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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Truncilla Rafinesque, 1819: cladomics
     4 valid species

Cladomics Summary

number of species = 4

analyzed in 38 cladograms in 24 publications
maximum number of terminals in an analysis = 4, mean = 1.55

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 9
maximum number of included terminals = 4, mean = 3.33
proportion of cladograms that support basic monophyly = 1.0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 37 32 19 0.0

*** Top Monophyly Cladogram ***

  +-------------------------- Obliquaria             
  |                                                  
  |                       +-- Medionidus             
  |                    +--+                          
  |                    |  +-- Lemiox                 
  |  +-----------------+                             
--+  |                 |  +-- Cyprogenia             
  |  |                 +--+                          
  |  |                    +-- Ptychobranchus         
  |  |                                               
  |  |     +----------------- Potamilus              
  |  |  +--+                                         
  +--+  |  |  +-------------- Atlanticoncha          
     |  |  +--+                                      
     |  |     |  +----------- Ellipsaria             
     |  |     +--+                                   
     |  |        |  +-------- Truncilla_truncata **     
     |  |        +--+                                
     |  |           |  +----- Truncilla_cognata **      
     +--+           +--+                             
        |              |  +-- Truncilla_macrodon **     
        |              +--+                          
        |                 +-- Truncilla_donaciformis ** 
        |                                            
        |                 +-- Hamiota                
        |     +-----------+                          
        |     |           +-- Villosa                
        |     |                                      
        +-----+  +----------- Ligumia                
              |  |                                   
              |  |     +----- Venustaconcha          
              +--+  +--+                             
                 |  |  |  +-- Obovaria               
                 |  |  +--+                          
                 +--+     +-- Epioblasma             
                    |                                
                    |  +----- Ortmanniana_pectorosa  
                    +--+                             
                       |  +-- Ortmanniana_ligamentina
                       +--+                          
                          +-- Lampsilis              

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 55 47 22 --

*** Top Sister-Group Cladogram ***

                                   +-- Cyrtonaias             
  +--------------------------------+                          
  |                                +-- Glebula                
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus 
           |  |              |  |                             
           |  |              +--+----- Lemiox                 
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla_truncata **     
              |  |              +--+                          
              |  |                 +-- Ellipsaria             
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia                

Smith et al., 2020 [Publ 4129], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 24 16 7 2.0
  +----------------------- Obliquaria            
  |                                              
--+  +-------------------- Cyrtonaias            
  |  |                                           
  |  |                 +-- Pachynaias            
  +--+  +--------------+                         
     |  |              +-- Lampsilis             
     |  |                                        
     +--+              +-- Atlanticoncha         
        |  +-----------+                         
        |  |           +-- Potamilus             
        +--+                                     
           |  +----------- Ellipsaria            
           +--+                                  
              |  +-------- Truncilla_truncata **    
              +--+                               
                 |  +----- Truncilla_donaciformis **
                 +--+                            
                    |  +-- Truncilla_cognata **     
                    +--+                         
                       +-- Truncilla_macrodon **    

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
4 4 0 4 29 22 17 2.0
  +----------------- Toxolasma             
  |                                        
  |  +-------------- Potamilus             
  |  |                                     
  |  |        +----- Medionidus            
  |  +--------+                            
  |  |        |  +-- Lemiox                
--+  |        +--+                         
  |  |           +-- Dromus                
  |  |                                     
  |  |        +----- Ellipsaria            
  |  |        |                            
  |  +--------+  +-- Truncilla_cognata **     
  |  |        |  |                         
  |  |        |  +-- Truncilla_donaciformis **
  +--+        +--+                         
     |           +-- Truncilla_macrodon **    
     |           |                         
     |           +-- Truncilla_truncata **    
     |                                     
     |  +----------- Cambarunio            
     |  |                                  
     |  +----------- Leaunio               
     |  |                                  
     |  +----------- Villosa               
     |  |                                  
     |  |        +-- Sagittunio            
     |  +--------+                         
     +--+        +-- Lampsilis_ovata       
        |                                  
        |     +----- Venustaconcha         
        +-----+                            
        |     |  +-- Epioblasma            
        |     +--+                         
        |        +-- Obovaria              
        |                                  
        |  +-------- Lampsilis_teres       
        +--+                               
           |  +----- Lampsilis_siliquoidea 
           +--+                            
              |  +-- Lampsilis_hydiana     
              +--+                         
                 +-- Lampsilis_straminea   

