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Page last updated
Sun 06 Oct 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Strophitus Rafinesque, 1820: cladomics
     8 valid species

Cladomics Summary

number of species = 8

analyzed in 51 cladograms in 34 publications
maximum number of terminals in an analysis = 5, mean = 1.35

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 5, mean = 4.0
proportion of cladograms that support basic monophyly = 1.0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Smith et al., 2018 [Publ 3790], fig. 1

iT tT bi riS oT roT roS iD
5 5 4 6 13 12 7 0.0

*** Top Monophyly Cladogram ***

                 +-- Utterbackia               
  +--------------+                             
  |              +-- Pyganodon                 
  |                                            
  |              +-- Alasmidonta               
  |           +--+                             
--+           |  +-- Alasminota                
  |  +--------+                                
  |  |        |  +-- Pseudodontoideus          
  |  |        +--+                             
  |  |           +-- Utterbackiana             
  +--+                                         
     |  +----------- Anodontoides              
     |  |                                      
     |  |     +----- Strophitus_pascagoulaensis **
     +--+  +--+                                
        |  |  |  +-- Strophitus_radiatus **       
        |  |  +--+                             
        +--+     +-- Strophitus_williamsi **      
           |                                   
           |  +----- Strophitus_howellsi__2 **    
           +--+                                
              |  +-- Strophitus_undulatus **      
              +--+                             
                 +-- Strophitus_howellsi__1 **    

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
4 4 3 5 37 29 12 5.0

*** Top Sister-Group Cladogram ***

                          +-- Utterbackia                  
  +-----------------------+                                
  |                       +-- Pyganodon                    
  |                                                        
  |                 +-------- Utterbackiana                
  |                 |                                      
  |  +--------------+     +-- Lasmigona_costata            
--+  |              |  +--+                                
  |  |              |  |  +-- Pseudodontoideus             
  |  |              +--+                                   
  |  |                 |  +-- Lasmigona_complanata         
  |  |                 +--+                                
  |  |                    +-- Arcidens                     
  +--+                                                     
     |                    +-- Prolasmidonta                
     |                 +--+                                
     |                 |  +-- Platynaias                   
     |  +--------------+                                   
     |  |              |  +-- Alasminota                   
     |  |              +--+                                
     +--+                 +-- Alasmidonta                  
        |                                                  
        |  +----------------- Pressodonta                  
        +--+                                               
           |  +-------------- Anodontoides                 
           +--+                                            
              |  +----------- Strophitus_undulatus **         
              +--+                                         
                 |  +-------- Strophitus_pascagoulaensis__2 **
                 +--+                                      
                    |  +----- Strophitus_radiatus **          
                    +--+                                   
                       |  +-- Strophitus_pascagoulaensis__1 **
                       +--+                                
                          +-- Strophitus_williamsi **         

Smith et al., 2023 [Publ 4291], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 2 2 1 1.0
  +----------- Anodontoides              
  |                                      
--+        +-- Strophitus_undulatus **      
  |  +-----+                             
  |  |     +-- Strophitus_howellsi **       
  +--+                                   
     |  +----- Strophitus_pascagoulaensis **
     +--+                                
        |  +-- Strophitus_radiatus **       
        +--+                             
           +-- Strophitus_williamsi **      

Whelan et al., 2023 [Publ 4287], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 23 21 10 4.0
  +----------------------- Pressodonta               
  |                                                  
  |              +-------- Utterbackiana             
--+  +-----------+                                   
  |  |           |  +----- Alasminota                
  |  |           +--+                                
  |  |              |  +-- Arcidens                  
  +--+              +--+                             
     |                 +-- Pseudodontoideus          
     |                                               
     |  +----------------- Alasmidonta               
     |  |                                            
     +--+  +-------------- Platynaias                
        |  |                                         
        |  |        +----- Pyganodon                 
        +--+  +-----+                                
           |  |     |  +-- Utterbackia               
           |  |     +--+                             
           +--+        +-- Anodontoides              
              |                                      
              |  +-------- Strophitus_pascagoulaensis **
              +--+                                   
                 |  +----- Strophitus_undulatus **      
                 +--+                                
                    |  +-- Strophitus_radiatus **       
                    +--+                             
                       +-- Strophitus_williamsi **      

