| Page last updated
Fri 04 Oct 2024 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Wu et al. (2024), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Wu, R., L. Zhang, L. Liu, J. Jia & X. Liu. 2024. Unraveling the phylogenetic relationships and taxonomic status of a puzzling freshwater mussel genus Inversidens (Bivalvia, Unionidae) through multilocus phylogeny and mitochondrial phylogenomics. Journal of Zoological Systematics and Evolutionary Research 2024, 1499508: 1-18.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 5a
fig. 5b
fig. 5c
fig. 6-7
fig. 8a_S1
fig. 8b_S2
Attributed Species Names
Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
(misidentified reference to Unio crassus Retzius, 1788)
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Lamellidens marginalis Bolotov et al., 2017
‘Lamarck’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 4’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207. [in part]
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
(misidentified reference to Unio marginalis Lamarck, 1819)
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Taxonomic Opinions
Aculamprotula coreana
Aculamprotula polysticta
Aculamprotula scripta
Aculamprotula tientsinensis
Aculamprotula tortuosa
Acuticosta chinensis
Alasmidonta varicosa
Amblema plicata
Anemina arcaeformis
Anemina euscaphys
Anodonta anatina
Anodonta cygnea
Anodonta exulcerata
Anodonta nuttalliana
Anodontites trapesialis
Beringiana fukuharai
Bineurus anodontinum (Rochebrune, 1882)
Bineurus loeiensis Konopleva et al., 2021
Chamberlainia somsakpanhai Kongim et al., 2023
Cristaria plicata
Cuneopsis celtiformis
Cuneopsis demangei
Cuneopsis heudei
Cuneopsis rufescens
Echyridella menziesii
Elliptio complanata
Ensidens sagittarius (Lea, 1856)
Gonidea angulata (Lea, 1838)
Hyriopsis bialata Simpson, 1900
Hyriopsis desowitzi Brandt, 1974
Indopseudodon bogani (Bolotov et al., 2017)
Indopseudodon kayinensis (Bolotov et al., 2020)
+ Pseudodon crebristriatus Pfeiffer et al., 2019
Inversidens brandtii (Kobelt, 1879)
Inversidens rentianensis Wu & Wu, 2021
Khairuloconcha lunbawangorum Zieritz et al., 2021
Khairuloconcha sahanae Zieritz et al., 2021
Koreosolenaia sitgyensis Lee et al., 2020
Lamellidens marginalis
Lamprotula caveata
Lamprotula cornuumlunae
Lamprotula leaii (Gray, 1833)
Lampsilis cardium
Lampsilis cariosa
Lampsilis ornata
Lampsilis powellii
Lampsilis siliquoidea
Lanceolaria gladiola
Lanceolaria grayii
Lanceolaria lanceolata
Lasmigona complanata
Leaunio lienosus
Lens contradens (Lea, 1838)
Lens eximius (Lea, 1856)
Lepidodesma aligera
Lepidodesma languilati
Margaritifera dahurica (Middendorff, 1850)
Margaritifera margaritifera (Linnaeus, 1758)
Microcondylaea bonellii (Férussac, 1827)
Mutela dubia
Neotrigonia margaritacea
Nodularia breviconcha
Nodularia douglasiae
Parvasolenaia rivularis (Heude, 1877)
Physunio superbus (Lea, 1843)
Pilsbryoconcha exilis (Lea, 1838)
Platynaias compressa
Pleurobema oviforme
Popenaias popeii
Postolata guangxiensis Dai et al., 2023
Potamilus alatus
Potamilus leptodon
Potamilus streckersoni
Potomida littoralis (Cuvier, 1798)
Prolasmidonta heterodon
Pronodularia japanensis
Pseudanodonta complanata
Pseudocuneopsis capitata
Pseudocuneopsis sichuanensis
Pseudodon mekongi (Bolotov et al., 2020)
Pseudodon vondembuschianus (Lea, 1840)
Ptychorhynchus pfisteri (Heude, 1874)
Pyganodon grandis
Quadrula quadrula
Rectidens sumatrensis (Dunker, 1852)
Schistodesmus lampreyanus
Schistodesmus sp.
