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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe COELATURINI Modell, 1942: cladomics
     10 genera

Cladomics Summary

number of species = 41
number of subtaxa (rank: genus) = 10

analyzed in 48 cladograms in 31 publications
maximum number of terminals in an analysis = 27, mean = 3.54
maximum number of subtaxa (genus) in an analysis = 7, mean = 2.1

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 26
maximum number of included terminals = 27, mean = 5.69
maximum number of subtaxa (genus) = 7, mean = 3.04
proportion of cladograms that support basic monophyly = 1.0

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 26
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 27, mean = 5.69
maximum number of subtaxa (genus) = 7, mean = 3.04
proportion of cladograms that support rigorous monophyly = 1.0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iT tT bi riS oT roT roS iD
27 27 16 7 9 9 2 0.0

*** Top Monophyly Cladogram ***

  +-------------------------------------- Leoparreysiini               
  |                                                                    
--+  +----------------------------------- Indochinellini               
  |  |                                                                 
  +--+  +-------------------------------- Grandidieria **                 
     |  |                                                              
     |  |                             +-- Pseudospatha **                 
     +--+  +--------------------------+                                
        |  |                          +-- Moncetia **                     
        |  |                                                           
        |  |  +-------------------------- Nitia **                        
        +--+  |                                                        
           |  |                       +-- Coelatura_aegyptiaca **         
           |  |                       |                                
           |  |                       +-- Coelatura_alluaudi **           
           +--+                       |                                
              |  +--------------------+-- Coelatura_bakeri **             
              |  |                    |                                
              |  |                    +-- Coelatura_hauttecoeuri **       
              |  |                    |                                
              |  |                    +-- Coelatura_stuhlmanni **         
              +--+                                                     
                 |                    +-- Coelatura_leopoldvillensis **   
                 |  +-----------------+                                
                 |  |                 +-- Coelatura_mesafricana **        
                 |  |                                                  
                 |  |                 +-- Coelatura_horei **              
                 +--+                 |                                
                    |  +--------------+-- Coelatura_ratidota **           
                    |  |              |                                
                    |  |              +-- Coelatura **                    
                    +--+                                               
                       |  +-------------- Coelatura_gabonensis **         
                       |  |                                            
                       |  +-------------- Coelatura_stanleyvillensis **   
                       +--+                                            
                          |  +----------- Coelatura_kunenensis **         
                          |  |                                         
                          |  |     +----- Prisodontopsis_aviculaeformis **
                          +--+  +--+                                   
                             |  |  |  +-- Coelatura_briarti **            
                             |  |  +--+                                
                             |  |     +-- Prisodontopsis_mweruensis **    
                             +--+                                      
                                |     +-- Coelatura_choziensis **         
                                |     |                                
                                |     +-- Coelatura_hypsiprymna **        
                                |     |                                
                                +-----+-- Coelatura_luapulaensis **       
                                      |                                
                                      +-- Coelatura_mossambicensis **     
                                      |                                
                                      +-- Nyassunio **                    

Bolotov et al., 2024 [Publ 4369], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 58 58 4 23.0

*** Top Sister-Group Cladogram ***

  +----------------- Lamellidentini      
  |                                      
--+              +-- Leoparreysiini      
  |  +-----------+                       
  |  |           +-- Parreysiini         
  +--+                                   
     |  +----------- Indochinellini      
     +--+                                
        |  +-------- Nitia **               
        +--+                             
           |  +----- Coelatura_aegyptiaca **
           +--+                          
              |  +-- Coelatura_kunenensis **
              +--+                       
                 +-- Nyassunio **           

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iT tT bi riS oT roT roS iD
27 27 16 7 9 9 2 5.0
  +-------------------------------- Leoparreysiini               
  |                                                              
  |  +----------------------------- Indochinellini               
--+  |                                                           
  |  |                       +----- Grandidieria **                 
  |  |  +--------------------+                                   
  +--+  |                    |  +-- Pseudospatha **                 
     |  |                    +--+                                
     |  |                       +-- Moncetia **                     
     |  |                                                        
     +--+  +----------------------- Nitia **                        
        |  |                                                     
        |  |                    +-- Coelatura_aegyptiaca **         
        |  |                    |                                
        |  |                    +-- Coelatura_alluaudi **           
        +--+                    |                                
           |  +-----------------+-- Coelatura_bakeri **             
           |  |                 |                                
           |  |                 +-- Coelatura_hauttecoeuri **       
           |  |                 |                                
           |  |                 +-- Coelatura_stuhlmanni **         
           +--+                                                  
              |                 +-- Coelatura_leopoldvillensis **   
              |  +--------------+                                
              |  |              +-- Coelatura_mesafricana **        
              |  |                                               
              |  |              +-- Coelatura_horei **              
              +--+              |                                
                 |  +-----------+-- Coelatura_ratidota **           
                 |  |           |                                
                 |  |           +-- Coelatura **                    
                 |  |                                            
                 +--+           +-- Coelatura_gabonensis **         
                    |  +--------+                                
                    |  |        +-- Coelatura_stanleyvillensis **   
                    |  |                                         
                    +--+  +-------- Coelatura_kunenensis **         
                       |  |                                      
                       |  |     +-- Coelatura_briarti **            
                       |  |     |                                
                       +--+  +--+-- Prisodontopsis_mweruensis **    
                          |  |  |                                
                          |  |  +-- Prisodontopsis_aviculaeformis **
                          |  |                                   
                          +--+  +-- Coelatura_choziensis **         
                             |  |                                
                             |  +-- Coelatura_hypsiprymna **        
                             |  |                                
                             +--+-- Coelatura_luapulaensis **       
                                |                                
                                +-- Coelatura_mossambicensis **     
                                |                                
                                +-- Nyassunio **                    

