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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Margaritiferidae | Margaritiferinae

genus Pseudunio Haas, 1910: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 74 cladograms in 39 publications
maximum number of terminals in an analysis = 5, mean = 1.5

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 28
maximum number of included terminals = 5, mean = 2.32
proportion of cladograms that support basic monophyly = 0.96


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Lyubas et al., 2019 [Publ 3963], fig. 10

iT tT bi riS oT roT roS iD
5 5 1 5 11 8 2 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------- Margaritifera             
  |                                      
--+  +-------- Cumberlandia              
  |  |                                   
  +--+  +----- Pseudunio_marocanus **       
     |  |                                
     +--+  +-- Pseudunio_auricularius **    
        |  |                             
        |  +-- Pseudunio_flabellata **      
        +--+                             
           +-- Pseudunio_flabellatiformis **
           |                             
           +-- Pseudunio_homsensis **       

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 195 8 2 2.0
  +----------- Cumberlandia          
--+                                  
  |  +-------- Margaritifera         
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Vikhrev et al., 2018 [Publ 3812], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 10 7 2 2.0
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 193 8 2 2.0
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 25 8 2 2.0
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 25 8 2 2.0
  +----------- Margaritifera         
--+                                  
  |  +-------- Cumberlandia          
  +--+                               
     |  +----- Pseudunio_marocanus **   
     +--+                            
        |  +-- Pseudunio_auricularius **
        +--+                         
           +-- Pseudunio_homsensis **   

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 195 8 2 2.0
  +-------- Cumberlandia          
  |                               
--+-------- Margaritifera         
  |                               
  |  +----- Pseudunio_marocanus **   
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_homsensis **   

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 67 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 137 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 67 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 67 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Inoue et al., 2013 [Publ 3579], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 7 6 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Riccardi et al., 2020 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 18 0 0 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Araujo et al., 2016 [Publ 3746], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 9 8 2 --
  +-- Pseudunio_marocanus **   
--+                         
  +-- Pseudunio_auricularius **

Bolotov et al., 2015 [Publ 3646], fig. 2B

iT tT bi riS oT roT roS iD
2 2 0 2 7 6 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 28 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 28 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 39 4 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Araujo et al., 2009 [Publ 2395], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 6 5 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 32 8 2 --
  +-------- Margaritifera         
--+                               
  |  +----- Cumberlandia          
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Huang et al., 2018 [Publ 3828], fig. 3_5

iT tT bi riS oT roT roS iD
2 2 0 2 27 8 2 --
  +-------- Margaritifera         
--+                               
  |  +----- Cumberlandia          
  +--+                            
     |  +-- Pseudunio_auricularius **
     +--+                         
        +-- Pseudunio_marocanus **   

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
2 2 0 2 74 7 2 --
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Pseudunio_marocanus **   

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
2 2 0 2 74 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 2 137 7 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Araujo et al., 2016 [Publ 3746], fig. 1_2b

iT tT bi riS oT roT roS iD
2 2 0 2 11 8 2 --
  +-------- Margaritifera         
--+                               
  |  +----- Cumberlandia          
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 32 8 2 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Pseudunio_marocanus **   

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 39 4 2 --
  +-------- Cumberlandia          
--+                               
  |  +----- Margaritifera         
  +--+                            
     |  +-- Pseudunio_marocanus **   
     +--+                         
        +-- Pseudunio_auricularius **

Takeuchi et al., 2015 [Publ 3661], fig. 2

iT tT bi riS oT roT roS iD
2 8 0 2 8 6 2 9.0
  +-------- Pseudunio_marocanus **   
--+                               
  |  +----- Pseudunio_auricularius **
  +--+                            
     |  +-- Cumberlandia          
     +--+                         
        +-- Margaritifera         

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 31 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 --
-- Pseudunio_marocanus

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 2 --
  +----- Pseudunio_marocanus **
--+                         
  |  +-- Margaritifera      
  +--+                      
     +-- Cumberlandia       

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
1 1 0 1 22 0 0 --
-- Pseudunio_marocanus

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 18 0 0 --
-- Pseudunio_marocanus

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 42 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 5 2 --
  +----- Margaritifera      
--+                         
  |  +-- Pseudunio_homsensis **
  +--+                      
     +-- Cumberlandia       

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 2 --
  +----- Cumberlandia          
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Margaritifera         

Araujo et al., 2009 [Publ 2589], fig. 7B

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Pseudunio_auricularius

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Araujo et al., 2018 [Publ 3779], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 0 0 --
-- Pseudunio_auricularius

