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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Rectidentini

genus Hyriopsis Conrad, 1853: cladomics
     10 valid species

Cladomics Summary

number of species = 10

analyzed in 50 cladograms in 31 publications
maximum number of terminals in an analysis = 9, mean = 2.82

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 27
maximum number of included terminals = 9, mean = 4.37
proportion of cladograms that support basic monophyly = 0.33


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
9 10 8 9 73 6 2 0.0

*** Top Monophyly Cladogram ***

  +----------------------- Ensidens               
  |                                               
--+                    +-- Hyriopsis_desowitzi **    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei **   
  +--+                                            
     |  +----------------- Rectidens              
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai **       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana **    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis **  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata **      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis **  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis **
                    +--+                          
                       +-- Hyriopsis_kratiensis **   

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iT tT bi riS oT roT roS iD
6 6 4 6 23 7 2 6.0

*** Top Sister-Group Cladogram ***

  +-------------------- Ensidens             
--+                                          
  |  +----------------- Rectidens            
  +--+                                       
     |  +-------------- Hyriopsis_delaportei ** 
     +--+                                    
        |  +----------- Hyriopsis_myersiana **  
        +--+                                 
           |  +-------- Hyriopsis_bialata **    
           +--+                              
              |  +----- Hyriopsis_khoratensis **
              +--+                           
                 |  +-- Hyriopsis_kratiensis ** 
                 +--+                        
                    +-- Hyriopsis_sakhonensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iT tT bi riS oT roT roS iD
9 9 5 9 2 1 1 7.0
  +-------------- Rectidens              
  |                                      
  |           +-- Hyriopsis_delaportei **   
--+  +--------+                          
  |  |        +-- Hyriopsis_desowitzi **    
  |  |                                   
  +--+  +-------- Hyriopsis_khoratensis **  
     |  |                                
     |  |     +-- Hyriopsis_kratiensis **   
     |  +-----+                          
     +--+     +-- Hyriopsis_phuphaniensis **
        |                                
        |  +----- Hyriopsis_sakhonensis **  
        |  |                             
        +--+  +-- Hyriopsis_myersiana **    
           |  |                          
           +--+-- Hyriopsis_panhai **       
              |                          
              +-- Hyriopsis_bialata **      

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iT tT bi riS oT roT roS iD
8 9 7 8 13 3 2 7.0
  +----------------- Ensidens               
  |                                         
  |              +-- Hyriopsis_desowitzi **    
--+  +-----------+                          
  |  |           +-- Hyriopsis_delaportei **   
  |  |                                      
  +--+  +----------- Rectidens              
     |  |                                   
     |  |     +----- Hyriopsis_myersiana **    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata **      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai **       
           |                                
           |  +----- Hyriopsis_khoratensis **  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis **
              +--+                          
                 +-- Hyriopsis_sakhonensis **  

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
5 6 4 5 107 7 2 4.0
  +----------------- Ensidens             
--+                                       
  |  +-------------- Hyriopsis_desowitzi **  
  +--+                                    
     |  +----------- Rectidens            
     +--+                                 
        |  +-------- Hyriopsis_panhai **     
        +--+                              
           |  +----- Hyriopsis_myersiana **  
           +--+                           
              |  +-- Hyriopsis_khoratensis **
              +--+                        
                 +-- Hyriopsis_bialata **    

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
5 6 4 5 134 6 2 10.0
  +-------------- Ensidens             
  |                                    
--+           +-- Hyriopsis_bialata **    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis **
  +--+                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Rectidens            
           +--+                        
              +-- Hyriopsis_desowitzi **  

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 11 6 5 193 6 2 12.0
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Hyriopsis_myersiana **  
  |                           
  +----- Hyriopsis_panhai **     
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
  |  +-- Hyriopsis_khoratensis **
  |                           
  |  +-- Rectidens            
  +--+                        
     +-- Ensidens             

Konopleva et al., 2019 [Publ 4041], fig. 3

iT tT bi riS oT roT roS iD
5 12 0 5 29 7 2 15.0
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Hyriopsis_bialata **    
  |                        
  +-- Hyriopsis_myersiana **  
  |                        
--+-- Hyriopsis_desowitzi **  
  |                        
  +-- Rectidens            
  |                        
  +-- Hyriopsis_panhai **     
  |                        
  +-- Ensidens_jaculus__1  
  |                        
  +-- Ensidens_jaculus__2  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius 

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 6 4 5 191 6 2 8.0
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_panhai **     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis **
        +--+                           
           |  +-- Hyriopsis_bialata **    
           +--+                        
              +-- Hyriopsis_myersiana **  

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
5 6 2 5 26 6 2 6.0
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis **
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_myersiana **  

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
5 6 3 5 47 6 2 5.0
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+----------- Rectidens            
     |                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Hyriopsis_khoratensis **
           +--+                        
              +-- Hyriopsis_bialata **    

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
5 6 2 5 120 6 2 6.0
  +----------- Ensidens             
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens            
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_khoratensis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
5 6 4 5 113 7 2 6.0
  +-------------- Ensidens             
  |                                    
--+  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens            
     |  |                              
     +--+     +-- Hyriopsis_khoratensis **
        |  +--+                        
        |  |  +-- Hyriopsis_bialata **    
        +--+                           
           |  +-- Hyriopsis_myersiana **  
           +--+                        
              +-- Hyriopsis_panhai **     

