Bivalvia | Unionoida | Hyrioidea | Hyriidae subfamily VELESUNIONINAE Iredale, 1934: cladomics 6 genera
Cladomics Summary
number of species = 23
number of subtaxa (rank: genus) = 6
analyzed in 62 cladograms in 37 publications
maximum number of terminals in an analysis = 7, mean = 2.31
maximum number of subtaxa (genus) in an analysis = 5, mean = 1.84
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 26
maximum number of included terminals = 7, mean = 4.12
maximum number of subtaxa (genus) = 5, mean = 3.0
proportion of cladograms that support basic monophyly = 0.69
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 25
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 7, mean = 4.2
maximum number of subtaxa (genus) = 5, mean = 3.08
proportion of cladograms that support rigorous monophyly = 0.68
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
19 |
20 |
5 |
14 |
13 |
1 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+----- Diplodon_suavidicus__1
+--------------+
| | +-- Hyriini
| +--+
| +-- Castaliini
|
| +-------- Westralunio **
| |
+-----------+ +-- Hyridella_drapeta
| | +--+
--+ | | +-- Hyridella_depressa
| +--+
| | +-- Diplodon_gratus
| +--+
| +-- Diplodon_suavidicus__2
|
| +-- Hyridella_australis
| +--------------+
| | +-- Cucumerunio
+--+
| +-------------- Echyridella
+--+
| +----------- Velesunio_ambiguus **
+--+
| +-------- Microdontia **
+--+
| +----- Lortiella **
+--+
| +-- Alathyria **
+--+
+-- Velesunio_angasi **
Graf et al., 2015 [Publ 3619], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
5 |
13 |
7 |
1 |
24.5 |
+-------------- Hyriinae
|
--+ +----------- Westralunio **
| |
| | +----- Alathyria_jacksoni **
+--+ +--+
| | | +-- Velesunio **
| | +--+
+--+ +-- Alathyria_profuga **
|
| +----- Alathyria_pertexta **
+--+
| +-- Microdontia **
+--+
+-- Lortiella **
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
14 |
9 |
5 |
191 |
7 |
1 |
23.5 |
+----------- Echyridella
|
+----------- Westralunio **
|
+----------- Hyridellini
|
| +----- Rhipidodontini
+-----+
--+ | +-- Castaliini
| +--+
| +-- Hyriini
|
| +----- Velesunio **
| +--+
| | | +-- Alathyria_jacksoni **
| | +--+
+--+ +-- Alathyria_profuga **
|
| +----- Alathyria_pertexta **
+--+
| +-- Lortiella **
+--+
+-- Microdontia **
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
14 |
12 |
5 |
191 |
7 |
1 |
26.5 |
+-- Westralunio **
+--------+
| +-- Hyriinae
|
--+ +----- Alathyria_jacksoni **
| +--+
| | | +-- Alathyria_profuga **
| | +--+
+--+ +-- Velesunio **
|
| +----- Alathyria_pertexta **
+--+
| +-- Lortiella **
+--+
+-- Microdontia **
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
14 |
12 |
5 |
189 |
7 |
1 |
26.5 |
+-- Westralunio **
+--------+
| +-- Hyriinae
|
--+ +----- Velesunio **
| +--+
| | | +-- Alathyria_jacksoni **
| | +--+
+--+ +-- Alathyria_profuga **
|
| +----- Alathyria_pertexta **
+--+
| +-- Lortiella **
+--+
+-- Microdontia **
Santos-Neto et al., 2016 [Publ 3739], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
13 |
11 |
5 |
14 |
13 |
1 |
16.0 |
+----- Diplodon_suavidicus__2
+-----------------+
| | +-- Hyriini
| +--+
| +-- Castaliini
|
| +-------- Westralunio **
--+ |
| +-----------+ +-- Hyridella_drapeta
| | | +--+
| | | | +-- Hyridella_depressa
| | +--+
| | | +-- Diplodon_suavidicus__1
+--+ +--+
| +-- Diplodon_gratus
|
| +----------------- Echyridella
| |
+--+ +-- Cucumerunio
| +-----------+
| | +-- Hyridella_australis
+--+
| +----------- Velesunio_ambiguus **
+--+
| +-------- Alathyria **
+--+
| +----- Velesunio_angasi **
+--+
| +-- Lortiella **
+--+
+-- Microdontia **
Bolotov et al., 2016 [Publ 3743], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
4 |
24 |
3 |
1 |
25.5 |
+----------- Hyriinae
|
| +----- Microdontia **
--+ +--+
| | | +-- Alathyria_pertexta **
| | +--+
+--+ +-- Lortiella **
|
| +----- Velesunio **
+--+
| +-- Alathyria_jacksoni **
+--+
+-- Alathyria_profuga **
Bolotov et al., 2017 [Publ 3761], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
4 |
133 |
3 |
1 |
25.5 |
+----------- Hyriinae
|
| +----- Velesunio **
--+ +--+
| | | +-- Alathyria_jacksoni **
| | +--+
+--+ +-- Alathyria_profuga **
|
| +----- Microdontia **
+--+
| +-- Alathyria_pertexta **
+--+
+-- Lortiella **
Bolotov et al., 2016 [Publ 3743], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
4 |
24 |
3 |
1 |
25.5 |
+----------- Hyriinae
|
| +----- Microdontia **
