MUSSELp
 
MUSSELpdb
  Introduction
Browse
d'basing
 
Taxonomy
  all genera
Unionidae
Margaritiferidae
Hyriidae
Etheriidae
Mycetopodidae
Iridinidae

Species & Genus Richness by Taxonomy
 
Geography
  Summary
North America
Central America
South America
Afrotropics
North Eurasia
East Asia
Australasia

Species Richness & Endemism by Geography
 
 

Page last updated
Sun 07 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validgen_cld&id=1405

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Aculamprotula Wu, Liang, Wang & Ouyang, 1999: cladomics
     9 valid species

Cladomics Summary

number of species = 9

analyzed in 103 cladograms in 57 publications
maximum number of terminals in an analysis = 5, mean = 2.62

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 61
maximum number of included terminals = 5, mean = 3.74
proportion of cladograms that support basic monophyly = 1.0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Dai et al., 2025 [Publ 4440], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 40 17 10 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +-------------------- Globunio                
  |                                             
  |           +-------- Aculamprotula_tortuosa **  
  |           |                                 
  |  +--------+     +-- Aculamprotula_fibrosa **   
--+  |        |  +--+                           
  |  |        |  |  +-- Aculamprotula_coreana **   
  |  |        +--+                              
  |  |           |  +-- Aculamprotula_scripta **   
  |  |           +--+                           
  +--+              +-- Aculamprotula_polysticta **
     |                                          
     |              +-- Middendorffinaia        
     |  +-----------+                           
     |  |           +-- Nodularia               
     |  |                                       
     +--+  +----------- Cuneopsis               
        |  |                                    
        |  |        +-- Diaurora                
        +--+--------+                           
           |        +-- Schistodesmus           
           |                                    
           |  +-------- Pseudocuneopsis         
           +--+                                 
              |  +----- Inversiunio             
              +--+                              
                 |  +-- Tchangsinaia            
                 +--+                           
                    +-- Unio

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 5 3 5 194 14 5 4.0 
  +-------- Aculamprotula_tortuosa **     
  |                                    
--+     +-- Aculamprotula_coreana **      
  |  +--+                              
  |  |  +-- Aculamprotula_scripta **      
  +--+                                 
     |  +-- Aculamprotula_fibrosa **      
     +--+                              
        +-- Aculamprotula_tientsinensis **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 44 9 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_coreana **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_tientsinensis **
  |        +--+                                 
--+           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Schistodesmus

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
5 5 3 5 40 10 4 4.0 
        +-------- Nodularia                  
  +-----+                                    
  |     |  +----- Tchangsinaia               
  |     +--+                                 
  |        |  +-- Schistodesmus              
--+        +--+                              
  |           +-- Cuneopsis                  
  |                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_tientsinensis **
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_coreana **      
           +--+                              
              +-- Aculamprotula_scripta **

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 5 35 15 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_polysticta **   
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_scripta **      
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 194 14 5 3.0 
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+  +----- Aculamprotula_coreana **      
  |     |  |                                 
  |     +--+----- Aculamprotula_scripta **      
  |        |                                 
--+        |  +-- Aculamprotula_fibrosa **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Schistodesmus              
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
5 5 3 5 90 17 6 2.0 
              +-------- Aculamprotula_tortuosa **      
              |                                     
  +-----------+     +-- Aculamprotula_coreana **       
  |           |  +--+                               
  |           |  |  +-- Aculamprotula_tientsinensis ** 
  |           +--+                                  
  |              |  +-- Aculamprotula_polysticta **    
--+              +--+                               
  |                 +-- Aculamprotula_scripta **       
  |                                                 
  |  +----------------- Unio                        
  |  |                                              
  +--+  +-------------- Pseudocuneopsis             
     |  |                                           
     +--+  +----------- Nodularia                   
        |  |                                        
        +--+        +-- Schistodesmus_lampreyanus__1
           |  +-----+                               
           |  |     +-- Cuneopsis                   
           +--+                                     
              |  +----- Tchangsinaia                
              +--+                                  
                 |  +-- Schistodesmus_lampreyanus__2
                 +--+                               
                    +-- Schistodesmus_spinosus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
5 5 3 5 116 22 7 2.0 
        +----------- Aculamprotula_tortuosa **     
  +-----+                                       
  |     |  +-------- Aculamprotula_scripta **      
  |     +--+                                    
  |        |  +----- Aculamprotula_coreana **      
  |        +--+                                 
--+           |  +-- Aculamprotula_fibrosa **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Middendorffinaia           
  |  |                                          
  +--+  +----------- Unio                       
     |  |                                       
     +--+  +-------- Nodularia                  
        |  |                                    
        +--+  +----- Schistodesmus              
           |  |                                 
           +--+----- Cuneopsis                  
              |                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Inversiunio

