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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Lopes-Lima et al. (2020), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Lopes-Lima, M., A. Hattori, T. Kondo, J.H. Lee, S.K. Kim et al. 2020. Freshwater mussels (Bivalvia: Unionidae) from the rising sun (Far East Asia): phylogeny, systematics, and distribution. Molecular Phylogenetics and Evolution 146: 1-27.

Click here for online access.

Cladogram Figures

(click the figure number to see the cladogram)

fig. 10
fig. 11
fig. 12
fig. 13
fig. 14
fig. 16
fig. 17
fig. 2-4
fig. 5
fig. 6
fig. 7
fig. 8
fig. 9

Attributed Genus Names

Koreosolenaia Lopes-Lima et al., 2020
Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020, MPE 146: [15, 23].
Type species: Koreosolenaia sitgyensis Lopes-Lima et al., 2020 (orig. desig.) = Koreosolenaia sitgyensis

Apppius Lopes-Lima et al., 2020
‘Menke’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Appius Menke, 1830)

Nipponihyra Lopes-Lima et al., 2020
‘Starobogatov’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Nipponihyria Starobogatov, 1970)

Attributed Species Names

Anodon randalii Conrad, 1857
‘Trask’ Conrad, 1857, Proc. Acad. Nat. Sci. 9: 165.
‘Trask’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Anodonta randalli Trask, 1854)

Lasmigona compressa Graf & Ó Foighil, 2000
‘Lea’ of Authors.
Graf & Ó Foighil, 2000, J. Moll. Studies 66: 160.
Graf & Cummings, 2006, Zool. J. Linn. Soc. 148: 348.
Hoeh et al., 2009, Malacologia 51(2): 310.
Jadhav & Jamkhedkar, 2009, Res. J. BioTech. 4(1): 43.
Whelan et al., 2011, MPE 61: 506.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Huang et al., 2019, MPE 130: Table S4.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Wu et al., 2023, ZooKeys 1166: 263, Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
(misidentified reference to Symphynota compressa Lea, 1829)

Middendorffinaia suifunensis Graf, 2007
‘Moskvicheva & Starobogatov’ of Authors.
Graf, 2007, Proc. Acad. Nat. Sci. 156: 85.
Lopes-Lima et al., 2020, MPE 146: Appendix 1.
Bolotov et al., 2020, Sci. Repts. 10: S32.
Bogatov, 2022, Lg. Bivalve Moll. Russia’s Fresh Waters: 59, 61.
(error for Middendorffinaia (Suifununio) sujfunensis Moskvicheva & Starobogatov, 1973)

Lamellidens corrianus Whelan et al., 2011
‘Lea’ of Authors.
Whelan et al., 2011, MPE 61: 506.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens corrianus sp. 1].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Pfeiffer et al., 2018, J. Moll. Studies 84(4): [3].
Pfeiffer et al., 2019, MPE 137: 117.
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Unio corrianus Lea, 1834)
Type(s): SEQUENCED SPECIMEN UMMZ_304642, Ayeryarwaddy River, mud bank on E side of river, Mandalay Division, Burma. 21.48187, 95.30501.

Pseudodon inoscularis Pfeiffer & Graf, 2013
‘Gould’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. inoscularis’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Huang et al., 2019, MPE 130: Table S4 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘ “Monodontina” cf. inoscularis sp. 2’].
(misidentified reference to Anodon inoscularis Gould, 1844)

Pilsbryoconcha exilis Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016, PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.

Ensidens sagittarius Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Ensidens aff. sagittarius sp. 1].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘E. aff. sagittarius sp. 1].
Huang et al., 2019, MPE 130: Table S4 [as ‘Ensidens aff. sagittarius sp. 1].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2242.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
(misidentified reference to Unio sagittarius Lea, 1856)

Pseudodon cumingii Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘”Bineurus” cumingii’].
Huang et al., 2019, MPE 130: Table S4 [as ‘”Bineurus” cumingii’].
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Anodonta cumingii Lea, 1851)
Type(s): SPECIMEN UMMZ_304648, Nam Phiat, Phonethong, ca. 2 km from confluence, Bolikhamxai Province, Laos. 18.08389, 104.97806.

Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
(misidentified reference to Unio hainesianus Lea, 1856)

Sinanodonta woodiana Ng et al., 2016
‘Lea’ of Authors.
Ng et al., 2016, PLoS One 11(8), e0161130: Table S2.
Bolotov et al., 2016, Biochem. Syst. & Ecol. 67: 60 [as ‘lineage A’ & ‘lineage B’]. [in part]
Zieritz et al., 2016, Sci. Total Environ. 571: 1071.
Vikhrev et al., 2017, Trop. Conserv. Sci. 10: Table 1 [as ‘lineage A’ & ‘lineage B’]. [in part]
Bespalaya et al., 2018, Limnologica 69: Table 1 [as ‘lineage A’ & ‘lineage B’]. [in part]
Konečný et al., 2018, Evol. Appl. 11: Table S1 [as ‘lineage A’ & ‘lineage B’]. [in part]
Zieritz et al., 2018, Biol. Conserv. 219: 131.
Kondakov et al., 2018, Zootaxa 4462(4): 517, Table 1.
Lopes-Lima et al., 2020, MPE 146: Table S7 [as ‘Sinanodonta cf. woodiana 1’]. [in part]
Bauer et al., 2021, Neotrop. Natur. 5: 1, figs. 1-2.
Sano et al., 2022, MPE 175: Table S1 [as ‘S. woodiana sp. 1’]. [in part]
Bolotov et al., 2022, Aquatic Conserv.: Mar. Freshw. Ecosyst. 32(9): 1498, Table S3. [in part]
Bogan et al., 2023, Diversity 15(6), 710: Tables 1, S3 [as ‘Sinanodonta cf. woodiana 1’]. [in part]
(misidentified reference to Symphynota woodiana Lea, 1834)

