The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

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Bivalvia | Unionoida | Unionoidea | Margaritiferidae | Gibbosulinae

genus Gibbosula Simpson, 1900: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 70 cladograms in 38 publications
maximum number of terminals in an analysis = 3, mean = 1.3

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included terminals = 3, mean = 2.17
proportion of cladograms that support basic monophyly = 0.78

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_crassa **        
  +--+                           
     +-- Gibbosula_laosensis **     

Lyubas et al., 2019 [Publ 3963], fig. 10

  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **        

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

  +----- Gibbosula_rochechouartii **
--+                              
  |  +-- Gibbosula_laosensis **     
  +--+                           
     +-- Gibbosula_crassa **        

Huang et al., 2019 [Publ 3843], fig. S4

  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Cumberlandia            
     +--+                              
        |  +-- Pseudunio               
        +--+                           
           +-- Margaritifera           

Wu et al., 2020 [Publ 4121], fig. 2

  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Huang et al., 2019 [Publ 3843], fig. S3

  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio               

Huang et al., 2018 [Publ 3828], fig. 3_5

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **     

Wu et al., 2021 [Publ 4366], fig. 4

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **        

Huang et al., 2019 [Publ 3843], fig. 2

  +-------- Gibbosula_rochechouartii **
  |                                 
--+  +----- Gibbosula_laosensis **     
  |  |                              
  +--+  +-- Cumberlandia            
     |  |                           
     +--+-- Pseudunio               
        |                           
        +-- Margaritifera           

Sano et al., 2020 [Publ 4138], fig. 3

  +-- Gibbosula_laosensis **     
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 2

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **     

Wu et al., 2021 [Publ 4366], fig. 3-BI

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **        

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

  +-- Gibbosula_crassa **        
--+                           
  +-- Gibbosula_rochechouartii **

Sano et al., 2020 [Publ 4138], fig. 4

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_laosensis **     

Wu et al., 2021 [Publ 4366], fig. 3-ML

  +-- Gibbosula_rochechouartii **
--+                           
  +-- Gibbosula_crassa **        

Wu et al., 2020 [Publ 4121], fig. S1

  +----------- Gibbosula_rochechouartii **
--+                                    
  |  +-------- Gibbosula_laosensis **     
  +--+                                 
     |  +----- Margaritifera           
     +--+                              
        |  +-- Cumberlandia            
        +--+                           
           +-- Pseudunio               

Sano et al., 2020 [Publ 4138], fig. 1

-- Gibbosula_rochechouartii

Dai et al., 2023 [Publ 4285], fig. 5

-- Gibbosula_rochechouartii

Wu et al., 2020 [Publ 4121], fig. 3

-- Gibbosula_rochechouartii

Dai et al., 2025 [Publ 4440], fig. 3

-- Gibbosula_rochechouartii

Huang et al., 2019 [Publ 3843], fig. S1

-- Gibbosula_rochechouartii

Konopleva et al., 2023 [Publ 4269], fig. S1_4

-- Gibbosula_laosensis

Dai et al., 2025 [Publ 4440], fig. 2

-- Gibbosula_rochechouartii

Dai et al., 2024 [Publ 4386], fig. 5-ML

-- Gibbosula_laosensis

Dai et al., 2024 [Publ 4386], fig. 5-BI

-- Gibbosula_laosensis

Wu et al., 2021 [Publ 4366], fig. 6

-- Gibbosula_laosensis

Huang et al., 2019 [Publ 3843], fig. 3

-- Gibbosula_rochechouartii

Konopleva et al., 2019 [Publ 3900], fig. 1

-- Gibbosula_laosensis

Bolotov et al., 2016 [Publ 3743], fig. 4

-- Gibbosula_laosensis

Bolotov et al., 2022 [Publ 4250], fig. 8

-- Gibbosula_laosensis

Bolotov et al., 2020 [Publ 4100], fig. 1

-- Gibbosula_laosensis

Wu et al., 2024 [Publ 4379], fig. 5

-- Gibbosula_crassa

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

-- Gibbosula_laosensis

Dai et al., 2025 [Publ 4441], fig. 1

-- Gibbosula_laosensis

Bogan et al., 2023 [Publ 4295], fig. 5

-- Gibbosula_crassa

Dai et al., 2024 [Publ 4389], fig. 3

-- Gibbosula_laosensis

Chen et al., 2023 [Publ 4302], fig. 1

-- Gibbosula_rochechouartii

Zhou et al., 2007 [Publ 2658], fig. 1

-- Gibbosula_rochechouartii

Dai et al., 2024 [Publ 4385], fig. 3-ML

-- Gibbosula_laosensis

Dai et al., 2023 [Publ 4285], fig. 3-BI

-- Gibbosula_laosensis

Vikhrev et al., 2018 [Publ 3812], fig. 3

-- Gibbosula_laosensis

Bogan et al., 2023 [Publ 4295], fig. 4

-- Gibbosula_crassa

Guerra et al., 2019 [Publ 4168], fig. 1

-- Gibbosula_crassa

Bogan et al., 2023 [Publ 4295], fig. 6

-- Gibbosula_crassa

Bolotov et al., 2017 [Publ 3761], fig. S1

-- Gibbosula_laosensis

Chen et al., 2020 [Publ 4170], fig. 1

-- Gibbosula_crassa

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

-- Gibbosula_laosensis

Araujo et al., 2016 [Publ 3746], fig. 2a

-- Gibbosula_laosensis

Araujo et al., 2016 [Publ 3746], fig. 1_2b

-- Gibbosula_laosensis

Fukata & Iigo, 2020 [Publ 4106], fig. 1

-- Gibbosula_crassa

Huang et al., 2018 [Publ 3828], fig. 6

-- Gibbosula_rochechouartii

Bolotov et al., 2017 [Publ 3761], fig. S2

-- Gibbosula_laosensis

Chen et al., 2023 [Publ 4302], fig. 2

-- Gibbosula_rochechouartii

Wu et al., 2019 [Publ 3864], fig. 3

-- Gibbosula_rochechouartii

Bolotov et al., 2016 [Publ 3743], fig. 2

-- Gibbosula_laosensis

Dai et al., 2025 [Publ 4440], fig. 1

-- Gibbosula_rochechouartii

Guerra et al., 2019 [Publ 4168], fig. 2

-- Gibbosula_crassa

Bolotov et al., 2016 [Publ 3732], fig. 2

-- Gibbosula_laosensis

Huang et al., 2019 [Publ 3843], fig. 1

-- Gibbosula_rochechouartii

Bespalaya et al., 2018 [Publ 3811], fig. 3

-- Gibbosula_laosensis

Wu et al., 2020 [Publ 4147], fig. 4

-- Gibbosula_rochechouartii

Wu et al., 2022 [Publ 4238], fig. 3

-- Gibbosula_rochechouartii

Dai et al., 2024 [Publ 4385], fig. 3-BI

-- Gibbosula_laosensis

Dai et al., 2023 [Publ 4285], fig. 3-ML

-- Gibbosula_laosensis

Vikhrev et al., 2017 [Publ 3810], fig. 4

-- Gibbosula_laosensis

Bogan et al., 2023 [Publ 4295], fig. 3

-- Gibbosula_crassa

Dai et al., 2024 [Publ 4358], fig. 4

-- Gibbosula_rochechouartii

Wu et al., 2021 [Publ 4366], fig. 2_I

-- Gibbosula_laosensis

Page updated: 08:16:51 Sun 07 Sep 2025