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
5 5 3 5 33 18 7 2.0 
              +-------- Aculamprotula_tortuosa **     
              |                                    
  +-----------+     +-- Aculamprotula_tientsinensis **
  |           |  +--+                              
  |           |  |  +-- Aculamprotula_coreana **      
  |           +--+                                 
--+              |  +-- Aculamprotula_polysticta **   
  |              +--+                              
  |                 +-- Aculamprotula_scripta **      
  |                                                
  |  +----------------- Unio                       
  +--+                                             
     |  +-------------- Pseudocuneopsis            
     +--+                                          
        |  +----------- Nodularia                  
        +--+                                       
           |  +-------- Schistodesmus              
           +--+                                    
              |  +----- Cuneopsis                  
              +--+                                 
                 |  +-- Tchangsinaia               
                 +--+                              
                    +-- Inversiunio

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 38 10 4 2.0 
  +----------- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Aculamprotula_scripta **      
  +--+                                    
     |  +----- Aculamprotula_coreana **      
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 68 15 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_tientsinensis **
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_coreana **      
  |        +--+                                 
--+           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 5 3 5 192 13 5 4.0 
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_scripta **      
  |     +--+                                 
--+        |  +-- Aculamprotula_fibrosa **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iT tT bi riS oT roT roS iD
5 5 3 5 4 2 1 4.0 
  +-------------- Nodularia                  
--+                                          
  |  +----------- Aculamprotula_tortuosa **     
  +--+                                       
     |  +-------- Aculamprotula_tientsinensis **
     +--+                                    
        |  +----- Aculamprotula_fibrosa **      
        +--+                                 
           |  +-- Aculamprotula_scripta **      
           +--+                              
              +-- Aculamprotula_coreana **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
5 5 3 5 68 10 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_scripta **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_polysticta **   
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Tchangsinaia

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 48 9 5 2.0 
     +----------- Aculamprotula_tortuosa **     
  +--+                                       
  |  |  +-------- Aculamprotula_scripta **      
  |  +--+                                    
  |     |  +----- Aculamprotula_coreana **      
  |     +--+                                 
--+        |  +-- Aculamprotula_fibrosa **      
  |        +--+                              
  |           +-- Aculamprotula_tientsinensis **
  |                                          
  |     +-------- Unio                       
  |     |                                    
  +-----+     +-- Nodularia                  
        |  +--+                              
        |  |  +-- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
5 5 3 5 90 17 6 2.0 
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_tientsinensis **
  |     +--+                                 
  |        |  +-- Aculamprotula_polysticta **   
--+        +--+                              
  |           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+        +-- Nodularia                  
     |  +-----+                              
     |  |     +-- Pseudocuneopsis            
     +--+                                    
        |  +----- Tchangsinaia               
        +--+                                 
           |  +-- Schistodesmus              
           +--+                              
              +-- Cuneopsis