Hyriopsis bialata Lopes-Lima et al., 2017
‘Simpson’ of Authors.
Lopes-Lima et al., 2017, MPE 106: 177.
Muanta et al., 2019, J. Moll. Studies 85(2): 225.
Lopes-Lima et al., 2020, MPE 146: Table 1.
(misidentified reference to Hyriopsis bialatus Simpson, 1900)

Pseudodon vondembuschianus Bolotov et al., 2017
‘Lea’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon aff. vondembuschianus sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Monodontina aff. vondembuschiana sp. 1’].
(misidentified reference to Margaritana vondembuschiana Lea, 1840)

Pseudodon avae Bolotov et al., 2017
‘Theobald’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. avae’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1 [as ‘Pseudodon cf. avae’].
Huang et al., 2019, MPE 130: Table S4.
Konopleva et al., 2019, Sci. Repts. 9: Table S1.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Dai et al., 2023, Invert. Syst. 37(2): 155, Table 1.
(misidentified reference to Monocondylaea avae Theobald, 1873)

Solenaia oleivora Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S4 [as ‘Solenaia oleivora sp. 2’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Solenaia cf. oleivora 3’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 2’].
(misidentified reference to Mycetopus oleivorus Heude, 1877)

Solenaia oleivora Huang et al., 2019
‘Heude’ of Authors.
Huang et al., 2019, MPE 130: Table S4 [as ‘Solenaia oleivora SDCFM189’].
Chen et al., 2020, mtDNA B 5(3): 2800 [as ‘Solenaia oleivora MG’]. [in part]
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘Solenaia cf. oleivora 1’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Solenaia oleivora sp. 1’].
Sano et al., 2020, Plankton Benthos Res. 15(2): 101.
Bolotov et al., 2022, Hindawi 6600359: Table S1.
(misidentified reference to Mycetopus oleivorus Heude, 1877)

Koreosolenaia sitgyensis Lopes-Lima et al., 2020
Lee, Kim, Lopes-Lima & Bogan in Lopes-Lima et al., 2020, MPE 146: [23], fig. 18D.

Unio osbreckii Lopes-Lima et al., 2020
‘Philippi’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Unio osbeckii Philippi, 1844)

Pronodularia japanensis Lopes-Lima et al., 2020
‘Lea’ of Authors.
Lopes-Lima et al., 2020, MPE 146: fig. 16 [as ‘Pronodularia cf. japanensis 3]. [in part]
(reference to Unio japanensis Lea, 1859)

Beringiana fukuharai Lopes-Lima et al., 2020
San, Hattori & Kondo in Lopes-Lima et al., 2020, MPE 146: [20], fig. 18A.

Beringiana gosannensis Lopes-Lima et al., 2020
San, Hattori & Kondo in Lopes-Lima et al., 2020, MPE 146: [20], fig. 18B.

Buldowskia kamiyai Lopes-Lima et al., 2020
San, Hattori & Kondo in Lopes-Lima et al., 2020, MPE 146: [4, 22], fig. 18C.

Nodularia breviconcha Lopes-Lima et al., 2020
Lee, Kim, Bogan & Kondo in Lopes-Lima et al., 2020, MPE 146: [5, 23].
(new name for Unio douglasiae var. sinuolatus Martens, 1905)

Taxonomic Opinions

Aculamprotula cf. fibrosa (Heude, 1877)

Aculamprotula coreana (Martens, 1886)

Aculamprotula scripta (Heude, 1875)

Aculamprotula tientsinensis

Aculamprotula tortuosa (Lea, 1865)

Acuticosta chinensis

Alasmidonta marginata

Amblema plicata

Amuranodonta kijaensis Moskvicheva, 1973
+ Amuranodonta parva Moskvicheva, 1973
+ Amuranodonta minima Moskvicheva, 1973
+ Buldowskia (Amuranodonta) lomakini Zatravkin & Bogatov, 1987
+ Buldowskia (Amuranodonta) boloniensis Zatravkin & Bogatov, 1987
+ Amuranodonta inflata Bogatov & Starobogatov, 1996
+ Amuranodonta pulchra Bogatov & Starobogatov, 1996
+ Anemina ussuriensis Qian et al., 2015

Anemina arcaeformis (Heude, 1877)
+ Anodon rivularis Heude, 1877
+ Anodonta heudei Neumayr, 1899
+ Anemina globosula Wu et al., 2018
+ Anemina euscaphys Huang et al., 2019