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
5 11 7 5 134 6 2 14.0
        +-- Hyriopsis_desowitzi **  
        |                        
  +-----+-- Rectidens            
  |     |                        
  |     +-- Ensidens             
--+                              
  |  +----- Hyriopsis_panhai **     
  |  |                           
  +--+----- Hyriopsis_khoratensis **
     |                           
     |  +-- Hyriopsis_myersiana **  
     +--+                        
        +-- Hyriopsis_bialata **    

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iT tT bi riS oT roT roS iD
5 5 2 5 1 1 1 8.0
  +----------- Rectidens            
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  +--+  +----- Hyriopsis_khoratensis **
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_panhai **     
           |                        
           +-- Hyriopsis_bialata **    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 11 5 5 193 6 2 12.0
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens            
  |                           
--+----- Ensidens             
  |                           
  |  +-- Hyriopsis_bialata **    
  |  |                        
  |  +-- Hyriopsis_myersiana **  
  +--+                        
     +-- Hyriopsis_panhai **     
     |                        
     +-- Hyriopsis_khoratensis **

Muanta et al., 2019 [Publ 3921], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 5 5 1 8.0
  +----------- Ensidens             
--+                                 
  |  +-------- Hyriopsis_desowitzi **  
  +--+                              
     |  +----- Hyriopsis_myersiana **  
     +--+                           
        |  +-- Hyriopsis_khoratensis **
        +--+                        
           +-- Hyriopsis_panhai **     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
3 7 2 3 77 4 2 10.0
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens            
--+                           
  +----- Ensidens             
  |                           
  |  +-- Hyriopsis_khoratensis **
  +--+                        
     +-- Hyriopsis_myersiana **  

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 8 0 3 80 5 1 13.0
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_jaculus     
  |                        
  +-- Hyriopsis_desowitzi **  
--+                        
  +-- Hyriopsis_myersiana **  
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius 

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 30 2 2 --
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
2 2 0 2 35 2 2 --
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
2 13 0 2 35 2 2 19.0
  +----------------------- Hyriopsis_bialata **  
  |                                           
--+                    +-- Rectidens          
  |  +-----------------+                      
  |  |                 +-- Khairuloconcha     
  +--+                                        
     |  +----------------- Ensidens           
     |  |                                     
     +--+  +-------------- Hyriopsis_desowitzi **
        |  |                                  
        +--+  +----------- Yaukthwa_elongatula
           |  |                               
           +--+  +-------- Trapezoideus       
              |  |                            
              +--+     +-- Physunio           
                 |  +--+                      
                 |  |  +-- Yaukthwa_inlenensis
                 +--+                         
                    |  +-- Yaukthwa_paiensis  
                    +--+                      
                       +-- Lens               

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
2 3 0 2 35 2 2 9.0
  +-------- Ensidens           
--+                            
  |  +----- Hyriopsis_bialata **  
  +--+                         
     |  +-- Hyriopsis_desowitzi **
     +--+                      
        +-- Rectidens          

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 44 3 2 --
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4388], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 36 2 2 --
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
2 2 0 2 35 2 2 --
  +-------- Ensidens           
--+                            
  |  +----- Rectidens          
  +--+                         
     |  +-- Hyriopsis_bialata **  
     +--+                      
        +-- Hyriopsis_desowitzi **

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
2 13 0 2 35 2 2 18.0
  +----- Ensidens           
  |                         
  +----- Hyriopsis_desowitzi **
  |                         
  +----- Rectidens          
  |                         
  +----- Hyriopsis_bialata **  
--+                         
  +----- Khairuloconcha     
  |                         
  |  +-- Physunio           
  |  |                      
  |  +-- Yaukthwa_paiensis  
  |  |                      
  |  +-- Yaukthwa_inlenensis
  +--+                      
     +-- Trapezoideus       
     |                      
     +-- Yaukthwa_elongatula
     |                      
     +-- Lens               

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 --
  +-- Ensidens         
--+                    
  +-- Hyriopsis_bialata **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 --
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens        

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 57 2 2 --
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens        

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Ensidens         

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens        

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 3 1 --
  +-- Hyriopsis_desowitzi **
--+                      
  +-- Ensidens           

Matos et al., 2024 [Publ 4384], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 --
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens        

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iT tT bi riS oT roT roS iD
1 3 0 2 6 2 2 11.0
  +-- Ensidens    
  |               
  +-- Hyriopsis__1 **
--+               
  +-- Hyriopsis__2 **
  |               
  +-- Rectidens   

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +-- Ensidens         
  |                    
--+-- Hyriopsis_bialata **
  |                    
  +-- Rectidens        

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Ensidens         
--+                       
  |  +-- Rectidens        
  +--+                    
     +-- Hyriopsis_bialata **

Matos et al., 2024 [Publ 4383], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Ensidens         
--+                       
  |  +-- Hyriopsis_bialata **
  +--+                    
     +-- Rectidens        

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 3 1 --
  +-- Hyriopsis_desowitzi **
--+                      
  +-- Ensidens           

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Ensidens         

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 --
  +-- Hyriopsis_bialata **
--+                    
  +-- Rectidens        

Page updated: 14:57:07 Wed 04 Dec 2024

 
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