Schistodesmus spinosus
Sinanodonta lucida
Sinanodonta tumens
Sinanodonta woodiana
Sinohyriopsis cumingii
Sinohyriopsis schlegelii
Sinosolenaia carinata (Heude, 1877)
Sinosolenaia iridinea
Sinosolenaia oleivora
Tchangsinaia piscicula
Thaiconcha callifera (Martens, 1860)
Toxolasma parvum
Trapezoideus foliaceus (Gould, 1843)
Unio crassus
Unio delphinus
Unio elongatulus
Unio mancus
Unio pictorum
Unio tumidus
Uniomerus tetralasmus
Utterbackia imbecillis
Utterbackia peninsularis
Venustaconcha ellipsiformis
Yaukthwa elongatula Bolotov et al., 2019
Yaukthwa inlenensis Konopleva et al., 2019
Yaukthwa paiensis Konopleva et al., 2019
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Cladogram Figure 5a
+-- Margaritifera_dahurica
+--------------------------+
| +-- Margaritifera_margaritifera
|
| +-------------------------- Chamberlainia_somsakpanhai
| |
| | +-- Khairuloconcha_lunbawangorum
| | +--------------+
--+ | | +-- Khairuloconcha_sahanae
| | |
| | | +-------- Ensidens_sagittarius
| | +-----+ +-----+
| | | | | | +----- Rectidens_sumatrensis
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata
| | | | | +--+
+--+ | +--+ +-- Hyriopsis_desowitzi
| | |
| | | +-- Physunio_superbus
| | | +--------+
| | | | +-- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_eximius
| | +--+
| | | +----- Yaukthwa_elongatula
| | +--+
+--+ | +-- Yaukthwa_inlenensis
| +--+
| +-- Yaukthwa_paiensis
|
| +----------------- Schepmania_nieuwenhuisi
| |
| +--+ +-- Indopseudodon_bogani
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis
| | +--+
| | | +----------- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Pseudodon_mekongi
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
| | | +----- Thaiconcha_callifera
+--+ +--+
| | +-- Bineurus_anodontinum
| +--+
| +-- Bineurus_loeiensis
|
| +-- Inversidens_brandtii
| +-----+
| | +-- Inversidens_rentianensis
| +--------+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +-- Gonidea_angulata
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Parvasolenaia_rivularis
+--+
| +----- Koreosolenaia_sitgyensis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 5b
+-- Margaritifera_dahurica
+-----------------------------------+
| +-- Margaritifera_margaritifera
|
| +----------------------------------- Schepmania_nieuwenhuisi
| |
| | +-- Indopseudodon_bogani
--+ | +-----------+
| | | +-- Indopseudodon_kayinensis
| | +-----------------+
| | | | +----------- Pilsbryoconcha_exilis
| | | | |
| | | +--+ +-- Bineurus_anodontinum
| | | | +-----+
+--+ | | | +-- Bineurus_loeiensis
| | +--+
| | | +----- Thaiconcha_callifera
| | +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +-------------------------- Hyriopsis_bialata
| | |
+--+ | +----- Rectidens_sumatrensis
| +--+ +-----------------+
| | | | | +-- Khairuloconcha_lunbawangorum
| | | | +--+
| | +--+ +-- Khairuloconcha_sahanae
| | |
| | | +-------------------- Ensidens_sagittarius
| | | |
| | +--+ +----------------- Hyriopsis_desowitzi
| | | |
| | +--+ +-------------- Yaukthwa_elongatula
| | | |
| | +--+ +----------- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Physunio_superbus
+--+ | +-----+
| | | +-- Yaukthwa_inlenensis
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_eximius
|
| +-- Inversidens_brandtii
| +-----------------+
| | +-- Inversidens_rentianensis
| |
| | +-- Chamberlainia_somsakpanhai
+--------+ +--+
| | +-- Potomida_littoralis
| +-----------+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +-- Gonidea_angulata
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 5c
+-- Margaritifera_dahurica
+-----------------------------+
| +-- Margaritifera_margaritifera
|
| +----------------------------- Schepmania_nieuwenhuisi__1
| |
| +----------------------------- Schepmania_nieuwenhuisi__2
| |
--+ | +-- Indopseudodon_bogani
| | +-----------+
| | | +-- Indopseudodon_kayinensis
| | +-----------+
| | | | +----------- Pilsbryoconcha_exilis
| | | | |