Anemina euscaphys

Anodonta anatina

Anodonta cygnea

Anodonta nuttalliana

Aspatharia pfeifferiana

Beringiana beringiana (Middendorff, 1851)
+ Anodonta youkanensis Lea, 1867
+ Anodonta beringiana var. taranetzi Zhadin, 1938
+ Amuranodonta sihotealinica Zatravkin & Starobogatov, 1984
+ Arsenievinaia coptzevi Zatravkin & Bogatov, 1987
+ Arsenievinaia zimini Zatravkin & Bogatov, 1987
+ Arsenievinaia zarjaensis Bogatov & Zatravkin, 1988
+ Arsenievinaia alimovi Bogatov & Zatravkin, 1988
+ Arsenievinaia compressa Bogatov & Starobogatov, 1996
+ Arsenievinaia zatravkini Bogatov & Starobogatov, 1996
+ Beringiana compressa Sayenko & Bogatov, 1998
+ Kunashiria japonica boreosakhalinensis Labay & Shulga, 1999
+ Anemina (Buldowskia) lacustris Labay & Shulga, 1999
+ Anemina (Buldowskia) adotymensys Labay & Shulga, 1999
+ Kunashiria iturupica Bogatov et al., 1999
+ Kunashiria sinanodontoides Bogatov et al., 1999
+ Beringiana kamchatica Bogatov & Starobogatov, 2001
+ Beringiana derzhavini Bogatov & Starobogatov, 2001
+ Beringiana chereshnevi Bogatov & Starobogatov, 2001

Beringiana fukuharai San, Hattori & Kondo sp. nov.

Beringiana gosannensis San, Hattori & Kondo sp. nov.

Beringiana japonica (Clessin, 1874)

Bineurus aff. mouhotii sp. 1

Bineurus cumingii

Buldowskia flavotincta (Martens, 1905)

Buldowskia iwakawai (Suzuki, 1939)

Buldowskia kamiyai San, Hattori & Kondo sp. nov.

Buldowskia shadini (Moskvicheva, 1973)
+ Anodonta (Haasiella) euscaphus Zhadin, 1938
+ Sinanodonta (Anemina) fuscoviridis Moskvicheva, 1973
+ Sinanodonta (Anemina) buldowskii Moskvicheva, 1973
+ Buldowskia (Amurbuldowskia) sitaensis Bogatov & Starobogatov, 1996

Buldowskia suifunica (Lindholm, 1925)
+ Anodonta beringiana var. suifunensis Zhadin, 1938
+ Anodonta woodiana var. elliptica Zhadin, 1938
+ Anodonta (Haasiella) arcaeformis Zhadin, 1938
+ Buldowskia suputinensis Moskvicheva, 1973
+ Amuranodonta starobogatovi Moskvicheva, 1973
+ Buldowskia cylindrica Moskvicheva, 1973
+ Anemina (Buldowskia) zatrawkini Martynov & Chernyshev, 1992
+ Anemina (s.s.) shadini deflexa Martynov & Chernyshev, 1992
+ Buldowskia (Buldowskia) possietica Bogatov & Starobogatov, 1996
+ Buldowskia (Buldowskia) koreana Bogatov & Starobogatov, 1996

Chambardia wahlbergi

Chamberlainia hainesiana

Coelatura aegyptiaca

Contradens contradens

Cristaria cf. radiata

Cristaria clessini (Kobelt, 1879)
+ Dipsas plicata var. japonica Kobelt, 1879

Cristaria plicata (Leach, 1814)
+ Cristaria tuberculata Schumacher, 1817
+ Anodonta cristata Blainville, 1825
+ Anodonta dipsas Blainville, 1825
+ Symphynota bialata Lea, 1829
+ Dianisotis chinensis Rafinesque, 1831
+ Anodonta (Dispsas) herculea Middendorff, 1848
+ Anodon gigantea Gould, 1849
+ Margaron (Anodonta) onowensis Lea, 1852
+ Anodonta bellua Morelet, 1866
+ Anodon herculeus Sowerby, 1867
+ Anodonta magnifica Clessin, 1873
+ Anodonta (Dipsas) spatiosa Clessin, 1876
+ Dipsas occidentalis Heude, 1885
+ Barbala plicatula Paetel, 1890

Cuneopsis celtiformis

Cuneopsis cf. heudei

Cuneopsis pisciculus

Cuneopsis rufescens

Elliptio complanata

Ensidens aff. sagittarius sp. 1

Etheria elliptica

Gonidea angulata (Lea, 1838)

Hyridella depressa

Hyriopsis bialata

Inversidens brandtii (Kobelt, 1879)

Inversidens pantoensis (Neumayr, 1899)

Inversiunio jokohamensis (Ihering, 1893)
+ Unio japanensis Kobelt, 1879
+ Unio haconensis Ihering, 1893

Inversiunio reinianus (Kobelt, 1879)
+ Nodularia hirasei Haas, 1911

Inversiunio verrucosus Kondo, Yang & Choi, 2007

Inversiunio yanagawensis (Kondo, 1982)

Koreosolenaia sitgyensis Lee, Kim, Lopes-Lima & Bogan gen. & sp. nov.
+ Solenaia triangularis Wu et al., 2018

Lamellidens corrianus

Lamproscapha ensiformis

Lamprotula caveata (Heude, 1877)
+ Lamprotula scripta Wang et al., 2018

Lamprotula leaii (Gray in Griffith & Pidgeon, 1833)
+ Unio nodulosus Reeve, 1864
+ Unio leleci Heude, 1875
+ Unio richthofeni Martens, 1875
+ Unio gottschei Martens, 1894
+ Unio leai var. truncatula Neumayr, 1899
+ Unio (Quadrula) leai var. ponderosa Dautzenberg & Fischer, 1905

Lamprotula seomjinensis (Kondo, Yang & Choi, 2007)

Lampsilis cardium

Lanceolaria acrorrhyncha (Martens, 1894)