Graf et al., 2014 [Publ 3591], fig. 3

iT tT bi riS oT roT roS iD
8 8 11 3 3 1 1 24.0
  +----------------------- Leoparreysiini           
  |                                                 
--+  +-------------------- Nitia **                    
  |  |                                              
  +--+  +----------------- Coelatura_aegyptiaca__2 **  
     |  |                                           
     +--+  +-------------- Coelatura_aegyptiaca__1 **  
        |  |                                        
        +--+  +----------- Coelatura_gabonensis **     
           |  |                                     
           +--+  +-------- Coelatura_kunenensis **     
              |  |                                  
              |  |  +----- Prisodontopsis **           
              +--+  |                               
                 |  |  +-- Coelatura_choziensis__1 **  
                 |  |  |                            
                 +--+  +-- Coelatura_choziensis__2 **  
                    |  |                            
                    |  +-- Coelatura_luapulaensis__1 **
                    |  |                            
                    +--+-- Coelatura_luapulaensis__2 **
                       |                            
                       +-- Coelatura_luapulaensis__3 **
                       |                            
                       +-- Coelatura_luapulaensis__4 **
                       |                            
                       +-- Coelatura_luapulaensis__5 **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
8 8 0 3 117 30 4 26.0
           +-------------- Trapezidens                  
           |                                            
  +--------+  +----------- Lamellidens_lamellatus       
  |        |  |                                         
  |        +--+  +-------- Lamellidens_jenkinsianus     
  |           |  |                                      
  |           |  |  +----- Lamellidens_generosus        
  |           +--+  |                                   
  |              |  +----- Lamellidens_mainwaringi      
  |              |  |                                   
  |              +--+  +-- Lamellidens_ferrugineus      
  |                 +--+                                
--+                 |  +-- Lamellidens_marginalis       
  |                 |                                   
  |                 |  +-- Lamellidens_savadiensis      
  |                 +--+                                
  |                    +-- Lamellidens_corrianus__1     
  |                                                     
  |  +-------------------- Parreysiini                  
  |  |                                                  
  |  |  +----------------- Leoparreysiini               
  |  |  |                                               
  |  |  |              +-- Nitia **                        
  +--+  |              |                                
     |  |              +-- Coelatura_gabonensis **         
     |  |              |                                
     |  |              +-- Prisodontopsis_mweruensis **    
     |  |              |                                
     +--+              +-- Coelatura_kunenensis **         
        |              |                                
        |  +-----------+-- Coelatura_aegyptiaca__1 **      
        |  |           |                                
        |  |           +-- Coelatura_aegyptiaca__2 **      
        |  |           |                                
        |  |           +-- Prisodontopsis_aviculaeformis **
        |  |           |                                
        |  |           +-- Coelatura_luapulaensis **       
        +--+           |                                
           |           +-- Coelatura_choziensis **         
           |                                            
           |        +----- Indochinella                 
           |  +-----+                                   
           |  |     |  +-- Scabies                      
           |  |     +--+                                
           +--+        +-- Radiatula                    
              |                                         
              |  +-------- Indonaia_shurtleffiana       
              |  |                                      
              +--+  +----- Indonaia_rugosa              
                 |  |                                   
                 |  |  +-- Indonaia_andersoniana        
                 +--+--+                                
                    |  +-- Indonaia_caerulea            
                    |                                   
                    |  +-- Indonaia_bonneaudii          
                    +--+                                
                       +-- Lamellidens_corrianus__2     

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 5 2 3 193 29 4 24.0
  +----------------- Parreysiini            
  |                                         
  +----------------- Lamellidentini         
--+                                         
  |  +-------------- Leoparreysiini         
  |  |                                      
  +--+  +----------- Indochinellini         
     |  |                                   
     +--+  +-------- Coelatura_aegyptiaca__2 **
        |  |                                
        |  +-------- Nitia **                  
        +--+                                
           |  +----- Coelatura_aegyptiaca__1 **
           |  |                             
           +--+  +-- Coelatura_gabonensis **   
              |  |                          
              +--+-- Coelatura_kunenensis **   
                 |                          
                 +-- Prisodontopsis **         