Whelan et al., 2023 [Publ 4287], fig. 3

iT tT bi riS oT roT roS iD
4 4 3 5 24 23 11 3.0
  +----------------------------- Utterbackiana                
  |                                                           
  |                          +-- Utterbackia                  
--+  +-----------------------+                                
  |  |                       +-- Pyganodon                    
  |  |                                                        
  |  |                    +----- Alasmidonta                  
  +--+  +-----------------+                                   
     |  |                 |  +-- Arcidens                     
     |  |                 +--+                                
     |  |                    +-- Pseudodontoideus             
     +--+                                                     
        |  +-------------------- Alasminota                   
        |  |                                                  
        |  |              +----- Pressodonta                  
        +--+  +-----------+                                   
           |  |           |  +-- Platynaias                   
           |  |           +--+                                
           +--+              +-- Prolasmidonta                
              |                                               
              |  +-------------- Anodontoides                 
              +--+                                            
                 |  +----------- Strophitus_undulatus **         
                 +--+                                         
                    |  +-------- Strophitus_pascagoulaensis__2 **
                    +--+                                      
                       |  +----- Strophitus_pascagoulaensis__1 **
                       +--+                                   
                          |  +-- Strophitus_radiatus **          
                          +--+                                
                             +-- Strophitus_williamsi **         

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 31 7 6 --
  +-- Strophitus_radiatus ** 
--+                       
  +-- Strophitus_undulatus **

Hewitt et al., 2018 [Publ 3758], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 5 4 --
  +-------- Pyganodon           
  |                             
  |     +-- Platynaias          
--+-----+                       
  |     +-- Strophitus_undulatus **
  |                             
  |  +----- Lasmigona_costata   
  +--+                          
     |  +-- Arcidens            
     +--+                       
        +-- Lasmigona_complanata

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 4 4 --
        +-- Pyganodon           
  +-----+                       
  |     +-- Simpsonaias         
--+                             
  |  +----- Strophitus_undulatus **
  +--+                          
     |  +-- Alasmidonta         
     +--+                       
        +-- Lasmigona           

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 5 4 --
        +-- Utterbackia             
  +-----+                           
  |     +-- Utterbackiana_couperiana
--+                                 
  |  +----- Strophitus_radiatus **     
  +--+                              
     |  +-- Lasmigona               
     +--+                           
        +-- Alasmidonta             

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 4 4 --
        +-- Pyganodon           
  +-----+                       
  |     +-- Simpsonaias         
--+                             
  |  +----- Strophitus_undulatus **
  +--+                          
     |  +-- Alasmidonta         
     +--+                       
        +-- Lasmigona           

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 4 4 --
  +----- Pyganodon           
  |                          
  +----- Simpsonaias         
--+                          
  +----- Strophitus_undulatus **
  |                          
  |  +-- Alasmidonta         
  +--+                       
     +-- Lasmigona           

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Strophitus_undulatus **
--+                       
  +-- Pyganodon           

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
-- Strophitus_undulatus

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pyganodon           
--+                       
  +-- Strophitus_undulatus **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 82 4 4 --
  +----------- Pyganodon           
--+                                
  |  +-------- Simpsonaias         
  +--+                             
     |  +----- Strophitus_undulatus **
     +--+                          
        |  +-- Alasmidonta         
        +--+                       
           +-- Lasmigona           

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
1 1 0 1 103 15 8 --
-- Strophitus_undulatus

King et al., 1999 [Publ 2699], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 5 3 --
  +----- Lasmigona           
  |                          
--+  +-- Prolasmidonta       
  |  |                       
  +--+-- Strophitus_undulatus **
     |                       
     +-- Platynaias          

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 6 5 --
  +-- Simpsonaias         
--+                       
  +-- Strophitus_undulatus **

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 --
  +-- Pyganodon           
--+                       
  +-- Strophitus_undulatus **

King et al., 1999 [Publ 2699], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 8 8 4 --
  +----------- Pyganodon           
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Strophitus_undulatus **
     +--+                          
        |  +-- Prolasmidonta       
        +--+                       
           +-- Platynaias          

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 3 --
  +-------- Lasmigona           
--+                             
  |  +----- Alasmidonta         
  +--+                          
     |  +-- Strophitus_undulatus **
     +--+                       
        +-- Pyganodon           

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 4 4 --
  +----- Pyganodon           
  |                          
  +----- Simpsonaias         
--+                          
  +----- Strophitus_undulatus **
  |                          
  |  +-- Alasmidonta         
  +--+                       
     +-- Lasmigona           

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 6 5 --
-- Strophitus_undulatus

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 2 --
  +----- Strophitus_undulatus **
--+                          
  |  +-- Platynaias          
  +--+                       
     +-- Pyganodon           

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 124 4 4 --
-- Strophitus_undulatus

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 --
-- Strophitus_radiatus

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 6 5 --
-- Strophitus_undulatus

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 3 --
  +-------- Lasmigona           
--+                             
  |  +----- Alasmidonta         
  +--+                          
     |  +-- Pyganodon           
     +--+                       
        +-- Strophitus_undulatus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 2 2 --
  +-- Pyganodon           
--+                       
  +-- Strophitus_undulatus **

