Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini genus Aculamprotula Wu, Liang, Wang & Ouyang, 1999: cladomics 9 valid species
Cladomics Summary
number of species = 9
analyzed in 97 cladograms in 54 publications
maximum number of terminals in an analysis = 5, mean = 2.62
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 57
maximum number of included terminals = 5, mean = 3.75
proportion of cladograms that support basic monophyly = 1.0
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
115 |
21 |
7 |
0.0 |
*** Top Monophyly Cladogram ***
+----------- Aculamprotula_tortuosa **
+-----+
| | +-------- Aculamprotula_scripta **
| +--+
| | +----- Aculamprotula_coreana **
| +--+
--+ | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------------- Middendorffinaia
| |
+--+ +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+ +----- Schistodesmus
| |
+--+----- Cuneopsis
|
| +-- Tchangsinaia
+--+
+-- Inversiunio
Chen et al., 2023 [Publ 4302], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
16 |
9 |
-- |
*** Top Sister-Group Cladogram ***
+----------------- Aculamprotula_fibrosa **
|
| +-- Middendorffinaia
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unio
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
91 |
17 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+-----+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| +--+
| | +-- Aculamprotula_polysticta **
--+ +--+
| +-- Aculamprotula_scripta **
|
| +----------- Unio
| |
+--+ +-- Nodularia
| +-----+
| | +-- Pseudocuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
64 |
10 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_polysticta **
| +--+
--+ | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
2 |
5 |
193 |
13 |
5 |
1.0 |
+-------- Aculamprotula_tortuosa **
|
+-----+ +----- Aculamprotula_coreana **
| | |
| +--+----- Aculamprotula_scripta **
| |
--+ | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Schistodesmus
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
191 |
12 |
5 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+-----+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_scripta **
| +--+
--+ | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Huang et al., 2019 [Publ 3843], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
40 |
9 |
4 |
4.0 |
+-------- Nodularia
+-----+
| | +----- Tchangsinaia
| +--+
| | +-- Schistodesmus
--+ +--+
| +-- Cuneopsis
|
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_tientsinensis **
+--+
| +----- Aculamprotula_fibrosa **
+--+
| +-- Aculamprotula_coreana **
+--+
+-- Aculamprotula_scripta **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
91 |
17 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+-----------+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| +--+
| | +-- Aculamprotula_polysticta **
--+ +--+
| +-- Aculamprotula_scripta **
|
| +----------------- Unio
| |
+--+ +-------------- Pseudocuneopsis
| |
+--+ +----------- Nodularia
| |
+--+ +-- Schistodesmus_lampreyanus__1
| +-----+
| | +-- Cuneopsis
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus_lampreyanus__2
+--+
+-- Schistodesmus_spinosus
Dai et al., 2024 [Publ 4386], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
35 |
15 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_polysticta **
| | +--+
| | | +-- Aculamprotula_scripta **
| +--+
--+ | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
2 |
5 |
29 |
2 |
2 |
2.0 |
+-- Unio
+--------+
| +-- Nodularia
--+
| +-------- Aculamprotula_tortuosa **
| |
+--+ +----- Aculamprotula_scripta **
| |
+--+ +-- Aculamprotula_tientsinensis **
| |
+--+-- Aculamprotula_fibrosa **
|
+-- Aculamprotula_zonata **
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
193 |
13 |
5 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
--+ +-- Aculamprotula_coreana **
| +--+
| | +-- Aculamprotula_scripta **
+--+
| +-- Aculamprotula_fibrosa **
+--+
+-- Aculamprotula_tientsinensis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
4 |
2 |
1 |
4.0 |
+-------------- Nodularia
--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_tientsinensis **
+--+
| +----- Aculamprotula_fibrosa **
+--+
| +-- Aculamprotula_scripta **
+--+
+-- Aculamprotula_coreana **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
87 |
17 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+-----+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| +--+
| | +-- Aculamprotula_polysticta **
--+ +--+
| +-- Aculamprotula_scripta **
|
| +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+-------- Pseudocuneopsis
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2024 [Publ 4350], fig. 8a_S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
87 |
17 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+-----+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| +--+
| | +-- Aculamprotula_polysticta **
--+ +--+
| +-- Aculamprotula_scripta **
|
| +----------- Unio
| |
+--+ +-------- Nodularia
| |
+--+-------- Pseudocuneopsis
|
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
69 |
10 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_polysticta **
| +--+