Hou et al., 2024 [Publ 4400], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 5 71 44 7 2.0 
              +-------- Aculamprotula_tortuosa **     
              |                                    
  +-----------+     +-- Aculamprotula_coreana **      
  |           |  +--+                              
  |           |  |  +-- Aculamprotula_tientsinensis **
  |           +--+                                 
--+              |  +-- Aculamprotula_polysticta **   
  |              +--+                              
  |                 +-- Aculamprotula_scripta **      
  |                                                
  |  +----------------- Unio                       
  +--+                                             
     |  +-------------- Middendorffinaia           
     +--+                                          
        |  +----------- Pseudocuneopsis            
        +--+                                       
           |  +-------- Nodularia                  
           +--+                                    
              |  +----- Schistodesmus              
              +--+                                 
                 |  +-- Tchangsinaia               
                 +--+                              
                    +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 83 24 8 2.0 
  +----------------- Globunio                   
  |                                             
  |        +-------- Aculamprotula_tortuosa **     
  |        |                                    
--+  +-----+     +-- Aculamprotula_polysticta **   
  |  |     |  +--+                              
  |  |     |  |  +-- Aculamprotula_scripta **      
  |  |     +--+                                 
  |  |        |  +-- Aculamprotula_coreana **      
  +--+        +--+                              
     |           +-- Aculamprotula_tientsinensis **
     |                                          
     |  +----------- Unio                       
     |  |                                       
     +--+        +-- Nodularia                  
        |  +-----+                              
        |  |     +-- Pseudocuneopsis            
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 5 63 10 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_scripta **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_polysticta **   
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
5 5 3 5 48 24 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_coreana **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_tientsinensis **
  |        +--+                                 
--+           |  +-- Aculamprotula_polysticta **   
  |           +--+                              
  |              +-- Aculamprotula_scripta **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
5 5 3 5 86 17 6 2.0 
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_tientsinensis **
  |     +--+                                 
  |        |  +-- Aculamprotula_polysticta **   
--+        +--+                              
  |           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+  +-------- Nodularia                  
     |  |                                    
     +--+-------- Pseudocuneopsis            
        |                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis

Dai et al., 2025 [Publ 4440], fig. 1

iT tT bi riS oT roT roS iD
5 5 2 5 40 17 10 1.0 
  +----------------------- Globunio                
  |                                                
  |              +-------- Aculamprotula_tortuosa **  
  |              |                                 
--+  +-----------+  +----- Aculamprotula_coreana **   
  |  |           |  |                              
  |  |           +--+----- Aculamprotula_fibrosa **   
  |  |              |                              
  |  |              |  +-- Aculamprotula_polysticta **
  +--+              +--+                           
     |                 +-- Aculamprotula_scripta **   
     |                                             
     |  +----------------- Middendorffinaia        
     |  |                                          
     +--+  +-------------- Nodularia               
        |  |                                       
        +--+  +----------- Cuneopsis               
           |  |                                    
           |  |        +-- Diaurora                
           +--+--------+                           
              |        +-- Schistodesmus           
              |                                    
              |  +-------- Pseudocuneopsis         
              +--+                                 
                 |  +----- Unio                    
                 +--+                              
                    |  +-- Tchangsinaia            
                    +--+                           
                       +-- Inversiunio

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
5 5 3 5 86 17 6 2.0 
        +-------- Aculamprotula_tortuosa **     
        |                                    
  +-----+     +-- Aculamprotula_coreana **      
  |     |  +--+                              
  |     |  |  +-- Aculamprotula_tientsinensis **
  |     +--+                                 
  |        |  +-- Aculamprotula_polysticta **   
--+        +--+                              
  |           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+  +-------- Nodularia                  
     |  |                                    
     +--+-------- Pseudocuneopsis            
        |                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
5 5 3 5 68 10 6 2.0 
           +-------- Aculamprotula_tortuosa **     
           |                                    
  +--------+     +-- Aculamprotula_scripta **      
  |        |  +--+                              
  |        |  |  +-- Aculamprotula_polysticta **   
  |        +--+                                 
--+           |  +-- Aculamprotula_coreana **      
  |           +--+                              
  |              +-- Aculamprotula_tientsinensis **
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Schistodesmus              
              +--+                              
                 +-- Cuneopsis

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 42 9 6 3.0 
           +-------- Aculamprotula_tortuosa **     
  +--------+                                    
  |        |  +----- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_tientsinensis **
--+           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Schistodesmus

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 18 2 2 3.0 
           +-- Unio                       
  +--------+                              
  |        +-- Nodularia                  
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_coreana **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 52 7 4 3.0 
     +-------- Aculamprotula_tortuosa **     
  +--+                                    
  |  |  +----- Aculamprotula_scripta **      
  |  +--+                                 
  |     |  +-- Aculamprotula_tientsinensis **
--+     +--+                              
  |        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Schistodesmus

