                      |  |                    +-------- Lanceolaria_gladiola         
                                      |  |  +-----------------+                                      
                                      |  |  |                 |  +----- Lanceolaria_grayii           
                                      |  |  |                 +--+                                   
                                      +--+  |                    |  +-- Lanceolaria_lanceolata       
                                         |  |                    +--+                                
                                         |  |                       +-- Lanceolaria_triformis        
                                         |  |                                                        
                                         |  |     +-------------------- Anemina_arcaeformis          
                                         |  |     |                                                  
                                         |  |  +--+  +----------------- Cristaria_plicata            
                                         +--+  |  |  |                                               
                                            |  |  +--+  +-------------- Pletholophus_tenuis          
                                            |  |     |  |                                            
                                            |  |     +--+           +-- Sinanodonta_angula           
                                            |  |        |  +--------+                                
                                            |  |        |  |        +-- Sinanodonta_lucida           
                                            |  |        +--+                                         
                                            |  |           |  +-------- Sinanodonta_woodiana         
                                            |  |           +--+                                      
                                            |  |              |  +----- Sinanodonta_jourdyi          
                                            +--+              +--+                                   
                                               |                 |  +-- Sinanodonta_elliptica        
                                               |                 +--+                                
                                               |                    +-- Sinanodonta_pacifica         
                                               |                                                     
                                               |                    +-- Alasmidonta_marginata        
                                               |           +--------+                                
                                               |           |        +-- Lasmigona_costata            
                                               |           |                                         
                                               |           |     +----- Anodonta_anatina             
                                               |           +-----+                                   
                                               |           |     |  +-- Anodonta_cygnea              
                                               +-----------+     +--+                                
                                                           |        +-- Anodonta_nuttalliana         
                                                           |                                         
                                                           |     +----- Pseudanodonta_complanata     
                                                           |  +--+                                   
                                                           |  |  |  +-- Simpsonaias_ambigua          
                                                           |  |  +--+                                
                                                           +--+     +-- Strophitus_undulatus         
                                                              |                                      
                                                              |  +----- Pyganodon_grandis            
                                                              +--+                                   
                                                                 |  +-- Utterbackia_imbecillis       
                                                                 +--+                                
                                                                    +-- Utterbackia_peninsularis     
Cladogram Figure S4
                                      +-- Neotrigonia_margaritacea   
  +-----------------------------------+                              
  |                                   +-- Neotrigonia_lamarckii      
  |                                                                  
  |                          +----------- Etheria_elliptica          
  |                          |                                       
  |                          |        +-- Aspatharia_pfeifferiana    
  |                          +--------+                              
  |                          |        +-- Chambardia_wahlbergi       
  |                          |                                       
  |  +-----------------------+     +----- Mycetopoda_siliquosa       
  |  |                       +-----+                                 
  |  |                       |     |  +-- Tamsiella_schroeteriana    
  |  |                       |     +--+                              
  |  |                       |        +-- Lamproscapha_ensiformis    
  |  |                       |                                       
  |  |                       |  +-------- Anodontites_trapesialis    
  |  |                       +--+                                    
  |  |                          |  +----- Mutela_hargeri             
  |  |                          +--+                                 
  |  |                             |  +-- Mutela_dubia               
--+  |                             +--+                              
  |  |                                +-- Mutela_rostrata            
  |  |                                                               
  |  |                       +----------- Echyridella_menziesii      
  |  |                       |                                       