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 4 --
  +----------- Utterbackiana      
--+                               
  |  +-------- Utterbackia        
  +--+                            
     |  +----- Strophitus_radiatus **
     +--+                         
        |  +-- Alasmidonta        
        +--+                      
           +-- Lasmigona          

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 24 3 3 --
  +----- Lasmigona           
  |                          
--+  +-- Strophitus_undulatus **
  |  |                       
  +--+-- Alasmidonta         
     |                       
     +-- Pyganodon           

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 --
  +----- Lasmigona           
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Pyganodon           

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 17 7 7 --
  +----- Strophitus_undulatus **
--+                          
  |  +-- Prolasmidonta       
  +--+                       
     +-- Platynaias          

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 4 3 --
     +-- Alasmidonta         
  +--+                       
  |  +-- Lasmigona           
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Pyganodon           

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 4 4 --
        +-- Alasmidonta         
  +-----+                       
  |     +-- Lasmigona           
--+                             
  |  +----- Pyganodon           
  +--+                          
     |  +-- Strophitus_undulatus **
     +--+                       
        +-- Simpsonaias         

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 3 --
  +-------- Lasmigona           
--+                             
  |  +----- Alasmidonta         
  +--+                          
     |  +-- Strophitus_undulatus **
     +--+                       
        +-- Pyganodon           

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 10 5 --
           +-- Pyganodon           
  +--------+                       
  |        +-- Utterbackia         
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Alasmidonta         
     +--+                          
        |  +-- Strophitus_undulatus **
        +--+                       
           +-- Platynaias          

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
-- Strophitus_undulatus

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
  +-- Strophitus_undulatus **
--+                       
  +-- Pyganodon           

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 3 --
  +-------- Lasmigona           
--+                             
  |  +----- Alasmidonta         
  +--+                          
     |  +-- Strophitus_undulatus **
     +--+                       
        +-- Pyganodon           

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pyganodon           
--+                       
  +-- Strophitus_undulatus **

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 8 7 --
  +-------------- Strophitus_undulatus **
  |                                   
  |        +----- Lasmigona_complanata
--+  +-----+                          
  |  |     |  +-- Lasmigona_costata   
  |  |     +--+                       
  +--+        +-- Arcidens            
     |                                
     |  +-------- Utterbackia         
     |  |                             
     +--+-------- Alasmidonta         
        |                             
        |  +----- Pyganodon           
        +--+                          
           |  +-- Anodontoides        
           +--+                       
              +-- Platynaias          

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 2 2 --
  +----- Lasmigona           
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Pyganodon           

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 10 5 --
  +----------- Lasmigona           
  |                                
--+  +-------- Strophitus_undulatus **
  |  |                             
  +--+     +-- Platynaias          
     |  +--+                       
     |  |  +-- Alasmidonta         
     +--+                          
        |  +-- Pyganodon           
        +--+                       
           +-- Utterbackia         

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 4 3 --
  +----- Strophitus_undulatus **
  |                          
--+  +-- Platynaias          
  |  |                       
  +--+-- Alasmidonta         
     |                       
     +-- Pyganodon           

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 4 4 --
        +-- Lasmigona           
  +-----+                       
  |     +-- Alasmidonta         
--+                             
  |  +----- Simpsonaias         
  +--+                          
     |  +-- Pyganodon           
     +--+                       
        +-- Strophitus_undulatus **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 4 3 --
     +-- Alasmidonta         
  +--+                       
  |  +-- Lasmigona           
--+                          
  |  +-- Strophitus_undulatus **
  +--+                       
     +-- Pyganodon           

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
1 1 0 1 19 13 7 --
     +-------- Pseudodontoideus    
  +--+                             
  |  |  +----- Lasmigona_costata   
  |  +--+                          
  |     |  +-- Lasmigona_complanata
--+     +--+                       
  |        +-- Arcidens            
  |                                
  |  +-------- Strophitus_undulatus **
  |  |                             
  +--+     +-- Pyganodon           
     |  +--+                       
     |  |  +-- Utterbackia         
     +--+                          
        |  +-- Platynaias          
        +--+                       
           +-- Alasmidonta         

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 4 4 --
  +----- Strophitus_radiatus **
--+                         
  |  +-- Lasmigona          
  +--+                      
     +-- Alasmidonta        

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 2 2 --
  +-- Pyganodon           
--+                       
  +-- Strophitus_undulatus **

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 5 4 --
  +----- Alasmidonta         
  |                          
  +----- Pyganodon           
--+                          
  +----- Lasmigona           
  |                          
  |  +-- Platynaias          
  +--+                       
     +-- Strophitus_undulatus **

Page updated: 10:16:04 Sun 06 Oct 2024

 
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