Saxon et al., 2023 [Publ 4338], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 1 1 1 2.0
  +----------- Ellipsaria            
--+                                  
  |  +-------- Truncilla_truncata **    
  +--+                               
     |  +----- Truncilla_macrodon **    
     +--+                            
        |  +-- Truncilla_cognata **     
        +--+                         
           +-- Truncilla_donaciformis **

Smith et al., 2020 [Publ 4129], fig. 2-3

iT tT bi riS oT roT roS iD
4 4 2 4 24 16 7 2.0
  +----------------------- Obliquaria            
  |                                              
--+  +-------------------- Cyrtonaias            
  |  |                                           
  |  |                 +-- Pachynaias            
  +--+  +--------------+                         
     |  |              +-- Lampsilis             
     |  |                                        
     +--+              +-- Atlanticoncha         
        |  +-----------+                         
        |  |           +-- Potamilus             
        +--+                                     
           |  +----------- Ellipsaria            
           +--+                                  
              |  +-------- Truncilla_truncata **    
              +--+                               
                 |  +----- Truncilla_donaciformis **
                 +--+                            
                    |  +-- Truncilla_cognata **     
                    +--+                         
                       +-- Truncilla_macrodon **    

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 39 24 6 3.0
  +----------------------------------- Toxolasma             
  |                                                          
  |  +-------------------------------- Potamilus             
--+  |                                                       
  |  |                       +-------- Ellipsaria            
  |  |  +--------------------+                               
  +--+  |                    |  +----- Truncilla_macrodon **    
     |  |                    +--+                            
     |  |                       |  +-- Truncilla_donaciformis **
     |  |                       +--+                         
     |  |                          +-- Truncilla_cognata **     
     +--+                                                    
        |  +-------------------------- Leaunio               
        |  |                                                 
        |  |     +-------------------- Lampsilis_bracteata   
        |  |     |                                           
        |  |  +--+  +----------------- Lampsilis_fasciola    
        |  |  |  |  |                                        
        +--+  |  +--+  +-------------- Lampsilis_ornata      
           |  |     |  |                                     
           |  |     +--+           +-- Lampsilis_ovata       
           |  |        |  +--------+                         
           |  |        |  |        +-- Lampsilis_cariosa     
           |  |        +--+                                  
           |  |           |  +-------- Lampsilis_cardium__3  
           +--+           +--+                               
              |              |  +----- Lampsilis_cardium__2  
              |              +--+                            
              |                 |  +-- Lampsilis_satura      
              |                 +--+                         
              |                    +-- Lampsilis_cardium__1  
              |                                              
              |                    +-- Lampsilis_higginsii   
              |     +--------------+                         
              |     |              +-- Ortmanniana           
              +-----+                                        
                    |  +-------------- Lampsilis_teres       
                    |  |                                     
                    +--+           +-- Lampsilis_radiata     
                       |  +--------+                         
                       |  |        +-- Lampsilis_siliquoidea 
                       +--+                                  
                          |  +-------- Lampsilis_bergmanni   
                          +--+                               
                             |  +----- Lampsilis_hydiana     
                             +--+                            
                                |  +-- Lampsilis_virescens   
                                +--+                         
                                   +-- Lampsilis_straminea   