  |  |                       +----------- Westralunio_carteri        
  |  |                       |                                       
  |  |                       |     +----- Diplodon_suavidicus        
  |  |                       +-----+                                 
  |  |                       |     |  +-- Castalia_ambigua           
  |  |                       |     +--+                              
  |  |                       |        +-- Triplodon_corrugatus       
  |  |                       |                                       
  |  |  +--------------------+     +----- Hyridella_australis        
  |  |  |                    +-----+                                 
  |  |  |                    |     |  +-- Hyridella_depressa         
  |  |  |                    |     +--+                              
  |  |  |                    |        +-- Hyridella_drapeta          
  |  |  |                    |                                       
  |  |  |                    |     +----- Velesunio_ambiguus         
  |  |  |                    |  +--+                                 
  +--+  |                    |  |  |  +-- Alathyria_jacksoni         
     |  |                    |  |  +--+                              
     |  |                    +--+     +-- Alathyria_profuga          
     |  |                       |                                    
     |  |                       |  +----- Alathyria_pertexta         
     |  |                       +--+                                 
     |  |                          |  +-- Lortiella_froggatti        
     |  |                          +--+                              
     |  |                             +-- Microdontia_anodontaeformis
     |  |                                                            
     |  |           +-------------------- Gibbosula_rochechouartii   
     |  |           |                                                
     |  |  +--------+  +----------------- Gibbosula_laosensis        
     |  |  |        |  |                                             
     |  |  |        +--+  +-------------- Cumberlandia_monodonta     
     |  |  |           |  |                                          
     |  |  |           |  |        +----- Pseudunio_marocanus        
     |  |  |           +--+  +-----+                                 
     |  |  |              |  |     |  +-- Pseudunio_auricularius     
     |  |  |              |  |     +--+                              
     |  |  |              |  |        +-- Pseudunio_homsensis        
     |  |  |              +--+                                       
     |  |  |                 |  +-------- Margaritifera_falcata      
     |  |  |                 |  |                                    
     |  |  |                 |  |     +-- Margaritifera_dahurica     
     |  |  |                 |  +-----+                              
     |  |  |                 +--+     +-- Margaritifera_margaritifera
     |  |  |                    |                                    
     |  |  |                    |  +----- Margaritifera_laevis       
     |  |  |                    |  |                                 
     |  |  |                    +--+----- Margaritifera_middendorffi 
     |  |  |                       |                                 
     +--+  |                       |  +-- Margaritifera_hembeli      
        |  |                       +--+                              
        |  |                          +-- Margaritifera_marrianae    
        |  |                                                         
        |  |                          +-- Parreysia_keralaensis      
        |  |                          |                              
        |  |     +--------------------+-- Parreysia_nagpoorensis     
        |  |     |                    |                              
        |  |     |                    +-- Parreysia_plagiosoma       
        |  |     |                                                   
        |  |     |                 +----- Trapezidens_exolescens     
        |  |     |        +--------+                                 
        |  |     |        |        |  +-- Trapezidens_angustior      
        |  |     |        |        +--+                              
        |  |     +--------+           +-- Trapezidens_dolichorhynchus
        |  |     |        |                                          
        |  |     |        |  +----------- Lamellidens_corrianus      
        |  |     |        |  |                                       
        |  |     |        +--+  +-------- Lamellidens_savadiensis    
        |  |     |           |  |                                    
        |  |     |           |  |  +----- Lamellidens_marginalis     
        |  |     |           +--+  |                                 
        |  |     |              |  +----- Lamellidens_ferrugineus    
        |  |     |              |  |                                 
        |  |  +--+              +--+  +-- Lamellidens_mainwaringi__1 
        |  |  |  |                 |  |                              
        |  |  |  |                 |  +-- Lamellidens_mainwaringi__2 
        |  |  |  |                 |  |                              
        |  |  |  |                 +--+-- Lamellidens_lamellatus     
        |  |  |  |                    |                              
        |  |  |  |                    +-- Lamellidens_generosus      
        |  |  |  |                    |                              
        |  |  |  |                    +-- Lamellidens_jenkinsianus   
        |  |  |  |                                                   
        |  |  |  |                 +----- Leoparreysia_tavoyensis    
        |  |  |  |  +--------------+                                 
        |  |  |  |  |              |  +-- Leoparreysia_canefrii      
        |  |  |  |  |              +--+                              
        |  |  |  |  |                 +-- Leoparreysia_olivacea      
        |  |  |  |  |                                                
        +--+  |  |  |           +-------- Coelatura_aegyptiaca__2    
           |  |  |  |           |                                    
           |  |  |  |           +-------- Nitia_teretiuscula         
           |  |  +--+  +--------+                                    
           |  |     |  |        |  +----- Coelatura_aegyptiaca__1    
           |  |     |  |        |  |                                 
           |  |     |  |        +--+  +-- Coelatura_gabonensis       