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 3 193 29 4 25.0
  +-------------------- Lamellidentini         
  |                                            
--+  +----------------- Parreysiini            
  |  |                                         
  +--+  +-------------- Leoparreysiini         
     |  |                                      
     +--+  +----------- Indochinellini         
        |  |                                   
        +--+  +-------- Coelatura_aegyptiaca__2 **
           |  |                                
           +--+  +----- Coelatura_aegyptiaca__1 **
              |  |                             
              |  +----- Nitia **                  
              +--+                             
                 |  +-- Coelatura_gabonensis **   
                 |  |                          
                 +--+-- Coelatura_kunenensis **   
                    |                          
                    +-- Prisodontopsis **         

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 5 4 3 191 29 4 27.0
  +----------------------- Lamellidentini         
  |                                               
--+  +-------------------- Parreysiini            
  |  |                                            
  +--+  +----------------- Leoparreysiini         
     |  |                                         
     +--+  +-------------- Indochinellini         
        |  |                                      
        +--+  +----------- Coelatura_aegyptiaca__2 **
           |  |                                   
           +--+        +-- Coelatura_aegyptiaca__1 **
              |  +-----+                          
              |  |     +-- Nitia **                  
              +--+                                
                 |  +----- Coelatura_kunenensis **   
                 +--+                             
                    |  +-- Coelatura_gabonensis **   
                    +--+                          
                       +-- Prisodontopsis **         

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
5 5 4 3 134 24 4 23.0
                       +-- Parreysiini            
  +--------------------+                          
  |                    +-- Lamellidentini         
--+                                               
  |  +-------------------- Leoparreysiini         
  +--+                                            
     |  +----------------- Indochinellini         
     +--+                                         
        |  +-------------- Nitia **                  
        +--+                                      
           |  +----------- Coelatura_aegyptiaca__1 **
           +--+                                   
              |  +-------- Coelatura_aegyptiaca__2 **
              +--+                                
                 |  +----- Coelatura_gabonensis **   
                 +--+                             
                    |  +-- Coelatura_kunenensis **   
                    +--+                          
                       +-- Prisodontopsis **         

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
5 5 4 3 134 24 4 25.0
  +-------------------------- Lamellidentini         
--+                                                  
  |  +----------------------- Parreysiini            
  +--+                                               
     |  +-------------------- Leoparreysiini         
     +--+                                            
        |  +----------------- Indochinellini         
        +--+                                         
           |  +-------------- Nitia **                  
           +--+                                      
              |  +----------- Coelatura_aegyptiaca__2 **
              +--+                                   
                 |  +-------- Coelatura_aegyptiaca__1 **
                 +--+                                
                    |  +----- Prisodontopsis **         
                    +--+                             
                       |  +-- Coelatura_gabonensis **   
                       +--+                          
                          +-- Coelatura_kunenensis **   

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
4 4 3 3 32 5 3 24.0
  +-------------------- Lamellidentini         
--+                                            
  |  +----------------- Leoparreysiini         
  +--+                                         
     |  +-------------- Indochinellini         
     +--+                                      
        |  +----------- Nitia **                  
        +--+                                   
           |  +-------- Coelatura_aegyptiaca__2 **
           +--+                                
              |  +----- Coelatura_aegyptiaca__1 **
              +--+                             
                 |  +-- Coelatura_gabonensis **   
                 +--+                          
                    +-- Prisodontopsis **         

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
4 4 2 3 54 50 4 23.0
  +-------------------- Parreysiini         
--+                                         
  |  +----------------- Lamellidentini      
  +--+                                      
     |  +-------------- Leoparreysiini      
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Coelatura_kunenensis **
                 +--+                       
                    +-- Nyassunio **           

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
4 4 2 3 32 5 3 24.0
  +----------------- Lamellidentini         
  |                                         
--+  +-------------- Leoparreysiini         
  |  |                                      
  +--+  +----------- Indochinellini         
     |  |                                   
     +--+  +-------- Nitia **                  
        |  |                                
        +--+  +----- Coelatura_aegyptiaca__1 **
           |  |                             
           +--+----- Coelatura_aegyptiaca__2 **
              |                             
              |  +-- Coelatura_gabonensis **   
              +--+                          
                 +-- Prisodontopsis **         

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 57 53 4 23.0
  +-------------------- Lamellidentini      
--+                                         
  |  +----------------- Parreysiini         
  +--+                                      
     |  +-------------- Leoparreysiini      
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Nyassunio **           
                 +--+                       
                    +-- Coelatura_kunenensis **