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 25 6 6 3.0 
           +-------- Aculamprotula_tortuosa **     
  +--------+                                    
  |        |  +----- Aculamprotula_scripta **      
  |        +--+                                 
  |           |  +-- Aculamprotula_tientsinensis **
--+           +--+                              
  |              +-- Aculamprotula_coreana **      
  |                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Cuneopsis                  
              +--+                              
                 +-- Tchangsinaia

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 30 1 1 3.0 
  +----------- Unio                       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 30 1 1 3.0 
  +----------- Unio                       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_polysticta **   
     +--+                                 
        |  +-- Aculamprotula_fibrosa **      
        +--+                              
           +-- Aculamprotula_tientsinensis **

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
4 4 2 4 52 8 5 3.0 
  +-------------- Middendorffinaia           
  |                                          
  |     +-------- Aculamprotula_tortuosa **     
--+  +--+                                    
  |  |  |  +----- Aculamprotula_scripta **      
  |  |  +--+                                 
  |  |     |  +-- Aculamprotula_tientsinensis **
  +--+     +--+                              
     |        +-- Aculamprotula_coreana **      
     |                                       
     |  +-------- Unio                       
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Cuneopsis

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 10 1 1 3.0 
  +----------- Unio                       
--+                                       
  |  +-------- Aculamprotula_tortuosa **     
  +--+                                    
     |  +----- Aculamprotula_scripta **      
     +--+                                 
        |  +-- Aculamprotula_tientsinensis **
        +--+                              
           +-- Aculamprotula_fibrosa **

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 41 7 4 4.0 
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 65 10 5 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Inversiunio                
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Schistodesmus              
           +--+                              
              +-- Tchangsinaia

Wu et al., 2024 [Publ 4379], fig. 4_6

iT tT bi riS oT roT roS iD
3 3 1 3 19 11 2 4.0 
  +----- Aculamprotula_tortuosa **     
--+                                 
  |  +-- Aculamprotula_scripta **      
  +--+                              
     +-- Aculamprotula_tientsinensis **

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 18 2 2 4.0 
        +-- Unio                       
  +-----+                              
  |     +-- Nodularia                  
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_coreana **      
     +--+                              
        +-- Aculamprotula_tientsinensis **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 28 2 2 4.0 
        +-- Unio                       
  +-----+                              
  |     +-- Nodularia                  
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_tientsinensis **
     +--+                              
        +-- Aculamprotula_scripta **

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 18 4 3 4.0 
        +-- Tchangsinaia               
  +-----+                              
  |     +-- Nodularia                  
--+                                    
  |  +----- Aculamprotula_tortuosa **     
  +--+                                 
     |  +-- Aculamprotula_tientsinensis **
     +--+                              
        +-- Aculamprotula_scripta **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
3 3 1 3 45 7 4 4.0 
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Schistodesmus              
        +--+                              
           +-- Tchangsinaia

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 41 7 4 4.0 
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Schistodesmus

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 20 7 4 4.0 
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_tientsinensis **
  |     +--+                              
--+        +-- Aculamprotula_coreana **      
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Schistodesmus              
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Nodularia

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 35 7 4 4.0 
        +----- Aculamprotula_tortuosa **     
  +-----+                                 
  |     |  +-- Aculamprotula_coreana **      
  |     +--+                              
--+        +-- Aculamprotula_tientsinensis **
  |                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Schistodesmus              
     +--+                                 
        |  +-- Nodularia                  
        +--+                              
           +-- Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 65 10 5 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Inversiunio                
        +--+                                 
           |  +-- Schistodesmus              
           +--+                              
              +-- Tchangsinaia

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
3 3 1 3 88 8 5 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_scripta **      
  |        +--+                              
--+           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Cuneopsis                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Schistodesmus

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 88 8 5 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_scripta **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Cuneopsis                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Schistodesmus

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 65 10 5 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Inversiunio                
     +--+                                    
        |  +----- Nodularia                  
        +--+                                 
           |  +-- Tchangsinaia               
           +--+                              
              +-- Schistodesmus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 50 9 6 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_tientsinensis **
  |        +--+                              
--+           +-- Aculamprotula_coreana **      
  |                                          
  |  +----------- Unio                       
  |  |                                       
  +--+        +-- Schistodesmus              
     |  +-----+                              
     |  |     +-- Cuneopsis                  
     +--+                                    
        |  +----- Pseudocuneopsis            
        +--+                                 
           |  +-- Nodularia                  
           +--+                              
              +-- Tchangsinaia