Lanceolaria cf. bilirata (Martens, 1902)

Lanceolaria gladiola

Lanceolaria grayii (Griffith & Pidgeon, 1833)
+ Unio grayanus Schrenck, 1867
+ Unio grayanus var. decurvata Schrenck, 1867
+ Nodularia cylindracea Simpson, 1900
+ Nodularia cylindrica Simpson, 1900
+ Lanceolaria ussuriensis Moskvicheva, 1973
+ Lanceolaria chankensis Moskvicheva, 1973
+ Lanceolaria maacki Moskvicheva, 1973
+ Lanceolaria (Pericylindrica) bogatovi Zatravkin & Starobogatov, 1984

Lanceolaria kihirai (Kondo & Hattori, 2019)

Lanceolaria lanceolata (Lea, 1856)

Lanceolaria oxyrhyncha (Martens, 1861)

Lanceolaria triformis (Heude, 1877)

Lasmigona compressa

Leguminaia wheatleyi

Leoparreysia bhamoensis

Lepidodesma languilati

Margaritifera dahurica

Margaritifera laevis

Margaritifera middendorffi

Microcondylaea bonellii

Middendorffinaia mongolica (Middendorff, 1851)
+ Unio (Nodularia) continentalis Zhadin, 1938
+ Middendorffinaia (Suifununio) sujfunensis Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) shadini Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) dulkeitiana Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) weliczkowskii Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) hassanica Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (Pseudopotomida) martensi Moskvicheva & Starobogatov, 1973
+ Middendorffinaia maihensis Moskvicheva, 1973
+ Middendorffinaia alimovi Bogatov, 2012

Monodontina aff. vondembuschiana sp. 1

Monodontina aff. “inoscularis” sp. 2

Mycetopoda siliquosa

Neotrigonia lamarckii

Neotrigonia margaritacea

Nodularia breviconcha Lee, Kim, Bogan & Kondo nom. nov.
+ Unio douglasiae var. sinuolatus Martens, 1905

Nodularia douglasiae (Griffith & Pidgeon, 1833)
+ Unio murchisonianus Lea, 1834
+ Unio osbeckii Philippi, 1844
+ Unio pictorum var. longirostris Middendorff, 1851
+ Unio shanghaiensis Lea, 1859
+ Unio wrightii Lea, 1865
+ Unio sculptus Deshayes, 1873
+ Unio biwae Kobelt, 1879
+ Unio douglasiae chloreus Heude, 1885
+ Unio douglasiae dactylinus Heude, 1885
+ Unio douglasiae deltae Heude, 1885
+ Unio douglasiae firmus Heude, 1885
+ Unio douglasiae hainanensis Heude, 1885
+ Unio pictorum var. amurensis Mousson, 1887
+ Unio middendorffi Westerlund, 1890
+ Unio verrucifer Martens, 1894
+ Unio pliculosus Martens, 1894
+ Unio abbreviatus Westerlund, 1897
+ Unio schrencki Westerlund, 1897
+ Nodularia douglasiae flavoviridis Haas, 1910
+ Nodularia sujfunica Moskvicheva, 1973
+ Nodularia vladivostokensis Moskvicheva, 1973
+ Middendorffinaia (s.s.) ussuriensis Moskvicheva & Starobogatov, 1973
+ Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973
+ Nodularia lebedevi Zatravkin & Starobogatov, 1984
+ Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992
+ Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992
+ Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000
+ Nodularia sakhalinensis Bogatov, 2001
+ Nodularia jourdyi Bolotov et al., 2018

Nodularia nipponensis (Martens, 1877)

Nodularia nuxpersicae

Obovalis omiensis (Heimburg, 1884)

Parvasolenaia neotriangularis He & Zhuang, 2013

Parvasolenaia rivularis (Heude, 1877)
+ Solenaia triangularis Lopes-Lima et al., 2017

Parvasolenaia triangularis (Heude, 1885)

Physunio modelli

Pilsbryoconcha aff. compressa sp. 1

Pilsbryoconcha aff. exilis sp. 1

Pletholophus reinianus (Martens, 1875)

Pletholophus tenuis (Gray in Griffith & Pidgeon, 1833)
+ Symphynota discoidea Lea, 1834

Potomida littoralis

Pronodularia cf. japanensis (Lea, 1859)

Pseudanodonta complanata

Pseudodon avae

Pseudodon manueli

Ptychorhynchus pfisteri (Heude, 1874)

Pyganodon grandis

Quadrula quadrula

Radiatula mouhoti

Rectidens sumatrensis

Schistodesmus lampreyanus

Schistodesmus spinosus

Simpsonaias ambigua

Sinanodonta angula (Tchang, Li & Liu, 1965)

Sinanodonta calipygos (Kobelt, 1879)

Sinanodonta cf. elliptica
+ Sinanodonta woodiana Bolotov et al., 2016
+ Sinanodonta joreti Huang et al., 2019
+ Sinanodonta pacifica Huang et al., 2019

Sinanodonta cf. woodiana 2 (Lea, 1834)
+ Sinanodonta elliptica Huang et al., 2019

Sinanodonta jourdyi
+ Sinanodonta woodiana Lopes-Lima et al., 2017

Sinanodonta lauta (Martens, 1877)
+ Sinanodonta woodiana fukudai Modell, 1945
+ Sinanodonta (Ellipsanodon) ovata Bogatov & Starobogatov, 1996
+ Sinanodonta (Ellipsanodon) manchurica Bogatov & Starobogatov, 1996