--+ +--+
| | | +-- Lortiella **
| | +--+
+--+ +-- Alathyria_pertexta **
|
| +----- Velesunio **
+--+
| +-- Alathyria_jacksoni **
+--+
+-- Alathyria_profuga **
Bolotov et al., 2017 [Publ 3761], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
4 |
133 |
3 |
1 |
25.5 |
+----------- Hyriinae
|
| +----- Alathyria_pertexta **
--+ +--+
| | | +-- Microdontia **
| | +--+
+--+ +-- Lortiella **
|
| +----- Velesunio **
+--+
| +-- Alathyria_jacksoni **
+--+
+-- Alathyria_profuga **
Walker et al., 2006 [Publ 2666], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
4 |
3 |
31 |
5 |
1 |
22.5 |
+----------------- Castaliini
|
+----------------- Rhipidodontini
--+
| +-------------- Hyridella_depressa
| |
| +-------------- Hyridella_australis
+--+
| +----------- Echyridella
| |
| | +-- Velesunio_ambiguus **
+--+ +-----+
| | +-- Velesunio__3 **
| |
+--+ +----- Velesunio__2 **
| |
| | +-- Alathyria **
+--+--+
| +-- Velesunio__1 **
|
| +-- Velesunio_angasi **
+--+
+-- Lortiella **
Klunzinger et al., 2021 [Publ 4086], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
4 |
2 |
2 |
3 |
1 |
1 |
21.0 |
+----- Westralunio **
--+
| +-- Velesunio **
+--+
+-- Hyriinae
Baker et al., 2004 [Publ 2685], fig. 2-3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
4 |
2 |
6 |
5 |
1 |
22.0 |
+----- Hyridella_depressa
+-----------+
| | +-- Hyridella_australis
| +--+
| +-- Rhipidodontini
--+
| +-- Echyridella
| +-----------+
| | +-- Castaliini
| |
+--+ +-- Velesunio__3 **
| +--------+
| | +-- Velesunio__4 **
+--+
| +-------- Velesunio__2 **
+--+
| +----- Velesunio_angasi **
+--+
| +-- Velesunio__1 **
+--+
+-- Alathyria **
Graf & Cummings, 2006 [Publ 1968], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
44 |
6 |
1 |
20.5 |
+-------- Hyriinae
--+
| +----- Velesunio_ambiguus **
+--+
| +-- Velesunio_angasi **
+--+
+-- Lortiella **
Klunzinger et al., 2022 [Publ 4259], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
1 |
1 |
1 |
22.0 |
+----- Hyriinae
--+
| +-- Velesunio **
+--+
+-- Westralunio **
Hoeh et al., 2009 [Publ 2716], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
44 |
6 |
1 |
20.5 |
+-------- Hyriinae
--+
| +----- Velesunio_ambiguus **
+--+
| +-- Velesunio_angasi **
+--+
+-- Lortiella **
Hoeh et al., 2009 [Publ 2716], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
43 |
6 |
1 |
20.5 |
+-------- Hyriinae
--+
| +----- Velesunio_ambiguus **
+--+
| +-- Velesunio_angasi **
+--+
+-- Lortiella **
Hoeh et al., 2009 [Publ 2716], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
43 |
6 |
1 |
20.5 |
+-------- Hyriinae
--+
| +----- Velesunio_ambiguus **
+--+
| +-- Velesunio_angasi **
+--+
+-- Lortiella **
Marshall et al., 2014 [Publ 3593], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
11 |
7 |
1 |
22.0 |
+----------- Echyridella
|
--+ +-- Hyridellini
| +-----+
| | +-- Rhipidodontini
+--+
| +----- Velesunio_ambiguus **
+--+
| +-- Velesunio **
+--+
+-- Lortiella **
Hoeh et al., 2002 [Publ 2653], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
3 |
1 |
-- |
+-- Castaliini
+-----+
| +-- Rhipidodontini
--+
| +----- Echyridella
+--+
| +-- Velesunio **
+--+
+-- Lortiella **
Hoeh et al., 2001 [Publ 2652], fig. 14f4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
3 |
1 |
-- |
+-- Castaliini
+-----+
| +-- Rhipidodontini
--+
| +----- Echyridella
+--+
| +-- Lortiella **
+--+
+-- Velesunio **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
5 |
0 |
2 |
29 |
3 |
1 |
21.5 |
+----- Rhipidodontini
|
+----- Echyridella
--+
+----- Lortiella **
|
| +-- Castaliini
+--+
+-- Velesunio **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
32 |
3 |
1 |
-- |
+-- Castaliini
+-----+
| +-- Rhipidodontini
--+
| +----- Echyridella
+--+
| +-- Velesunio **
+--+
+-- Lortiella **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
3 |
1 |
-- |
+-- Castaliini
+-----+
| +-- Rhipidodontini
--+
| +----- Echyridella
+--+
| +-- Velesunio **
+--+
+-- Lortiella **
Baker et al., 2003 [Publ 2690], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
6 |
5 |
1 |
-- |
+-- Hyridella_australis
+--------+
| +-- Rhipidodontini
--+
| +-------- Hyridella_depressa
+--+
| +----- Velesunio **
+--+
| +-- Echyridella
+--+
+-- Castaliini
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
5 |
0 |
2 |
29 |
3 |
1 |
22.5 |
+-- Castaliini
+--+
| +-- Velesunio **
--+
| +-- Echyridella
| |
+--+-- Lortiella **
|