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 39 6 3 4.0 
     +----- Aculamprotula_tortuosa **     
  +--+                                 
  |  |  +-- Aculamprotula_coreana **      
  |  +--+                              
--+     +-- Aculamprotula_tientsinensis **
  |                                    
  |  +----- Unio                       
  +--+                                 
     |  +-- Tchangsinaia               
     +--+                              
        +-- Nodularia

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 15 2 2 4.0 
  +----- Aculamprotula_tortuosa **     
--+                                 
  |  +-- Aculamprotula_tientsinensis **
  +--+                              
     +-- Aculamprotula_scripta **

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 37 8 5 4.0 
           +----- Aculamprotula_tortuosa **     
  +--------+                                 
  |        |  +-- Aculamprotula_coreana **      
  |        +--+                              
--+           +-- Aculamprotula_tientsinensis **
  |                                          
  |  +----------- Unio                       
  +--+                                       
     |  +-------- Nodularia                  
     +--+                                    
        |  +----- Tchangsinaia               
        +--+                                 
           |  +-- Cuneopsis                  
           +--+                              
              +-- Schistodesmus

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 14 1 1 -- 
  +-- Aculamprotula_coreana ** 
--+                         
  +-- Aculamprotula_tortuosa **

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 21 11 7 -- 
  +----------------- Unio                       
  |                                             
--+              +-- Aculamprotula_scripta **      
  |  +-----------+                              
  |  |           +-- Aculamprotula_tientsinensis **
  +--+                                          
     |  +----------- Cuneopsis                  
     |  |                                       
     +--+        +-- Nodularia                  
        |  +-----+                              
        |  |     +-- Schistodesmus              
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Inversiunio                
              +--+                              
                 +-- Pseudocuneopsis

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 32 1 1 -- 
  +-- Aculamprotula_coreana ** 
--+                         
  +-- Aculamprotula_tortuosa **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 30 5 4 -- 
           +-- Aculamprotula_tortuosa **
  +--------+                         
  |        +-- Aculamprotula_coreana ** 
--+                                  
  |  +-------- Unio                  
  +--+                               
     |  +----- Schistodesmus         
     +--+                            
        |  +-- Tchangsinaia          
        +--+                         
           +-- Nodularia

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 14 2 2 -- 
     +-- Nodularia                  
  +--+                              
  |  +-- Tchangsinaia               
--+                                 
  |  +-- Aculamprotula_tientsinensis **
  +--+                              
     +-- Aculamprotula_tortuosa **

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 29 5 4 -- 
           +-- Aculamprotula_tortuosa **
  +--------+                         
  |        +-- Aculamprotula_coreana ** 
--+                                  
  |  +-------- Unio                  
  +--+                               
     |  +----- Schistodesmus         
     +--+                            
        |  +-- Nodularia             
        +--+                         
           +-- Tchangsinaia

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 2 36 4 4 -- 
  +-- Aculamprotula_tientsinensis **
--+                              
  +-- Aculamprotula_tortuosa **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 34 4 4 -- 
           +-- Aculamprotula_tientsinensis **
  +--------+                              
  |        +-- Aculamprotula_tortuosa **     
--+                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Schistodesmus              
     +--+                                 
        |  +-- Nodularia                  
        +--+                              
           +-- Tchangsinaia

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 -- 
  +-------- Unio                       
  |                                    
  |     +-- Aculamprotula_tientsinensis **
  +-----+                              
  |     +-- Aculamprotula_scripta **      
--+                                    
  |  +----- Cuneopsis                  
  |  |                                 
  |  |  +-- Nodularia                  
  |  +--+                              
  +--+  +-- Schistodesmus              
     |                                 
     |  +-- Tchangsinaia               
     |  |                              
     +--+-- Inversiunio                
        |                              
        +-- Pseudocuneopsis

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 -- 
  +----------------- Unio                       
  |                                             
--+              +-- Aculamprotula_scripta **      
  |  +-----------+                              
  |  |           +-- Aculamprotula_tientsinensis **
  +--+                                          
     |  +----------- Cuneopsis                  
     |  |                                       
     +--+        +-- Nodularia                  
        |  +-----+                              
        |  |     +-- Schistodesmus              
        +--+                                    
           |  +----- Tchangsinaia               
           +--+                                 
              |  +-- Inversiunio                
              +--+                              
                 +-- Pseudocuneopsis