Sinanodonta lucida

Sinanodonta schrenkii (Lea, 1870)
+ Anodonta magnifica Schrenck, 1867
+ Sinanodonta (s.s.) schrencki Moskvicheva, 1973
+ Sinanodonta (s.s.) amurensis Moskvicheva, 1973
+ Sinanodonta (Cristariopsis) crassitesta Moskvicheva, 1973
+ Sinanodonta (s.s.) likharevi Moskvicheva, 1973
+ Sinanodonta (s.s.) renzini Bogatov & Zatravkin, 1988
+ Sinanodonta (Cristariopsis) primorjensis Bogatov & Zatravkin, 1988
+ Sinanodonta woodiana Soroka, 2010

Sinanodonta tumens (Haas, 1910)
+ Oguranodonta ogurae Kuroda & Habe, 1987

Sinohyriopsis cf. cumingii (Lea, 1852)

Sinohyriopsis goliath (Rolle, 1904)

Sinohyriopsis schlegelii (Martens, 1861)

Solenaia carinata

Solenaia cf. oleivora 2

Solenaia sp.

Strophitus undulatus

Trapezoideus nesemanni

Unio crassus

Unio gibbus

Unio pictorum

Unio tumidus

Velesunio ambiguus

Cladogram Figure 10
                    +-- Margaritifera_middendorffi
  +-----------------+                             
  |                 +-- Margaritifera_laevis      
  |                                               
--+                 +-- Beringiana_japonica       
  |  +--------------+                             
  |  |              +-- Beringiana_beringiana     
  |  |                                            
  +--+              +-- Sinanodonta_angula        
     |  +-----------+                             
     |  |           +-- Sinanodonta_lucida        
     |  |                                         
     +--+  +----------- Sinanodonta_schrenkii     
        |  |                                      
        |  |        +-- Sinanodonta_calipygos     
        |  +--------+                             
        +--+        +-- Sinanodonta_tumens        
           |                                      
           |  +-------- Sinanodonta_woodiana      
           |  |                                   
           +--+-------- Sinanodonta_jourdyi       
              |                                   
              |  +----- Sinanodonta_elliptica     
              +--+                                
                 |  +-- Sinanodonta_lauta         
                 +--+                             
                    +-- Sinanodonta               
Cladogram Figure 11
                       +-- Margaritifera_laevis      
  +--------------------+                             
  |                    +-- Margaritifera_middendorffi
  |                                                  
--+                    +-- Nodularia_breviconcha     
  |  +-----------------+                             
  |  |                 +-- Nodularia_douglasiae      
  +--+                                               
     |  +----------------- Lanceolaria_oxyrhyncha    
     |  |                                            
     +--+  +-------------- Lanceolaria_gladiola      
        |  |                                         
        +--+           +-- Lanceolaria_grayii        
           |  +--------+                             
           |  |        +-- Lanceolaria_bilirata      
           +--+                                      
              |  +-------- Lanceolaria_acrorrhyncha  
              +--+                                   
                 |  +----- Lanceolaria_kihirai       
                 +--+                                
                    |  +-- Lanceolaria_lanceolata    
                    +--+                             
                       +-- Lanceolaria_triformis     
Cladogram Figure 12
              +-- Margaritifera_laevis      
  +-----------+                             
  |           +-- Margaritifera_middendorffi
  |                                         
--+           +-- Nodularia_breviconcha     
  |  +--------+                             
  |  |        +-- Nodularia_douglasiae      
  +--+                                      
     |  +-------- Inversiunio_jokohamensis  
     |  |                                   
     +--+  +----- Inversiunio_verrucosus__1 
        |  |                                
        +--+----- Inversiunio_verrucosus__2 
           |                                
           |  +-- Inversiunio_reinianus     
           +--+                             
              +-- Inversiunio_yanagawensis  
Cladogram Figure 13
                       +-- Margaritifera_laevis      
  +--------------------+                             
  |                    +-- Margaritifera_middendorffi
--+                                                  
  |  +-------------------- Middendorffinaia_mongolica
  +--+                                               
     |  +----------------- Nodularia_breviconcha     
     +--+                                            
        |  +-------------- Nodularia_fusiformans     
        +--+                                         
           |  +----------- Nodularia_huana           
           +--+                                      
              |  +-------- Nodularia_nuxpersicae     
              +--+                                   
                 |  +----- Nodularia_nipponensis     
                 +--+                                
                    |  +-- Nodularia_douglasiae      
                    +--+                             
                       +-- Nodularia_dualobtusa      
Cladogram Figure 14
        +-- Margaritifera_laevis      
  +-----+                             
  |     +-- Margaritifera_middendorffi
  |                                   
--+     +-- Parvasolenaia_rivularis   
  |  +--+                             
  |  |  +-- Koreosolenaia_sitgyensis  
  +--+                                
     |  +-- Lamprotula_leaii          
     +--+                             
        +-- Lamprotula_seomjinensis   
Cladogram Figure 16
                    +-- Margaritifera_laevis      
  +-----------------+                             
  |                 +-- Margaritifera_middendorffi
--+                                               
  |  +----------------- Chamberlainia_somsakpanhai
  +--+                                            
     |  +-------------- Sinohyriopsis_schlegelii  
     +--+                                         
        |  +----------- Pronodularia_japanensis   
        +--+                                      
           |  +-------- Lamprotula_caveata        
           +--+                                   
              |  +----- Lamprotula_leaii__2       
              +--+                                
                 |  +-- Lamprotula_seomjinensis   
                 +--+                             
                    +-- Lamprotula_leaii__1       
Cladogram Figure 17
              +-- Margaritifera_laevis      
  +-----------+                             
  |           +-- Margaritifera_middendorffi
  |                                         
--+  +----------- Chamberlainia_somsakpanhai
  |  |                                      
  |  |  +-------- Sinohyriopsis_cumingii__10
  +--+  |                                   
     |  |  +----- Sinohyriopsis_cumingii__1 
     |  |  |                                
     |  |  +----- Sinohyriopsis_cumingii__2 
     +--+  |                                
        |  +----- Sinohyriopsis_cumingii__3 
        |  |                                
        |  +----- Sinohyriopsis_cumingii__4 
        |  |                                
        |  +----- Sinohyriopsis_cumingii__5 
        +--+                                
           +----- Sinohyriopsis_cumingii__6 
           |                                
           +----- Sinohyriopsis_cumingii__7 
           |                                
           +----- Sinohyriopsis_cumingii__8 
           |                                
           +----- Sinohyriopsis_cumingii__9 
           |                                
           |  +-- Sinohyriopsis_schlegelii  
           +--+                             