+-- Rhipidodontini
Graf, 2002 [Publ 2651], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
17 |
3 |
1 |
-- |
+-- Velesunio **
--+
+-- Hyriinae
Whelan et al., 2011 [Publ 3380], fig. 1-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
35 |
3 |
1 |
-- |
+-- Velesunio **
--+
+-- Hyriinae
Wu et al., 2021 [Publ 4366], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
90 |
2 |
1 |
-- |
+-- Alathyria **
--+
+-- Hyriinae
Bieler et al., 2014 [Publ 3622], fig. 28
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Sonowal et al., 2021 [Publ 4221], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Velesunio **
+--+
+-- Hyridellini
Sonowal et al., 2021 [Publ 4221], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
+-- Hyriini
|
--+-- Hyridellini
|
+-- Velesunio **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Combosch et al., 2017 [Publ 3806], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
+-- Velesunio **
--+
+-- Hyriinae
Wu et al., 2020 [Publ 4121], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Hyridellini
+--+
+-- Velesunio **
Combosch et al., 2017 [Publ 3806], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Velesunio **
+--+
+-- Hyridellini
Whelan et al., 2011 [Publ 3380], fig. 1-MP
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
35 |
3 |
1 |
-- |
+-- Velesunio **
--+
+-- Hyriinae
González & Giribet, 2015 [Publ 3632], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Wu et al., 2020 [Publ 4121], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Hyridellini
+--+
+-- Velesunio **
Huang et al., 2018 [Publ 3828], fig. 3_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
28 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Velesunio **
+--+
+-- Hyridellini
González & Giribet, 2015 [Publ 3632], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Guerra et al., 2019 [Publ 4168], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
1 |
1 |
-- |
+-- Hyriinae
--+
+-- Westralunio **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
79 |
1 |
1 |
-- |
+-- Hyriinae
--+
+-- Velesunio **
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
68 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Hyridellini
+--+
+-- Velesunio **
Sonowal et al., 2021 [Publ 4221], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
+-- Hyriini
|
--+-- Velesunio **
|
+-- Hyridellini
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
68 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Hyridellini
+--+
+-- Velesunio **
Elderkin et al., 2016 [Publ 3693], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
119 |
1 |
1 |
-- |
+-- Hyriinae
--+
+-- Velesunio **
Wu et al., 2021 [Publ 4366], fig. 2_I
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
90 |
2 |
1 |
-- |
+-- Alathyria **
--+
+-- Hyriinae
Plazzi et al., 2011 [Publ 3676], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
17 |
0 |
0 |
-- |
-- Alathyria_jacksoni
González & Giribet, 2015 [Publ 3632], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Velesunio_ambiguus
González & Giribet, 2015 [Publ 3632], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Velesunio_ambiguus
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
3 |
1 |
-- |
+-- Velesunio **
--+
+-- Hyriinae
Guerra et al., 2019 [Publ 4168], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
40 |
4 |
1 |
-- |
+-- Echyridella
+-----+
| +-- Westralunio **
--+
| +----- Rhipidodontini
+--+
| +-- Castaliini
+--+
+-- Hyriini
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Velesunio **
+--+
+-- Hyridellini
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
68 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Hyridellini
+--+
+-- Velesunio **
Graf, 2000 [Publ 2647], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
3 |
1 |
-- |
+-------- Castaliini
--+
| +----- Rhipidodontini
+--+
| +-- Hyridellini
+--+
+-- Velesunio **
Guerra et al., 2019 [Publ 4168], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
40 |
4 |
1 |
-- |
+-- Westralunio **
--+
+-- Hyriinae
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
5 |
1 |
-- |
+-- Velesunio **
--+
+-- Hyriinae
Guerra et al., 2019 [Publ 4168], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
1 |
1 |
-- |
+-- Hyriinae
--+
+-- Westralunio **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
2 |
1 |
-- |
+----- Hyriini
--+
| +-- Velesunio **
+--+
+-- Hyridellini
|