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 50 15 6 -- 
                 +-- Aculamprotula_scripta **      
  +--------------+                              
  |              +-- Aculamprotula_tientsinensis **
--+                                             
  |  +-------------- Unio                       
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Nodularia                  
        +--+                                    
           |  +----- Schistodesmus              
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Cuneopsis

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 -- 
  +-------------- Aculamprotula_tientsinensis **
  |                                          
--+  +----------- Unio                       
  |  |                                       
  +--+        +-- Pseudocuneopsis            
     |  +-----+                              
     |  |     +-- Nodularia                  
     +--+                                    
        |  +----- Schistodesmus              
        +--+                                 
           |  +-- Cuneopsis                  
           +--+                              
              +-- Tchangsinaia

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 6 -- 
  +----------------- Aculamprotula_tientsinensis **
--+                                             
  |  +-------------- Nodularia                  
  +--+                                          
     |  +----------- Pseudocuneopsis            
     +--+                                       
        |  +-------- Schistodesmus              
        +--+                                    
           |  +----- Cuneopsis                  
           +--+                                 
              |  +-- Tchangsinaia               
              +--+                              
                 +-- Inversiunio

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 4 2 -- 
  +----- Aculamprotula_tortuosa **
--+                            
  |  +-- Schistodesmus         
  +--+                         
     +-- Unio

Huang et al., 2018 [Publ 3828], fig. 3_5

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 -- 
  +-- Unio                       
--+                              
  +-- Aculamprotula_tientsinensis **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 -- 
-- Aculamprotula_scripta

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 10 4 -- 
-- Aculamprotula_tientsinensis

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 16 9 -- 
  +----------------- Aculamprotula_fibrosa **
  |                                       
  |              +-- Middendorffinaia     
--+  +-----------+                        
  |  |           +-- Nodularia            
  |  |                                    
  +--+  +----------- Cuneopsis            
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 2 2 -- 
  +-- Schistodesmus         
--+                         
  +-- Aculamprotula_tortuosa **

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 11 6 -- 
-- Aculamprotula_tientsinensis

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 9 5 -- 
  +----------- Unio                  
  |                                  
--+  +-------- Aculamprotula_tortuosa **
  |  |                               
  |  +-------- Schistodesmus         
  +--+                               
     +-------- Nodularia             
     |                               
     |  +----- Cuneopsis_szechenyii  
     +--+                            
        |  +-- Tchangsinaia          
        +--+                         
           +-- Cuneopsis_rufescens

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 -- 
-- Aculamprotula_tortuosa

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 6 4 -- 
-- Aculamprotula_tortuosa

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 -- 
  +----------------- Aculamprotula_scripta **
--+                                       
  |  +-------------- Unio                 
  +--+                                    
     |  +----------- Middendorffinaia     
     +--+                                 
        |  +-------- Nodularia            
        +--+                              
           |  +----- Cuneopsis            
           +--+                           
              |  +-- Inversiunio          
              +--+                        
                 +-- Schistodesmus

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 1 -- 
-- Aculamprotula_tientsinensis

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 8 6 -- 
  +----------------- Aculamprotula_tortuosa **
--+                                        
  |  +-------------- Unio                  
  +--+                                     
     |  +----------- Pseudocuneopsis       
     +--+                                  
        |  +-------- Nodularia             
        +--+                               
           |  +----- Tchangsinaia          
           +--+                            
              |  +-- Cuneopsis             
              +--+                         
                 +-- Schistodesmus

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 6 3 -- 
  +-------- Aculamprotula_tientsinensis **
--+                                    
  |  +----- Unio                       
  +--+                                 
     |  +-- Tchangsinaia               
     +--+                              
        +-- Nodularia

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 -- 
  +----- Aculamprotula_tortuosa **
--+                            
  |  +-- Schistodesmus         
  +--+                         
     +-- Unio

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 9 5 -- 
  +-------------- Unio                  
--+                                     
  |  +----------- Aculamprotula_tortuosa **
  +--+                                  
     |  +-------- Nodularia             
     +--+                               
        |  +----- Cuneopsis             
        +--+                            
           |  +-- Tchangsinaia          
           +--+                         
              +-- Schistodesmus

