|
Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Ptychobranchus Simpson, 1900: cladomics
8 valid species
Cladomics Summary
number of species = 8
analyzed in 31 cladograms in 18 publications
maximum number of terminals in an analysis = 5, mean = 1.13
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 1
maximum number of included terminals = 5, mean = 5.0
proportion of cladograms that support basic monophyly = 1.0
Roe, 2013 [Publ 3571], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
4 |
6 |
6 |
6 |
5 |
0.0 |
*** Top Monophyly Cladogram ***
+----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Medionidus
| |
+--+ +-- Cyprogenia
| +--------------+
| | +-- Dromus
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Ptychobranchus_jonesi **
+--+
| +-------- Ptychobranchus_greenii **
+--+
| +----- Ptychobranchus_occidentalis__2 **
+--+
| +-- Ptychobranchus_occidentalis__1 **
+--+
+-- Ptychobranchus_fasciolaris **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
55 |
47 |
22 |
-- |
*** Top Sister-Group Cladogram ***
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus_fasciolaris **
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Smith et al., 2019 [Publ 3951], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
40 |
35 |
19 |
-- |
+----------------------- Obliquaria
|
| +-- Medionidus
| +--+
| | +-- Lemiox
--+ +--------------+
| | | +-- Cyprogenia
| | +--+
| | +-- Ptychobranchus_fasciolaris **
| |
+--+ +-------- Potamilus
| +--------+
| | | +----- Atlanticoncha
| | +--+
| | | +-- Ellipsaria
| | +--+
+--+ +-- Truncilla
|
| +-- Hamiota
| +-----------+
| | +-- Villosa
| |
+--+ +----------- Ligumia
| |
| | +----- Venustaconcha
+--+ +--+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +----- Ortmanniana_pectorosa
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
197 |
13 |
11 |
-- |
+----- Ellipsaria
|
+----- Potamilus_leptodon
|
+----- Medionidus
|
+----- Potamilus_alatus
|
+----- Ptychobranchus_subtentus **
--+
+----- Truncilla
|
+----- Venustaconcha
|
+----- Cambarunio
|
+----- Leaunio
|
| +-- Ortmanniana
+--+
+-- Lampsilis
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
7 |
7 |
-- |
+----------------- Ptychobranchus_fasciolaris **
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
47 |
22 |
-- |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus_fasciolaris **
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
6 |
4 |
-- |
+----------------------- Obliquaria
|
+----------------------- Lampsilis
--+
| +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__5
| |
+--+ +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
7 |
6 |
-- |
+----------------- Medionidus
--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus_subtentus **
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis
+--+
| +-- Villosa
+--+
+-- Ellipsaria
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
7 |
7 |
-- |
+----------------- Ptychobranchus_fasciolaris **
--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
6 |
4 |
-- |
+-------------------------- Lampsilis
|
--+ +----------------------- Obliquaria
| |
+--+ +-------------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__6
| +--------------+
| | +-- Dromus
+--+
| +-------------- Cyprogenia_aberti__4
| |
+--+ +----------- Cyprogenia_aberti__3
| |
+--+ +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
11 |
8 |
-- |
+----------- Ligumia
|
+----------- Truncilla
|
| +-- Ptychobranchus_fasciolaris **
+--------+
--+ +-- Sagittunio
|
| +-- Lampsilis_siliquoidea
| +-----+
| | +-- Epioblasma
| |
+--+ +----- Ortmanniana
| |
| +----- Cambarunio
+--+
+----- Leaunio
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
195 |
13 |
11 |
-- |
+-------------------- Potamilus_leptodon
|
--+ +----------------- Potamilus_alatus
| |
| | +-- Medionidus
+--+ +-----------+
| | +-- Ptychobranchus_subtentus **
| |
+--+ +-- Ellipsaria
| +--------+
| | +-- Truncilla
+--+
| +-------- Leaunio
| |
+--+ +-- Venustaconcha
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
8 |
6 |
-- |
+-------------------- Obliquaria
--+
| +----------------- Medionidus
+--+
| +-------------- Ptychobranchus_subtentus **
+--+
| +----------- Potamilus_fragilis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ellipsaria
+--+
| +-- Villosa
+--+
+-- Lampsilis