              +-- Sinohyriopsis_goliath     
Cladogram Figure 2-4
                                                  +-- Neotrigonia_lamarckii      
  +-----------------------------------------------+                              
  |                                               +-- Neotrigonia_margaritacea   
  |                                                                              
  |                                               +-- Hyridella_depressa         
  |  +--------------------------------------------+                              
  |  |                                            +-- Velesunio_ambiguus         
--+  |                                                                           
  |  |                                            +-- Margaritifera_laevis       
  |  |  +-----------------------------------------+                              
  |  |  |                                         +-- Margaritifera_middendorffi 
  |  |  |                                                                        
  |  |  |                                         +-- Lamproscapha_ensiformis    
  +--+  |                                   +-----+                              
     |  |                                   |     +-- Mycetopoda_siliquosa       
     |  |  +--------------------------------+                                    
     |  |  |                                |  +----- Etheria_elliptica          
     |  |  |                                +--+                                 
     |  |  |                                   |  +-- Aspatharia_pfeifferiana    
     |  |  |                                   +--+                              
     |  |  |                                      +-- Chambardia_wahlbergi       
     |  |  |                                                                     
     +--+  |                                      +-- Coelatura_aegyptiaca       
        |  |                                      |                              
        |  |  +-----------------------------------+-- Leoparreysia_tavoyensis    
        |  |  |                                   |                              
        |  |  |                                   +-- Radiatula_mouhoti          
        |  |  |                                                                  
        |  |  |                                   +-- Lens_contradens            
        |  |  |                                   |                              
        |  |  |                                   +-- Ensidens_sagittarius       
        |  |  |                                   |                              
        |  |  |           +-----------------------+-- Physunio_modelli           
        |  |  |           |                       |                              
        |  |  |           |                       +-- Rectidens_sumatrensis      
        |  |  |           |                       |                              
        +--+  |           |                       +-- Yaukthwa_nesemanni         
           |  |           |                                                      
           |  |           |                       +-- Amblema_plicata            
           |  |           |                       |                              
           |  |           |                       +-- Elliptio_complanata        
           |  |           |  +--------------------+                              
           |  |  +--------+  |                    +-- Lampsilis_cardium          
           |  |  |        |  |                    |                              
           |  |  |        |  |                    +-- Quadrula_quadrula          
           |  |  |        |  |                                                   
           |  |  |        |  |                 +----- Chamberlainia_somsakpanhai 
           |  |  |        |  |     +-----------+                                 
           |  |  |        |  |     |           |  +-- Sinohyriopsis_cumingii     
           |  |  |        |  |     |           +--+                              
           |  |  |        |  |  +--+              +-- Sinohyriopsis_schlegelii   
           |  |  |        |  |  |  |                                             
           |  |  |        |  |  |  |  +-------------- Pronodularia_japanensis    
           |  |  |        +--+  |  |  |                                          
           |  |  |           |  |  +--+  +----------- Potomida_littoralis        
           |  |  |           |  |     |  |                                       
           |  |  |           |  |     +--+  +-------- Lamprotula_caveata         
           |  |  |           |  |        |  |                                    
           +--+  |           |  |        +--+  +----- Lamprotula_leaii__2        
              |  |           |  |           |  |                                 
              |  |           |  |           +--+----- Lamprotula_leaii__3        
              |  |           |  |              |                                 
              |  |           |  |              |  +-- Lamprotula_seomjinensis    
              |  |           |  |              +--+                              
              |  |           |  |                 +-- Lamprotula_leaii__1        
              |  |           +--+                                                
              |  |              |                 +-- Indopseudodon_indawgyiensis
              |  |              |        +--------+                              
              |  |              |        |        +-- Indopseudodon_salwenianus  
              |  |              |        |                                       
              |  |              |  +-----+        +-- Pilsbryoconcha_exilis      
              |  |              |  |     |  +-----+                              
              |  |              |  |     |  |     +-- Pilsbryoconcha_carinifera  
              |  |              |  |     +--+                                    
              |  |              |  |        |  +----- Pseudodon_mekongi          
              |  |              |  |        |  |                                 
              |  |              |  |        +--+  +-- Nyeinchanconcha_nyeinchani 
              |  |              |  |           |  |                              
              |  |              |  |           +--+-- Bineurus_mouhotii          
              |  |              |  |              |                              
              |  |              +--+              +-- Sundadontina_tumida        
              |  |                 |                                             
              |  |                 |        +-------- Inversidens_brandtii       
              |  |                 |        |                                    
              |  |                 |  +-----+     +-- Obovalis_omiensis          
              |  |                 |  |     |  +--+                              
              |  |                 |  |     |  |  +-- Ptychorhynchus_pfisteri    
              |  |                 |  |     +--+                                 
              |  |                 |  |        |  +-- Parvasolenaia_rivularis    
              |  |                 |  |        +--+                              
              |  |                 +--+           +-- Koreosolenaia_sitgyensis   
              |  |                    |                                          
              +--+                    |        +----- Gonidea_angulata           
                 |                    |  +-----+                                 
                 |                    |  |     |  +-- Leguminaia_wheatleyi       
                 |                    |  |     +--+                              