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 10 5 -- 
  +-------------- Unio                  
--+                                     
  |  +----------- Aculamprotula_tortuosa **
  +--+                                  
     |  +-------- Nodularia             
     +--+                               
        |  +----- Schistodesmus         
        +--+                            
           |  +-- Tchangsinaia          
           +--+                         
              +-- Cuneopsis

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 -- 
-- Aculamprotula_tortuosa

Dai et al., 2024 [Publ 4386], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 9 5 -- 
-- Aculamprotula_scripta

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 4 2 -- 
  +----- Schistodesmus         
--+                            
  |  +-- Aculamprotula_tortuosa **
  +--+                         
     +-- Unio

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 16 3 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 7 4 -- 
  +----------- Aculamprotula_tientsinensis **
--+                                       
  |  +-------- Unio                       
  +--+                                    
     |  +----- Nodularia                  
     +--+                                 
        |  +-- Tchangsinaia               
        +--+                              
           +-- Cuneopsis

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 -- 
  +-- Unio                  
--+                         
  +-- Aculamprotula_tortuosa **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 5 -- 
  +----------------- Unio                  
--+                                        
  |  +-------------- Aculamprotula_tortuosa **
  +--+                                     
     |  +----------- Nodularia             
     +--+                                  
        |  +-------- Schistodesmus         
        +--+                               
           |  +----- Cuneopsis_szechenyii  
           +--+                            
              |  +-- Tchangsinaia          
              +--+                         
                 +-- Cuneopsis_rufescens

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 4 2 -- 
  +----- Schistodesmus         
--+                            
  |  +-- Aculamprotula_tortuosa **
  +--+                         
     +-- Unio

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 3 3 -- 
  +-------- Aculamprotula_tortuosa **
--+                               
  |  +----- Schistodesmus         
  +--+                            
     |  +-- Nodularia             
     +--+                         
        +-- Unio

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 -- 
  +-- Unio                  
--+                         
  +-- Aculamprotula_tortuosa **

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 16 9 -- 
  +----------------- Aculamprotula_fibrosa **
  |                                       
  |              +-- Middendorffinaia     
--+  +-----------+                        
  |  |           +-- Nodularia            
  |  |                                    
  +--+  +----------- Cuneopsis            
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 124 9 5 -- 
  +-- Unio_crassus          
  |                         
  +-- Unio_pictorum         
  |                         
  +-- Unio_tumidus          
  |                         
  +-- Aculamprotula_tortuosa **
  |                         
  +-- Cuneopsis_szechenyii  
--+                         
  +-- Tchangsinaia          
  |                         
  +-- Cuneopsis_rufescens   
  |                         
  +-- Nodularia_douglasiae  
  |                         
  +-- Nodularia_nuxpersicae 
  |                         
  +-- Schistodesmus

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 9 4 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 2 1 -- 
-- Aculamprotula_tientsinensis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 5 -- 
  +----------------- Unio                  
--+                                        
  |  +-------------- Aculamprotula_tortuosa **
  +--+                                     
     |  +----------- Nodularia             
     +--+                                  
        |  +-------- Schistodesmus         
        +--+                               
           |  +----- Cuneopsis_szechenyii  
           +--+                            
              |  +-- Tchangsinaia          
              +--+                         
                 +-- Cuneopsis_rufescens

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 2 -- 
-- Aculamprotula_tortuosa

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 19 4 1 -- 
  +-- Aculamprotula_tortuosa **
--+                         
  +-- Unio

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 -- 
-- Aculamprotula_tortuosa

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 1 -- 
-- Aculamprotula_tientsinensis

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
1 1 0 1 16 11 6 -- 
  +----------------- Aculamprotula_tortuosa **
--+                                        
  |  +-------------- Unio                  
  +--+                                     
     |  +----------- Pseudocuneopsis       
     +--+                                  
        |  +-------- Nodularia             
        +--+                               
           |  +----- Tchangsinaia          
           +--+                            
              |  +-- Cuneopsis             
              +--+                         
                 +-- Schistodesmus

Page updated: 08:18:20 Sun 07 Sep 2025
 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."