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 38 23 6 1.0
  +----------------------------------- Toxolasma             
  |                                                          
--+  +-------------------------------- Ellipsaria            
  |  |                                                       
  |  |                          +----- Truncilla_cognata **     
  +--+  +-----------------------+                            
     |  |                       |  +-- Truncilla_macrodon **    
     |  |                       +--+                         
     |  |                          +-- Truncilla_donaciformis **
     +--+                                                    
        |  +-------------------------- Potamilus             
        |  |                                                 
        |  |  +----------------------- Leaunio               
        |  |  |                                              
        +--+  |     +----------------- Lampsilis_bracteata   
           |  |     |                                        
           |  |  +--+              +-- Lampsilis_ornata      
           |  |  |  |  +-----------+                         
           +--+  |  |  |           +-- Lampsilis_fasciola    
              |  |  +--+                                     
              |  |     |  +----------- Lampsilis_ovata       
              |  |     +--+                                  
              |  |        |  +-------- Lampsilis_cardium__3  
              |  |        +--+                               
              +--+           |  +----- Lampsilis_cardium__2  
                 |           +--+                            
                 |              |  +-- Lampsilis_satura      
                 |              +--+                         
                 |                 +-- Lampsilis_cardium__1  
                 |                                           
                 |     +-------------- Lampsilis_teres       
                 |     |                                     
                 |     |           +-- Lampsilis_higginsii   
                 +-----+  +--------+                         
                       |  |        +-- Ortmanniana           
                       |  |                                  
                       +--+        +-- Lampsilis_radiata     
                          |  +-----+                         
                          |  |     +-- Lampsilis_siliquoidea 
                          |  |                               
                          +--+     +-- Lampsilis_hydiana     
                             |  +--+                         
                             |  |  +-- Lampsilis_bergmanni   
                             +--+                            
                                |  +-- Lampsilis_virescens   
                                +--+                         
                                   +-- Lampsilis_straminea   

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 36 14 9 --
  +----------------- Potamilus             
  |                                        
  |           +----- Ellipsaria            
--+  +--------+                            
  |  |        |  +-- Truncilla_donaciformis **
  |  |        +--+                         
  |  |           +-- Truncilla_truncata **    
  +--+                                     
     |  +----------- Lampsilis_cardium     
     |  |                                  
     |  |     +----- Ligumia               
     |  +-----+                            
     +--+     |  +-- Obovaria              
        |     +--+                         
        |        +-- Epioblasma            
        |                                  
        |  +-------- Lampsilis_teres       
        +--+                               
           |  +----- Lampsilis_siliquoidea 
           +--+                            
              |  +-- Ortmanniana           
              +--+                         
                 +-- Lampsilis_higginsii   

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 33 14 9 --
  +-------------- Toxolasma             
  |                                     
  |  +----------- Obliquaria            
--+  |                                  
  |  |  +-------- Obovaria              
  |  |  |                               
  |  |  +-------- Lampsilis_cardium     
  +--+  |                               
     |  +-------- Potamilus             
     |  |                               
     |  |     +-- Epioblasma            
     |  +-----+                         
     |  |     +-- Lampsilis_siliquoidea 
     +--+                               
        |     +-- Ortmanniana           
        +-----+                         
        |     +-- Lampsilis_higginsii   
        |                               
        |     +-- Ligumia               
        +-----+                         
        |     +-- Lampsilis_teres       
        |                               
        |  +----- Ellipsaria            
        +--+                            
           |  +-- Truncilla_truncata **    
           +--+                         
              +-- Truncilla_donaciformis **

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla_truncata **            
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 24 7 7 --
  +----------- Truncilla_truncata **
  |                              
--+        +-- Lampsilis         
  |  +-----+                     
  |  |     +-- Epioblasma        
  +--+                           
     |  +----- Ligumia           
     +--+                        
        |  +-- Obliquaria        
        +--+                     
           +-- Ortmanniana       

Inoue et al., 2020 [Publ 4128], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 24 10 9 --
  +-- Epioblasma        
--+                     
  +-- Truncilla_truncata **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Truncilla_truncata **
--+                     
  +-- Lampsilis         