Bolotov et al., 2024 [Publ 4369], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 3 59 58 4 23.0
  +-------------------- Parreysia           
  |                                         
--+                 +-- Leoparreysiini      
  |  +--------------+                       
  |  |              +-- Lamellidentini      
  +--+                                      
     |  +-------------- Assamnaia           
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Nyassunio **           
                 +--+                       
                    +-- Coelatura_kunenensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 54 50 4 23.0
  +----------------- Lamellidentini      
  |                                      
--+              +-- Parreysiini         
  |  +-----------+                       
  |  |           +-- Leoparreysiini      
  +--+                                   
     |  +----------- Indochinellini      
     +--+                                
        |  +-------- Nitia **               
        +--+                             
           |  +----- Coelatura_aegyptiaca **
           +--+                          
              |  +-- Coelatura_kunenensis **
              +--+                       
                 +-- Nyassunio **           

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
4 4 0 3 57 53 4 25.0
  +-------------- Parreysiini         
  |                                   
--+  +----------- Lamellidentini      
  |  |                                
  +--+  +-------- Leoparreysiini      
     |  |                             
     +--+  +----- Indochinellini      
        |  |                          
        +--+  +-- Coelatura_aegyptiaca **
           |  |                       
           |  +-- Coelatura_kunenensis **
           +--+                       
              +-- Nitia **               
              |                       
              +-- Nyassunio **           

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 3 54 50 4 23.0
  +-------------------- Lamellidentini      
--+                                         
  |  +----------------- Parreysiini         
  +--+                                      
     |  +-------------- Leoparreysiini      
     +--+                                   
        |  +----------- Indochinellini      
        +--+                                
           |  +-------- Nitia **               
           +--+                             
              |  +----- Coelatura_aegyptiaca **
              +--+                          
                 |  +-- Coelatura_kunenensis **
                 +--+                       
                    +-- Nyassunio **           

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 25 24 4 24.0
  +----------------- Lamellidentini
--+                                
  |  +-------------- Parreysiini   
  +--+                             
     |  +----------- Leoparreysiini
     +--+                          
        |  +-------- Indochinellini
        +--+                       
           |  +----- Nitia **         
           +--+                    
              |  +-- Coelatura **     
              +--+                 
                 +-- Prisodontopsis **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 81 22 4 --
  +-------------- Lamellidentini
--+                             
  |  +----------- Parreysiini   
  +--+                          
     |  +-------- Leoparreysiini
     +--+                       
        |  +----- Indochinellini
        +--+                    
           |  +-- Coelatura **     
           +--+                 
              +-- Nitia **         

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 55 8 3 --
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Prisodontopsis **

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 44 3 3 --
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Nitia **         

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 116 39 4 --
  +----------- Lamellidentini
  |                          
--+        +-- Parreysiini   
  |  +-----+                 
  |  |     +-- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Nitia **         

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
2 2 0 2 55 8 3 --
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Prisodontopsis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 110 38 4 --
  +----------- Lamellidentini
  |                          
--+        +-- Parreysiini   
  |  +-----+                 
  |  |     +-- Leoparreysiini
  +--+                       
     |  +----- Indochinellini
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Nitia **         

Bolotov et al., 2019 [Publ 3974], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 16 11 2 --
  +-------- Leoparreysiini
--+                       
  |  +----- Indochinellini
  +--+                    
     |  +-- Nitia **         
     +--+                 
        +-- Coelatura **     

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 12 3 --
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 10 3 --
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
  +-- Lamellidentini
--+                 
  +-- Coelatura **     

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 0 0 --
-- Coelatura_aegyptiaca

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 3 3 --
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 22 4 --
  +----------- Lamellidentini
--+                          
  |  +-------- Leoparreysiini
  +--+                       
     |  +----- Parreysiini   
     +--+                    
        |  +-- Coelatura **     
        +--+                 
           +-- Indochinellini

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 0 0 --
-- Coelatura_aegyptiaca

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 --
  +-- Lamellidentini
--+                 
  +-- Coelatura **     

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 2 2 --
  +-- Coelatura **     
  |                 
--+-- Leoparreysiini
  |                 
  +-- Indochinellini

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 0 0 --
-- Coelatura_aegyptiaca

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 12 3 --
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +-- Coelatura **     
  |                 
--+-- Indochinellini
  |                 
  +-- Parreysiini   

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 7 3 --
  +-------- Lamellidentini
--+                       
  |  +----- Leoparreysiini
  +--+                    
     |  +-- Coelatura **     
     +--+                 
        +-- Indochinellini

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +----- Parreysiini   
--+                    
  |  +-- Coelatura **     
  +--+                 
     +-- Indochinellini

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
-- Grandidieria

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Coelatura_aegyptiaca

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Coelatura_aegyptiaca

Page updated: 08:34:08 Wed 06 Nov 2024

 
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