Serb, 2006 [Publ 2678], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
6 |
4 |
-- |
+-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
6 |
6 |
-- |
+----- Truncilla
|
+----- Ptychobranchus_fasciolaris **
--+
| +-- Ortmanniana
| |
| +-- Cambarunio
+--+
+-- Ligumia
|
+-- Epioblasma
Serb, 2006 [Publ 2678], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
6 |
4 |
-- |
+-------------- Obliquaria
|
| +-- Ptychobranchus_fasciolaris **
+-----------+
--+ +-- Lampsilis
|
| +-- Cyprogenia_aberti__6
| +--------+
| | +-- Dromus
+--+
| +-------- Cyprogenia_aberti__5
| |
+--+ +----- Cyprogenia_aberti__4
| |
| | +-- Cyprogenia_stegaria__1
+--+ |
| +-- Cyprogenia_stegaria__2
| |
+--+-- Cyprogenia_aberti__1
|
+-- Cyprogenia_aberti__2
|
+-- Cyprogenia_aberti__3
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
12 |
10 |
-- |
+-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Serb, 2006 [Publ 2678], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
6 |
4 |
-- |
+-------------------- Obliquaria
|
+-------------------- Lampsilis
--+
| +----------------- Ptychobranchus_fasciolaris **
| |
+--+ +-- Cyprogenia_aberti__5
| +-----------+
| | +-- Dromus
+--+
| +----------- Cyprogenia_aberti__4
| |
+--+ +-------- Cyprogenia_aberti__3
| |
+--+ +----- Cyprogenia_stegaria__2
| |
+--+----- Cyprogenia_aberti__2
|
| +-- Cyprogenia_stegaria__1
+--+
+-- Cyprogenia_aberti__1
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
26 |
8 |
6 |
-- |
+-------------- Potamilus_fragilis
|
--+ +----------- Ptychobranchus_subtentus **
| |
+--+ +-------- Medionidus
| |
+--+ +----- Potamilus_purpuratus
| |
+--+----- Lampsilis
|
| +-- Ellipsaria
+--+
+-- Villosa
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
7 |
7 |
-- |
+-- Cambarunio
--+
+-- Ptychobranchus_fasciolaris **
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
32 |
10 |
7 |
-- |
+----- Ptychobranchus_subtentus **
+--------+
| | +-- Potamilus_fragilis
| +--+
| +-- Medionidus
--+
| +-- Villosa
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Inoue et al., 2020 [Publ 4128], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
25 |
11 |
10 |
-- |
+-- Ptychobranchus_fasciolaris **
--+
+-- Truncilla
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
101 |
42 |
22 |
-- |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus_fasciolaris **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Inoue et al., 2020 [Publ 4128], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
10 |
9 |
-- |
-- Ptychobranchus_fasciolaris
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
12 |
10 |
-- |
+-------------------------- Toxolasma
|
--+ +----------------------- Obliquaria
| |
| | +-- Cyrtonaias
+--+ +-----------------+
| | +-- Glebula
| |
+--+ +-- Ptychobranchus_fasciolaris **
| +--------------+
| | +-- Lemiox
| |
+--+ +-- Venustaconcha
| +-----------+
| | +-- Obovaria
+--+
| +----------- Hamiota
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
11 |
8 |
-- |
+-------------------- Truncilla
|
--+ +----------------- Ligumia
| |
| | +-- Ptychobranchus_fasciolaris **
+--+ +-----------+
| | +-- Sagittunio
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_fasciola
|
| +----- Epioblasma
+--+
| +-- Cambarunio
+--+
+-- Leaunio
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
7 |
6 |
-- |
+-- Villosa
|
+-- Ellipsaria
|
--+-- Lampsilis
|
+-- Potamilus
|
+-- Ptychobranchus_subtentus **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
101 |
42 |
22 |
-- |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus_fasciolaris **
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
34 |
11 |
10 |
-- |
+-- Truncilla
--+
+-- Ptychobranchus_fasciolaris **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
197 |
13 |
11 |
-- |
+-------------- Potamilus_leptodon
|
| +----------- Potamilus_alatus
--+ |
| | +-------- Venustaconcha
| | |
+--+ +-------- Leaunio
| |
| | +-- Ellipsaria
| +-----+
+--+ +-- Truncilla
|
| +-- Medionidus
+-----+
| +-- Ptychobranchus_subtentus **
|
| +----- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
7 |
7 |
-- |
+-------------- Obliquaria
|
+-------------- Truncilla
--+
+-------------- Ptychobranchus_fasciolaris **
|
| +----------- Ligumia
+--+
| +-------- Cambarunio
+--+
| +----- Epioblasma
+--+
| +-- Lampsilis
+--+
+-- Ortmanniana
| 