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 3 3 --
  +-------- Obliquaria        
--+                           
  |  +----- Potamilus         
  +--+                        
     |  +-- Lampsilis         
     +--+                     
        +-- Truncilla_truncata **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 11 8 --
  +----------- Ligumia              
  |                                 
  +----------- Truncilla_truncata **   
  |                                 
  |        +-- Ptychobranchus       
  +--------+                        
--+        +-- Sagittunio           
  |                                 
  |        +-- Lampsilis_siliquoidea
  |  +-----+                        
  |  |     +-- Epioblasma           
  |  |                              
  +--+  +----- Ortmanniana          
     |  |                           
     |  +----- Cambarunio           
     +--+                           
        +----- Leaunio              
        |                           
        |  +-- Lampsilis_cardium    
        +--+                        
           +-- Lampsilis_fasciola   

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
  +-- Lampsilis         
--+                     
  +-- Truncilla_truncata **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
  +----- Ellipsaria        
  |                        
  +----- Potamilus_leptodon
  |                        
  +----- Medionidus        
  |                        
  +----- Potamilus_alatus  
  |                        
  +----- Ptychobranchus    
--+                        
  +----- Truncilla_truncata **
  |                        
  +----- Venustaconcha     
  |                        
  +----- Cambarunio        
  |                        
  +----- Leaunio           
  |                        
  |  +-- Ortmanniana       
  +--+                     
     +-- Lampsilis         

Inoue et al., 2020 [Publ 3941], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 15 15 1 --
  +-- Truncilla_macrodon **
--+                     
  +-- Lampsilis         

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Lampsilis         
--+                     
  +-- Truncilla_truncata **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 7 7 --
  +-------------- Obliquaria        
  |                                 
  +-------------- Truncilla_truncata **
--+                                 
  +-------------- Ptychobranchus    
  |                                 
  |  +----------- Ligumia           
  +--+                              
     |  +-------- Cambarunio        
     +--+                           
        |  +----- Epioblasma        
        +--+                        
           |  +-- Lampsilis         
           +--+                     
              +-- Ortmanniana       

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
  +-------------- Potamilus_leptodon
  |                                 
  |  +----------- Potamilus_alatus  
--+  |                              
  |  |  +-------- Venustaconcha     
  |  |  |                           
  +--+  +-------- Leaunio           
     |  |                           
     |  |     +-- Ellipsaria        
     |  +-----+                     
     +--+     +-- Truncilla_truncata **
        |                           
        |     +-- Medionidus        
        +-----+                     
        |     +-- Ptychobranchus    
        |                           
        |  +----- Cambarunio        
        +--+                        
           |  +-- Ortmanniana       
           +--+                     
              +-- Lampsilis         

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 13 11 --
  +-------------------- Potamilus_leptodon
  |                                       
--+  +----------------- Potamilus_alatus  
  |  |                                    
  |  |              +-- Medionidus        
  +--+  +-----------+                     
     |  |           +-- Ptychobranchus    
     |  |                                 
     +--+           +-- Ellipsaria        
        |  +--------+                     
        |  |        +-- Truncilla_truncata **
        +--+                              
           |  +-------- Leaunio           
           |  |                           
           +--+     +-- Venustaconcha     
              |  +--+                     
              |  |  +-- Cambarunio        
              +--+                        
                 |  +-- Ortmanniana       
                 +--+                     
                    +-- Lampsilis         

Szumowski et al., 2012 [Publ 3569], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 14 9 --
  +----------------------- Ellipsaria           
  |                                             
--+  +-------------------- Truncilla_truncata **   
  |  |                                          
  +--+  +----------------- Obliquaria           
     |  |                                       
     |  |  +-------------- Potamilus            
     +--+  |                                    
        |  |        +----- Lampsilis_ornata     
        |  |  +-----+                           
        |  |  |     |  +-- Lampsilis_cardium    
        +--+  |     +--+                        
           |  |        +-- Lampsilis_ovata      
           |  |                                 
           |  |  +-------- Lampsilis_teres      
           |  +--+                              
           +--+  |  +----- Lampsilis_siliquoidea
              |  +--+                           
              |     |  +-- Ortmanniana          
              |     +--+                        
              |        +-- Lampsilis_higginsii  
              |                                 
              |     +----- Hamiota              
              |     |                           
              +-----+  +-- Epioblasma           
                    |  |                        
                    +--+-- Cambarunio           
                       |                        
                       +-- Ligumia              