           |  |     |  |           |  |                              
           |  |     |  |           +--+-- Coelatura_kunenensis       
           |  |     |  |              |                              
           |  |     |  |              +-- Prisodontopsis_mweruensis  
           |  |     +--+                                             
           |  |        |           +----- Indonaia_shurtleffiana     
           |  |        |           |                                 
           |  |        |           |  +-- Indonaia_bonneaudii        
           |  |        |  +--------+--+                              
           |  |        |  |        |  +-- Indonaia_rugosa            
           |  |        |  |        |                                 
           |  |        |  |        |  +-- Indonaia_andersoniana      
           |  |        |  |        +--+                              
           |  |        +--+           +-- Indonaia_caerulea          
           |  |           |                                          
           |  |           |  +----------- Indochinella_pugio         
           |  |           |  |                                       
           |  |           |  |     +----- Scabies_humilis            
           |  |           +--+  +--+                                 
           |  |              |  |  |  +-- Scabies_phaselus           
           |  |              |  |  +--+                              
           |  |              +--+     +-- Scabies_mandarinus         
           |  |                 |                                    
           |  |                 |  +----- Radiatula_mouhoti          
           |  |                 +--+                                 
           |  |                    |  +-- Radiatula_myitkyinae       
           |  |                    +--+                              
           |  |                       +-- Radiatula_chaudhurii       
           |  |                                                      
           |  |     +-------------------- Chamberlainia_somsakpanhai 
           |  |     |                                                
           |  |     +-------------------- Megalonaias_nervosa        
           |  |     |                                                
           |  |     +-------------------- Uniomerus_carolinianus     
           |  |     |                                                
           |  |     |                 +-- Quadrula_quadrula          
           |  |     +-----------------+                              
           |  |     |                 +-- Quadrula_verrucosa         
           |  |     |                                                
           |  |     |                 +-- Sinohyriopsis_cumingii     
           |  |     +-----------------+                              
           |  |     |                 +-- Sinohyriopsis_schlegelii   
           |  |     |                                                
           |  |     |                 +-- Potomida_littoralis        
           |  |     |           +-----+                              
           |  |     |           |     +-- Pronodularia_japanensis    
           |  |     +-----------+                                    
           |  |     |           |  +----- Lamprotula_leaii           
           |  |     |           +--+                                 
           |  |     |              |  +-- Lamprotula_caveata         
           |  |     |              +--+                              
           |  |     |                 +-- Lamprotula_cornuumlunae    
           |  |     |                                                
           |  |     |           +-------- Sinosolenaia_carinata      
           +--+     |           |                                    
              |     |           |  +----- Gonidea_angulata           
              |     |           +--+                                 
              |     |           |  |  +-- Leguminaia_wheatleyi       
              |     |           |  +--+                              
              |     |           |     +-- Microcondylaea_bonellii    
              |     +-----------+                                    
              |     |           |  +----- Ptychorhynchus_pfisteri    
              |     |           +--+                                 
              |     |           |  |  +-- Parvasolenaia_rivularis    
              |     |           |  +--+                              
              |     |           |     +-- Koreosolenaia_sitgyensis   
              |     |           |                                    
              |     |           |  +----- Sinosolenaia_recognita     
              |     |           +--+                                 
              |     |              |  +-- Sinosolenaia_iridinea      
              |     |              +--+                              
              |     |                 +-- Sinosolenaia_oleivora      
              |     |                                                
              |     |              +----- Indopseudodon_salwenianus  
              |     |        +-----+                                 
              |     |        |     |  +-- Indopseudodon_indawgyiensis
              |     |        |     +--+                              
              |     |        |        +-- Indopseudodon_bogani       
              |     |        |                                       
              |     |        |  +-------- Nyeinchanconcha_nyeinchani 
              |     |        |  |                                    
              |     |        |  +-------- Thaiconcha_callifera       
              |     +--------+  |                                    
              |     |        |  +-------- Sundadontina_cumingii      
              |     |        |  |                                    
              |     |        |  +-------- Sundadontina_tumida__1     
              |     |        |  |                                    
              |     |        |  +-------- Sundadontina_tumida__2     
              |     |        |  |                                    
              |     |        |  |  +----- Pilsbryoconcha_exilis__2   
              |     |        +--+--+                                 
              |     |           |  |  +-- Pilsbryoconcha_carinifera  
              |     |           |  +--+                              
              |     |           |     +-- Pilsbryoconcha_exilis__1   