                 |                    +--+        +-- Microcondylaea_bonellii    
                 |                       |                                       
                 |                       |  +-------- Sinosolenaia_carinata      
                 |                       +--+                                    
                 |                          |  +----- Sinosolenaia_recognita     
                 |                          +--+                                 
                 |                             |  +-- Sinosolenaia_iridinea      
                 |                             +--+                              
                 |                                +-- Sinosolenaia_oleivora      
                 |                                                               
                 |  +-------------------------------- Acuticosta_chinensis       
                 |  |                                                            
                 |  +-------------------------------- Lepidodesma_languilati     
                 |  |                                                            
                 |  |                    +----------- Aculamprotula_tortuosa     
                 |  |     +--------------+                                       
                 |  |     |              |  +-------- Aculamprotula_scripta      
                 |  |     |              +--+                                    
                 |  |     |                 |  +----- Aculamprotula_coreana      
                 |  |     |                 +--+                                 
                 |  |     |                    |  +-- Aculamprotula_fibrosa      
                 |  |     |                    +--+                              
                 |  +-----+                       +-- Aculamprotula_tientsinensis
                 |  |     |                                                      
                 |  |     |  +----------------------- Middendorffinaia_mongolica 
                 |  |     |  |                                                   
                 |  |     |  |                 +----- Unio_gibbus                
                 |  |     |  |                 |                                 
                 |  |     |  |  +--------------+----- Unio_tumidus               
                 |  |     +--+  |              |                                 
                 |  |        |  |              |  +-- Unio_crassus               
                 |  |        |  |              +--+                              
                 |  |        |  |                 +-- Unio_pictorum              
                 |  |        |  |                                                
                 |  |        |  |     +-------------- Nodularia_breviconcha      
                 |  |        +--+  +--+                                          
                 |  |           |  |  |  +----------- Nodularia_huana            
                 |  |           |  |  +--+                                       
                 |  |           |  |     |  +-------- Nodularia_nuxpersicae      
                 |  |           |  |     +--+                                    
                 |  |           |  |        |  +----- Nodularia_nipponensis      
                 |  |           |  |        +--+                                 
                 |  |           |  |           |  +-- Nodularia_douglasiae       
                 |  |           +--+           +--+                              
                 |  |              |              +-- Nodularia_dualobtusa       
                 |  |              |                                             
                 +--+              |              +-- Schistodesmus_lampreyanus  
                    |              |  +-----------+                              
                    |              |  |           +-- Schistodesmus_spinosus     
                    |              |  |                                          
                    |              |  |        +----- Cuneopsis_rufescens        
                    |              |  +--------+                                 
                    |              +--+        |  +-- Cuneopsis_heudei           
                    |                 |        +--+                              
                    |                 |           +-- Cuneopsis_celtiformis      
                    |                 |                                          
                    |                 |  +----------- Tchangsinaia_piscicula     
                    |                 +--+                                       
                    |                    |  +-------- Inversiunio_jokohamensis   
                    |                    +--+                                    
                    |                       |  +----- Inversiunio_verrucosus     
                    |                       +--+                                 
                    |                          |  +-- Inversiunio_reinianus      
                    |                          +--+                              
                    |                             +-- Inversiunio_yanagawensis   
                    |                                                            
                    |              +----------------- Lanceolaria_oxyrhyncha     
                    |              |                                             
                    |  +-----------+  +-------------- Lanceolaria_gladiola       
                    |  |           |  |                                          
                    |  |           +--+           +-- Lanceolaria_bilirata       
                    |  |              |  +--------+                              
                    |  |              |  |        +-- Lanceolaria_grayii         
                    |  |              +--+                                       
                    |  |                 |  +-------- Lanceolaria_acrorrhyncha   
                    |  |                 +--+                                    
                    |  |                    |  +----- Lanceolaria_kihirai        
                    |  |                    +--+                                 
                    |  |                       |  +-- Lanceolaria_lanceolata     
                    |  |                       +--+                              
                    |  |                          +-- Lanceolaria_triformis      
                    |  |                                                         
                    |  |                    +-------- Pseudanodonta_complanata   
                    +--+                    |                                    
                       |                    |  +----- Anodonta_cygnea            
                       |                    +--+                                 
                       |                    |  |  +-- Anodonta_anatina           
                       |  +-----------------+  +--+                              
                       |  |                 |     +-- Anodonta_nuttalliana       
                       |  |                 |                                    
                       |  |                 |  +----- Pyganodon_grandis          
                       |  |                 |  |                                 
                       |  |                 |  +----- Simpsonaias_ambigua        
                       |  |                 +--+                                 
                       |  |                    +----- Strophitus_undulatus       
                       |  |                    |                                 