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 5 --
        +-- Cyrtonaias        
  +-----+                     
  |     +-- Glebula           
  |                           
--+     +-- Lampsilis         
  |  +--+                     
  |  |  +-- Leaunio           
  +--+                        
     |  +-- Truncilla_macrodon **
     +--+                     
        +-- Potamilus         

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 56 47 22 --
                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla_truncata **     
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis 

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 18 3 3 --
  +-------- Obliquaria        
--+                           
  |  +----- Truncilla_truncata **
  +--+                        
     |  +-- Lampsilis         
     +--+                     
        +-- Potamilus         

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 7 7 --
  +----------------- Ptychobranchus    
--+                                    
  |  +-------------- Truncilla_truncata **
  +--+                                 
     |  +----------- Ortmanniana       
     +--+                              
        |  +-------- Lampsilis         
        +--+                           
           |  +----- Cambarunio        
           +--+                        
              |  +-- Ligumia           
              +--+                     
                 +-- Epioblasma        

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 11 7 --
  +----------------- Toxolasma         
  |                                    
--+  +-------------- Ellipsaria        
  |  |                                 
  +--+  +----------- Truncilla_macrodon **
     |  |                              
     +--+        +-- Lemiox            
        |  +-----+                     
        |  |     +-- Potamilus         
        +--+                           
           |  +----- Lampsilis         
           +--+                        
              |  +-- Cambarunio        
              +--+                     
                 +-- Venustaconcha     

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 3 3 --
  +-------- Obliquaria        
--+                           
  |  +----- Potamilus         
  +--+                        
     |  +-- Lampsilis         
     +--+                     
        +-- Truncilla_truncata **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +----- Potamilus         
--+                        
  |  +-- Lampsilis         
  +--+                     
     +-- Truncilla_truncata **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 34 11 10 --
  +-- Truncilla_truncata **
--+                     
  +-- Ptychobranchus    

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 11 8 --
  +-------------------- Truncilla_truncata **   
  |                                          
--+  +----------------- Ligumia              
  |  |                                       
  |  |              +-- Ptychobranchus       
  +--+  +-----------+                        
     |  |           +-- Sagittunio           
     |  |                                    
     +--+  +----------- Lampsilis_siliquoidea
        |  |                                 
        |  |     +----- Ortmanniana          
        +--+  +--+                           
           |  |  |  +-- Lampsilis_cardium    
           |  |  +--+                        
           +--+     +-- Lampsilis_fasciola   
              |                              
              |  +----- Epioblasma           
              +--+                           
                 |  +-- Cambarunio           
                 +--+                        
                    +-- Leaunio              

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 6 6 --
  +----- Truncilla_truncata **
  |                        
  +----- Ptychobranchus    
--+                        
  |  +-- Ortmanniana       
  |  |                     
  |  +-- Cambarunio        
  +--+                     
     +-- Ligumia           
     |                     
     +-- Epioblasma        

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla_truncata **            
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 7 7 --
  +----------------- Ptychobranchus    
--+                                    
  |  +-------------- Truncilla_truncata **
  +--+                                 
     |  +----------- Ortmanniana       
     +--+                              
        |  +-------- Lampsilis         
        +--+                           
           |  +----- Cambarunio        
           +--+                        
              |  +-- Ligumia           
              +--+                     
                 +-- Epioblasma        

Inoue et al., 2020 [Publ 4128], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 25 11 10 --
  +-- Ptychobranchus    
--+                     
  +-- Truncilla_truncata **

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 11 7 --
  +----------------- Toxolasma         
  |                                    
--+  +-------------- Venustaconcha     
  |  |                                 
  +--+           +-- Lemiox            
     |  +--------+                     
     |  |        +-- Lampsilis         
     +--+                              
        |  +-------- Potamilus         
        +--+                           
           |  +----- Cambarunio        
           +--+                        
              |  +-- Ellipsaria        
              +--+                     
                 +-- Truncilla_macrodon **

Page updated: 08:36:43 Wed 06 Nov 2024

 
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