--+ | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Wu et al., 2022 [Publ 4238], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
69 |
15 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_coreana **
| +--+
--+ | +-- Aculamprotula_polysticta **
| +--+
| +-- Aculamprotula_scripta **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2020 [Publ 4121], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
45 |
9 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| +--+
--+ | +-- Aculamprotula_polysticta **
| +--+
| +-- Aculamprotula_scripta **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2018 [Publ 3836], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
48 |
9 |
5 |
0.0 |
+----------- Aculamprotula_tortuosa **
+--+
| | +-------- Aculamprotula_scripta **
| +--+
| | +----- Aculamprotula_coreana **
| +--+
--+ | +-- Aculamprotula_fibrosa **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------- Unio
| |
+-----+ +-- Nodularia
| +--+
| | +-- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2021 [Publ 4366], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
69 |
10 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_polysticta **
| +--+
--+ | +-- Aculamprotula_coreana **
| +--+
| +-- Aculamprotula_tientsinensis **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Schistodesmus
+--+
+-- Cuneopsis
Wu et al., 2022 [Publ 4238], fig. 2d
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
34 |
18 |
7 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+-----------+ +-- Aculamprotula_tientsinensis **
| | +--+
| | | +-- Aculamprotula_coreana **
| +--+
--+ | +-- Aculamprotula_polysticta **
| +--+
| +-- Aculamprotula_scripta **
|
| +----------------- Unio
+--+
| +-------------- Pseudocuneopsis
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Inversiunio
Huang et al., 2019 [Publ 3843], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
37 |
9 |
4 |
0.0 |
+----------- Aculamprotula_tortuosa **
--+
| +-------- Aculamprotula_scripta **
+--+
| +----- Aculamprotula_coreana **
+--+
| +-- Aculamprotula_fibrosa **
+--+
+-- Aculamprotula_tientsinensis **
Wu et al., 2024 [Publ 4379], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
49 |
24 |
6 |
2.0 |
+-------- Aculamprotula_tortuosa **
|
+--------+ +-- Aculamprotula_coreana **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
| +--+
--+ | +-- Aculamprotula_polysticta **
| +--+
| +-- Aculamprotula_scripta **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Zhou et al., 2007 [Publ 2658], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
18 |
4 |
3 |
2.0 |
+-- Tchangsinaia
+-----------+
| +-- Nodularia
--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Aculamprotula_scripta **
+--+
| +----- Aculamprotula_tientsinensis **
+--+
| +-- Aculamprotula_zonata **
+--+
+-- Aculamprotula_fibrosa **
Zhou et al., 2007 [Publ 2658], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
16 |
2 |
2 |
2.0 |
+----------- Aculamprotula_tortuosa **
--+
| +-------- Aculamprotula_scripta **
+--+
| +----- Aculamprotula_fibrosa **
+--+
| +-- Aculamprotula_zonata **
+--+
+-- Aculamprotula_tientsinensis **
Wu et al., 2020 [Publ 4147], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
10 |
1 |
1 |
1.0 |
+----------- Unio
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_scripta **
+--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_fibrosa **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
52 |
8 |
5 |
1.0 |
+-------------- Middendorffinaia
|
| +-------- Aculamprotula_tortuosa **
--+ +--+
| | | +----- Aculamprotula_scripta **
| | +--+
| | | +-- Aculamprotula_tientsinensis **
+--+ +--+
| +-- Aculamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2018 [Publ 4143], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
18 |
2 |
2 |
1.0 |
+-- Unio
+--------+
| +-- Nodularia
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_scripta **
+--+
| +-- Aculamprotula_coreana **
+--+
+-- Aculamprotula_tientsinensis **
Wu et al., 2019 [Publ 3864], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
43 |
9 |
6 |
1.0 |
+-------- Aculamprotula_tortuosa **
+--------+
| | +----- Aculamprotula_scripta **
| +--+
| | +-- Aculamprotula_tientsinensis **
--+ +--+
| +-- Aculamprotula_coreana **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4121], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
30 |
1 |
1 |
3.0 |
+----------- Unio
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_polysticta **
+--+
| +-- Aculamprotula_fibrosa **
+--+
+-- Aculamprotula_tientsinensis **
Wu et al., 2020 [Publ 4121], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
30 |
1 |
1 |
3.0 |
+----------- Unio
--+
| +-------- Aculamprotula_tortuosa **
+--+
| +----- Aculamprotula_polysticta **
+--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_fibrosa **
Huang et al., 2019 [Publ 3843], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
53 |
7 |
4 |
1.0 |
+-------- Aculamprotula_tortuosa **
+--+
| | +----- Aculamprotula_scripta **
| +--+
| | +-- Aculamprotula_tientsinensis **
--+ +--+
| +-- Aculamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2020 [Publ 4147], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