                       |  |                    |  +-- Alasmidonta_marginata      
                       |  |                    +--+                              
                       |  |                       +-- Lasmigona_costata          
                       +--+                                                      
                          |                       +-- Cristaria_plicata          
                          |  +--------------------+                              
                          |  |                    +-- Cristaria_clessini         
                          |  |                                                   
                          |  |        +-------------- Anemina_arcaeformis        
                          |  |        |                                          
                          |  +--------+  +----------- Amuranodonta_kijaensis     
                          |  |        |  |                                       
                          |  |        +--+  +-------- Buldowskia_suifunica       
                          |  |           |  |                                    
                          |  |           +--+     +-- Buldowskia_flavotincta     
                          +--+              |  +--+                              
                             |              |  |  +-- Buldowskia_shadini         
                             |              +--+                                 
                             |                 |  +-- Buldowskia_iwakawai        
                             |                 +--+                              
                             |                    +-- Buldowskia_kamiyai         
                             |                                                   
                             |  +-------------------- Pletholophus_tenuis        
                             |  |                                                
                             |  |           +-------- Beringiana_japonica        
                             |  |  +--------+                                    
                             +--+  |        |  +----- Beringiana_beringiana      
                                |  |        +--+                                 
                                |  |           |  +-- Beringiana_fukuharai       
                                |  |           +--+                              
                                +--+              +-- Beringiana_gosannensis     
                                   |                                             
                                   |              +-- Sinanodonta_angula         
                                   |  +-----------+                              
                                   |  |           +-- Sinanodonta_lucida         
                                   |  |                                          
                                   +--+  +----------- Sinanodonta_schrenkii      
                                      |  |                                       
                                      |  |        +-- Sinanodonta_calipygos      
                                      |  +--------+                              
                                      +--+        +-- Sinanodonta_tumens         
                                         |                                       
                                         |  +-------- Sinanodonta_woodiana       
                                         |  |                                    
                                         +--+  +----- Sinanodonta_elliptica      
                                            |  |                                 
                                            +--+----- Sinanodonta_jourdyi        
                                               |                                 
                                               |  +-- Sinanodonta_lauta          
                                               +--+                              
                                                  +-- Sinanodonta_pacifica       
Cladogram Figure 5
                 +-- Margaritifera_laevis       
  +--------------+                              
  |              +-- Margaritifera_middendorffi 
  |                                             
--+              +-- Nodularia_breviconcha      
  |  +-----------+                              
  |  |           +-- Nodularia_douglasiae       
  +--+                                          
     |  +----------- Aculamprotula_tortuosa     
     +--+                                       
        |  +-------- Aculamprotula_tientsinensis
        +--+                                    
           |  +----- Aculamprotula_fibrosa      
           +--+                                 
              |  +-- Aculamprotula_scripta      
              +--+                              
                 +-- Aculamprotula_coreana      
Cladogram Figure 6
                    +-- Margaritifera_laevis      
  +-----------------+                             
  |                 +-- Margaritifera_middendorffi
  |                                               
--+                 +-- Sinanodonta_lauta         
  |  +--------------+                             
  |  |              +-- Sinanodonta_woodiana      
  |  |                                            
  +--+  +-------------- Amuranodonta_kijaensis    
     |  |                                         
     |  |           +-- Anemina_arcaeformis       
     +--+  +--------+                             
        |  |        +-- Anemina_euscaphys         
        |  |                                      
        +--+        +-- Buldowskia_iwakawai       
           |  +-----+                             
           |  |     +-- Buldowskia_kamiyai        
           +--+                                   
              |  +----- Buldowskia_shadini        
              +--+                                
                 |  +-- Buldowskia_suifunica      
                 +--+                             
                    +-- Buldowskia_flavotincta    
Cladogram Figure 7
              +-- Margaritifera_laevis  
  +-----------+                         
  |           +-- Margaritifera_dahurica
  |                                     
--+           +-- Sinanodonta_woodiana  
  |  +--------+                         
  |  |        +-- Sinanodonta_tumens    
  +--+                                  
     |  +-------- Beringiana_gosannensis
     +--+                               
        |  +----- Beringiana_fukuharai  
        +--+                            
           |  +-- Beringiana_beringiana 
           +--+                         
              +-- Beringiana_japonica   
Cladogram Figure 8
           +-- Margaritifera_laevis      
  +--------+                             
  |        +-- Margaritifera_middendorffi
  |                                      
--+        +-- Sinanodonta_woodiana      
  |  +-----+                             
  |  |     +-- Sinanodonta_lucida        
  +--+                                   
     |  +----- Cristaria_clessini        
     +--+                                
        |  +-- Cristaria_plicata         
        +--+                             
           +-- Cristaria_radiata         
Cladogram Figure 9
           +-- Margaritifera_laevis      
  +--------+                             
  |        +-- Margaritifera_middendorffi
--+                                      
  |  +-------- Cristaria_plicata         
  +--+                                   
     |  +----- Sinanodonta_schrenkii     
     +--+                                
        |  +-- Pletholophus_tenuis       
        +--+                             
           +-- Pletholophus_reinianus    

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