              |     |           |                                    
              |     |           |  +----- Pseudodon_vagulus          
              |     |           +--+                                 
              |     |           |  |  +-- Pseudodon_mekongi          
              |     |           |  +--+                              
              |     |           |     +-- Pseudodon_vondembuschianus 
              |     |           |                                    
              |     |           |  +----- Bineurus_loeiensis         
              |     |           +--+                                 
              |     |              |  +-- Bineurus_mouhotii          
              |     |              +--+                              
              |     |                 +-- Bineurus_exilis            
              |     |                                                
              |     |     +-------------- Amblema_plicata            
              |     |     |                                          
              |     |     +-------------- Toxolasma_parvum           
              |     |     |                                          
              |     |     |           +-- Obliquaria_reflexa         
              |     |     +-----------+                              
              |     |     |           +-- Plectomerus_dombeyanus     
              |     |     |                                          
              |     |     |  +----------- Elliptoideus_sloatianus    
              |     |     +--+                                       
              |     |     |  |  +-------- Elliptio_complanata        
              |     |     |  +--+                                    
              |     |     |     |  +----- Eurynia_dilatata           
              |     +-----+     +--+                                 
              |     |     |        |  +-- Fusconaia_flava            
              |     |     |        +--+                              
              |     |     |           +-- Pleurobema_sintoxia        
              |     |     |                                          
              |     |     |     +-------- Ellipsaria_lineolata       
              |     |     |     |                                    
              |     |     |     +-------- Potamilus_leptodon         
              |     |     |     |                                    
              |     |     |     +-------- Medionidus_conradicus      
              |     |     |     |                                    
              +-----+     |     +-------- Potamilus_alatus           
                    |     |     |                                    
                    |     |     +-------- Ptychobranchus_subtentus   
                    |     +-----+                                    
                    |           +-------- Truncilla_truncata         
                    |           |                                    
                    |           +-------- Venustaconcha_ellipsiformis
                    |           |                                    
                    |           +-------- Cambarunio_iris            
                    |           |                                    
                    |           +-------- Leaunio_lienosus           
                    |           |                                    
                    |           |  +----- Ortmanniana_ligamentina    
                    |           +--+                                 
                    |              |  +-- Lampsilis_cardium          
                    |              +--+                              
                    |                 +-- Lampsilis_ornata           
                    |                                                
                    |     +-------------- Hyriopsis_desowitzi        
                    |     |                                          
                    |     +-------------- Hyriopsis_myersiana        
                    |     |                                          
                    |     +-------------- Hyriopsis_panhai           
                    |     |                                          
                    |  +--+           +-- Hyriopsis_bialata          
                    |  |  +-----------+                              
                    |  |  |           +-- Hyriopsis_khoratensis      
                    |  |  |                                          
                    |  |  |  +----------- Rectidens_sumatrensis      
                    |  |  |  |                                       
                    |  |  +--+        +-- Ensidens_telus             
                    |  |     |  +-----+                              
                    |  |     |  |     +-- Ensidens_ingallsianus      
                    |  |     +--+                                    
                    |  |        |  +----- Ensidens_sagittarius       
                    |  |        +--+                                 
                    +--+           |  +-- Ensidens_spiculus          
                    |  |           +--+                              
                    |  |              +-- Ensidens_jaculus           
                    |  |                                             
                    |  |     +----------- Physunio_modelli           
                    |  |     |                                       
                    |  |     +----------- Lens                       
                    |  |     |                                       
                    |  |     +----------- Physunio_superbus          
                    |  |     |                                       
                    |  |     |  +-------- Yaukthwa_inlenensis        
                    |  |     +--+                                    
                    |  +-----+  |  +----- Yaukthwa_panhai            
                    |        |  +--+                                 
                    |        |     |  +-- Yaukthwa_nesemanni         
                    |        |     +--+                              
                    |        |        +-- Yaukthwa_rectangularis     
                    |        |                                       
                    |        |        +-- Lens_comptus               
                    |        |  +-----+                              
                    |        |  |     +-- Lens_novoselovi            
                    |        +--+                                    