| | | +--+ +-------- Bineurus_anodontinum
+--+ | | |
| | +--+-------- Bineurus_loeiensis
| | |
| | | +----- Thaiconcha_callifera
| | +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +-------- Ensidens_sagittarius
| | |
| | +-------- Hyriopsis_desowitzi
+--+ |
| +-------- Rectidens_sumatrensis
| |
| +-------- Hyriopsis_bialata
| |
| +--------------+ +-- Khairuloconcha_lunbawangorum
| | +-----+
| | | +-- Khairuloconcha_sahanae
| | |
| | | +----- Physunio_superbus
| | | |
| | | +----- Yaukthwa_paiensis
| | | |
| | | +----- Yaukthwa_inlenensis
| | +--+
| | +----- Trapezoideus_foliaceus
+--+ |
| +----- Yaukthwa_elongatula
| |
| | +-- Lens_contradens
| +--+
| +-- Lens_eximius
|
| +-- Inversidens_rentianensis
| +-----------------+
| | +-- Inversidens_brandtii
| |
| | +-- Chamberlainia_somsakpanhai
+--+ +--+
| | +-- Potomida_littoralis
| +-----------+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-------------- Sinosolenaia_carinata
| |
+--+ +----------- Gonidea_angulata
| |
+--+----------- Microcondylaea_bonellii
|
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 6-7
+-- Margaritifera_dahurica
+-----------------------------+
| +-- Margaritifera_margaritifera
|
| +----------------------------- Chamberlainia_somsakpanhai
| |
| | +-- Khairuloconcha_lunbawangorum
| | +--------------+
--+ | | +-- Khairuloconcha_sahanae
| | |
| | | +-------- Ensidens_sagittarius
| | +--------+ +-----+
| | | | | | +----- Rectidens_sumatrensis
| | | | | +--+
| | | | | | +-- Hyriopsis_bialata
+--+ | | | +--+
| | +--+ +-- Hyriopsis_desowitzi
| | |
| | | +-- Physunio_superbus
| | | +--------+
| | | | +-- Trapezoideus_foliaceus
| | | |
| | +--+ +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_eximius
| | +--+
| | | +----- Yaukthwa_elongatula
+--+ +--+
| | +-- Yaukthwa_inlenensis
| +--+
| +-- Yaukthwa_paiensis
|
| +----------------- Schepmania_nieuwenhuisi
| |
| +-----+ +-- Indopseudodon_bogani
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis
| | +--+
| | | +----------- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Pseudodon_mekongi
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus
| | +--+
+--+ | +----- Thaiconcha_callifera
| +--+
| | +-- Bineurus_anodontinum
| +--+
| +-- Bineurus_loeiensis
|
| +----- Potomida_littoralis
| +--------------+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_leaii
+--+
| +-- Inversidens_brandtii
| +--------------+
| | +-- Inversidens_rentianensis
+--+
| +-------------- Sinosolenaia_carinata
| |
+--+ +-- Gonidea_angulata
| +--------+
| | +-- Microcondylaea_bonellii
+--+
| +-------- Koreosolenaia_sitgyensis
+--+
| +----- Parvasolenaia_rivularis
+--+
| +-- Postolata_guangxiensis
+--+
+-- Ptychorhynchus_pfisteri
Cladogram Figure 8a_S1
+-------------------------------------------------- Neotrigonia_margaritacea
|
| +----------------------------------------------- Echyridella_menziesii
| |
--+ | +-- Anodontites_trapesialis
| | +-----------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Margaritifera_dahurica
+--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_nanus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Cladogram Figure 8b_S2
+-------------------------------------------------- Neotrigonia_margaritacea
|
| +----------------------------------------------- Echyridella_menziesii
| |
--+ | +-- Anodontites_trapesialis
| | +-----------------------------------------+
| | | +-- Mutela_dubia
| | |
| | | +-- Margaritifera_dahurica
+--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | +--+ +-- Potamilus_streckersoni
| | |
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-------- Venustaconcha_ellipsiformis
| | | |
| | | | +-- Lampsilis_powellii
| | +--+-----+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_cariosa
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_nanus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
|
Page updated: 09:49:14 Fri 04 Oct 2024 |
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