Huff et al., 2004 [Publ 2697], fig. 5-6

iT tT bi riS oT roT roS iD
1 1 0 1 11 6 2 --
  +----- Margaritifera         
--+                            
  |  +-- Pseudunio_auricularius **
  +--+                         
     +-- Cumberlandia          

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Cumberlandia       

Bashê, 2022 [Publ 4265], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Pseudunio_homsensis

Välilä et al., 2017 [Publ 3793], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Margaritifera      
  +--+                      
     +-- Pseudunio_marocanus **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Bolotov et al., 2015 [Publ 3646], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 8 7 2 --
  +----------- Margaritifera_margaritifera
  |                                       
  |     +----- Margaritifera_dahurica__1  
  |     |                                 
  +-----+----- Margaritifera_dahurica__2  
--+     |                                 
  |     |  +-- Pseudunio_auricularius **     
  |     +--+                              
  |        +-- Cumberlandia               
  |                                       
  |  +-------- Margaritifera_falcata      
  +--+                                    
     |  +----- Margaritifera_laevis       
     +--+                                 
        |  +-- Margaritifera_middendorffi 
        +--+                              
           +-- Margaritifera_marrianae    

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 16 0 0 --
-- Pseudunio_auricularius

Huff et al., 2004 [Publ 2697], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 6 2 --
  +-- Pseudunio_auricularius **
  |                         
--+-- Cumberlandia          
  |                         
  +-- Margaritifera         

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 2 2 --
  +----- Margaritifera      
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Cumberlandia       

Huff et al., 2004 [Publ 2697], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 11 6 2 --
  +-------- Margaritifera_dahurica        
  |                                       
  +-------- Margaritifera_margaritifera__1
  |                                       
  +-------- Margaritifera_margaritifera__2
  |                                       
--+-------- Margaritifera_margaritifera__3
  |                                       
  |     +-- Pseudunio_auricularius **        
  +-----+                                 
  |     +-- Cumberlandia                  
  |                                       
  |  +----- Margaritifera_falcata         
  +--+                                    
     |  +-- Margaritifera_middendorffi    
     +--+                                 
        +-- Margaritifera_marrianae       

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 18 0 0 --
-- Pseudunio_marocanus

Machordom et al., 2003 [Publ 2688], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 1 1 --
  +-- Margaritifera         
--+                         
  +-- Pseudunio_auricularius **

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 5 2 --
  +----- Margaritifera      
--+                         
  |  +-- Cumberlandia       
  +--+                      
     +-- Pseudunio_homsensis **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Cumberlandia       

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Takeuchi et al., 2015 [Publ 3661], fig. 3

iT tT bi riS oT roT roS iD
1 8 0 2 9 7 2 12.0
  +----- Pseudunio_auricularius__1 **
  |                               
--+----- Pseudunio_auricularius__2 **
  |                               
  |  +-- Cumberlandia             
  +--+                            
     +-- Margaritifera            

Huff et al., 2004 [Publ 2697], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 7 5 2 --
  +-------- Margaritifera_margaritifera  
  |                                      
  |  +----- Cumberlandia                 
--+  |                                   
  |  |  +-- Pseudunio_auricularius **       
  |  |  |                                
  +--+  +-- Margaritifera_dahurica       
     |  |                                
     |  +-- Margaritifera_middendorffi__1
     |  |                                
     |  +-- Margaritifera_middendorffi__2
     +--+                                
        +-- Margaritifera_falcata__1     
        |                                
        +-- Margaritifera_falcata__2     
        |                                
        +-- Margaritifera_falcata__3     
        |                                
        +-- Margaritifera_falcata__4     

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Pseudunio_marocanus

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 2 1 --
  +-- Pseudunio_marocanus **
--+                      
  +-- Margaritifera      

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 9 4 2 --
  +----- Cumberlandia       
--+                         
  |  +-- Pseudunio_marocanus **
  +--+                      
     +-- Margaritifera      

Machordom et al., 2003 [Publ 2688], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 2 --
  +----- Cumberlandia          
--+                            
  |  +-- Margaritifera         
  +--+                         
     +-- Pseudunio_auricularius **

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
  +-- Pseudunio_auricularius **
--+                         
  +-- Margaritifera         

Araujo et al., 2009 [Publ 2589], fig. 7A

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Pseudunio_auricularius

Araujo et al., 2009 [Publ 2589], fig. 7C

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Pseudunio_auricularius

Page updated: 14:55:48 Wed 04 Dec 2024

 
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