                    |           |  +----- Lens_eximius               
                    |           +--+                                 
                    |              |  +-- Lens_contradens            
                    |              +--+                              
                    |                 +-- Lens_micropterus           
                    |                                                
                    |  +----------------- Acuticosta_chinensis       
                    |  |                                             
                    |  +----------------- Alasmidonta_marginata      
                    |  |                                             
                    |  +----------------- Anodonta_anatina           
                    |  |                                             
                    |  +----------------- Anemina_arcaeformis        
                    |  |                                             
                    |  +----------------- Anodonta_cygnea            
                    |  |                                             
                    |  +----------------- Cristaria_plicata          
                    |  |                                             
                    |  +----------------- Anodonta_nuttalliana       
                    |  |                                             
                    |  +----------------- Lasmigona_costata          
                    |  |                                             
                    |  +----------------- Lepidodesma_languilati     
                    |  |                                             
                    |  +----------------- Pseudanodonta_complanata   
                    |  |                                             
                    |  +----------------- Pyganodon_grandis          
                    |  |                                             
                    |  +----------------- Simpsonaias_ambigua        
                    |  |                                             
                    |  +----------------- Strophitus_undulatus       
                    |  |                                             
                    |  |              +-- Schistodesmus_lampreyanus  
                    |  +--------------+                              
                    |  |              +-- Schistodesmus_spinosus     
                    |  |                                             
                    |  |              +-- Utterbackia_imbecillis     
                    |  +--------------+                              
                    |  |              +-- Utterbackia_peninsularis   
                    |  |                                             
                    |  |        +-------- Lanceolaria_gladiola       
                    |  +--------+                                    
                    |  |        |  +----- Lanceolaria_grayii         
                    |  |        +--+                                 
                    +--+           |  +-- Lanceolaria_lanceolata     
                       |           +--+                              
                       |              +-- Lanceolaria_triformis      
                       |                                             
                       |        +-------- Unio_tumidus               
                       +--------+                                    
                       |        |  +----- Unio_crassus               
                       |        +--+                                 
                       |           |  +-- Unio_delphinus             
                       |           +--+                              
                       |              +-- Unio_pictorum              
                       |                                             
                       |        +-------- Aculamprotula_tortuosa     
                       |        |                                    
                       +--------+     +-- Aculamprotula_coreana      
                       |        |  +--+                              
                       |        |  |  +-- Aculamprotula_scripta      
                       |        +--+                                 
                       |           |  +-- Aculamprotula_fibrosa      
                       |           +--+                              
                       |              +-- Aculamprotula_tientsinensis
                       |                                             
                       |        +-------- Cuneopsis_heudei           
                       |        |                                    
                       |        +-------- Cuneopsis_celtiformis      
                       |        |                                    
                       +--------+-------- Tchangsinaia_piscicula     
                       |        |                                    
                       |        +-------- Cuneopsis_rufescens        
                       |        |                                    
                       |        |  +----- Nodularia_huana            
                       |        +--+                                 
                       |           |  +-- Nodularia_douglasiae       
                       |           +--+                              
                       |              +-- Nodularia_nuxpersicae      
                       |                                             
                       |  +-------------- Pletholophus_tenuis        
                       |  |                                          
                       +--+           +-- Sinanodonta_angula         
                          |  +--------+                              
                          |  |        +-- Sinanodonta_lucida         
                          +--+                                       
                             |  +-------- Sinanodonta_woodiana       
                             +--+                                    
                                |  +----- Sinanodonta_jourdyi        
                                +--+                                 
                                   |  +-- Sinanodonta_elliptica      
                                   +--+                              
                                      +-- Sinanodonta_pacifica       

Page updated: 11:09:25 Wed 27 Nov 2024

 
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