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Page last updated
Sun 07 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validgen_cld&id=68

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Velesunioninae

genus Lortiella Modell, 1942: cladomics
     3 valid species

Cladomics Summary

number of species = 3

analyzed in 22 cladograms in 13 publications
maximum number of terminals in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = 0, mean = 0
proportion of cladograms that support basic monophyly = 0

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 6 4 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------- Westralunio        
  |                               
  |     +----- Alathyria_jacksoni 
--+  +--+                         
  |  |  |  +-- Velesunio          
  |  |  +--+                      
  +--+     +-- Alathyria_profuga  
     |                            
     |  +----- Alathyria_pertexta 
     +--+                         
        |  +-- Microdontia        
        +--+                      
           +-- Lortiella_froggatti **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 198 6 4 -- 
     +----- Velesunio          
  +--+                         
  |  |  +-- Alathyria_jacksoni 
  |  +--+                      
--+     +-- Alathyria_profuga  
  |                            
  |  +----- Alathyria_pertexta 
  +--+                         
     |  +-- Lortiella_froggatti **
     +--+                      
        +-- Microdontia

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 -- 
  +-- Velesunio       
--+                   
  +-- Lortiella_rugata **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 19 5 4 -- 
  +----------- Velesunio_ambiguus
--+                              
  |  +-------- Alathyria         
  +--+                           
     |  +----- Velesunio_angasi  
     +--+                        
        |  +-- Lortiella **         
        +--+                     
           +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 196 6 4 -- 
     +----- Velesunio          
  +--+                         
  |  |  +-- Alathyria_jacksoni 
  |  +--+                      
--+     +-- Alathyria_profuga  
  |                            
  |  +----- Alathyria_pertexta 
  +--+                         
     |  +-- Lortiella_froggatti **
     +--+                      
        +-- Microdontia

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 -- 
  +-- Lortiella_rugata **
--+                   
  +-- Velesunio

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 2 1 -- 
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 198 6 4 -- 
     +----- Alathyria_jacksoni 
  +--+                         
  |  |  +-- Alathyria_profuga  
  |  +--+                      
--+     +-- Velesunio          
  |                            
  |  +----- Alathyria_pertexta 
  +--+                         
     |  +-- Lortiella_froggatti **
     +--+                      
        +-- Microdontia

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 -- 
-- Lortiella_rugata

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 29 5 3 -- 
     +----- Microdontia        
  +--+                         
  |  |  +-- Alathyria_pertexta 
  |  +--+                      
--+     +-- Lortiella_froggatti **
  |                            
  |  +----- Velesunio          
  +--+                         
     |  +-- Alathyria_jacksoni 
     +--+                      
        +-- Alathyria_profuga

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 2 1 -- 
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 5 3 -- 
     +----- Alathyria_pertexta 
  +--+                         
  |  |  +-- Microdontia        
  |  +--+                      
--+     +-- Lortiella_froggatti **
  |                            
  |  +----- Velesunio          
  +--+                         
     |  +-- Alathyria_jacksoni 
     +--+                      
        +-- Alathyria_profuga

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 -- 
  +-- Velesunio
--+            
  +-- Lortiella **

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 2 1 -- 
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio         
  +--+                     
     +-- Lortiella **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 -- 
-- Lortiella_rugata

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 5 3 -- 
     +----- Velesunio          
  +--+                         
  |  |  +-- Alathyria_jacksoni 
  |  +--+                      
--+     +-- Alathyria_profuga  
  |                            
  |  +----- Microdontia        
  +--+                         
     |  +-- Alathyria_pertexta 
     +--+                      
        +-- Lortiella_froggatti **

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 29 5 3 -- 
     +----- Microdontia        
  +--+                         
  |  |  +-- Lortiella_froggatti **
  |  +--+                      
--+     +-- Alathyria_pertexta 
  |                            
  |  +----- Velesunio          
  +--+                         
     |  +-- Alathyria_jacksoni 
     +--+                      
        +-- Alathyria_profuga

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 35 4 2 -- 
-- 2889

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iT tT bi riS oT roT roS iD
1 1 0 1 19 5 4 -- 
  +----------- Velesunio_ambiguus
--+                              
  |  +-------- Microdontia       
  +--+                           
     |  +----- Lortiella **         
     +--+                        
        |  +-- Alathyria         
        +--+                     
           +-- Velesunio_angasi

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 2 1 -- 
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 -- 
  +-- Velesunio       
--+                   
  +-- Lortiella_rugata **

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 2 1 -- 
  +----- Velesunio_ambiguus
--+                        
  |  +-- Velesunio_angasi  
  +--+                     
     +-- Lortiella_rugata **

Page updated: 08:16:37 Sun 07 Sep 2025
 
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