25 |
6 |
6 |
1.0 |
+-------- Aculamprotula_tortuosa **
+--------+
| | +----- Aculamprotula_scripta **
| +--+
| | +-- Aculamprotula_tientsinensis **
--+ +--+
| +-- Aculamprotula_coreana **
|
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Wu et al., 2021 [Publ 4366], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
88 |
8 |
5 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_scripta **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Huang et al., 2018 [Publ 3828], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
40 |
6 |
3 |
2.0 |
+----- Aculamprotula_tortuosa **
+--+
| | +-- Aculamprotula_coreana **
| +--+
--+ +-- Aculamprotula_tientsinensis **
|
| +----- Unio
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Wu et al., 2024 [Publ 4379], fig. 4_6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
19 |
11 |
2 |
2.0 |
+----- Aculamprotula_tortuosa **
--+
| +-- Aculamprotula_scripta **
+--+
+-- Aculamprotula_tientsinensis **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
46 |
7 |
4 |
2.0 |
+----- Aculamprotula_tortuosa **
+-----+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
21 |
7 |
4 |
2.0 |
+----- Aculamprotula_tortuosa **
+-----+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Cha et al., 2018 [Publ 3916], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
36 |
7 |
4 |
2.0 |
+----- Aculamprotula_tortuosa **
+-----+
| | +-- Aculamprotula_coreana **
| +--+
--+ +-- Aculamprotula_tientsinensis **
|
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Fukata & Iigo, 2020 [Publ 4106], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
51 |
9 |
6 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +----------- Unio
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Cuneopsis
+--+
| +----- Pseudocuneopsis
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Froufe et al., 2019 [Publ 3936], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
42 |
7 |
4 |
2.0 |
+----- Aculamprotula_tortuosa **
+-----+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Aunins et al., 2018 [Publ 3863], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
38 |
8 |
5 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_coreana **
| +--+
--+ +-- Aculamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
66 |
10 |
5 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +----------- Unio
+--+
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Wu et al., 2021 [Publ 4366], fig. 2_I
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
88 |
8 |
5 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_scripta **
| +--+
--+ +-- Aculamprotula_tientsinensis **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Wu et al., 2016 [Publ 3711], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
18 |
2 |
2 |
2.0 |
+-- Unio
+-----+
| +-- Nodularia
--+
| +----- Aculamprotula_tortuosa **
+--+
| +-- Aculamprotula_coreana **
+--+
+-- Aculamprotula_tientsinensis **
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
66 |
10 |
5 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +----------- Unio
+--+
| +-------- Inversiunio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Froufe et al., 2019 [Publ 3936], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
42 |
7 |
4 |
2.0 |
+----- Aculamprotula_tortuosa **
+-----+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
66 |
10 |
5 |
2.0 |
+----- Aculamprotula_tortuosa **
+--------+
| | +-- Aculamprotula_tientsinensis **
| +--+
--+ +-- Aculamprotula_coreana **
|
| +----------- Unio
+--+
| +-------- Nodularia
+--+
| +----- Inversiunio
+--+
| +-- Schistodesmus
+--+
+-- Tchangsinaia
Fonseca et al., 2016 [Publ 3775], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
30 |
5 |
4 |
-- |
+-- Aculamprotula_tortuosa **
+--------+
| +-- Aculamprotula_coreana **
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
31 |
5 |
4 |
-- |
+-- Aculamprotula_tortuosa **
+--------+
| +-- Aculamprotula_coreana **
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Dai et al., 2024 [Publ 4358], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
50 |
15 |
6 |
-- |
+-- Aculamprotula_scripta **
+--------------+
| +-- Aculamprotula_tientsinensis **
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Wu et al., 2023 [Publ 4296], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
21 |
11 |
7 |
-- |
+----------------- Unio
|
--+ +-- Aculamprotula_scripta **
| +-----------+
| | +-- Aculamprotula_tientsinensis **
+--+
| +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Wen et al., 2017 [Publ 3795], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
14 |
1 |
1 |
-- |
+-- Aculamprotula_coreana **
--+
+-- Aculamprotula_tortuosa **
Zhou et al., 2017 [Publ 3788], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
15 |
2 |
2 |
-- |
+-- Nodularia
+--+
| +-- Tchangsinaia
--+
| +-- Aculamprotula_tientsinensis **
+--+
+-- Aculamprotula_tortuosa **
Liu et al., 2023 [Publ 4310], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
22 |
12 |
7 |
-- |
+-------- Unio
|
| +-- Aculamprotula_tientsinensis **
+-----+
| +-- Aculamprotula_scripta **
--+
| +----- Cuneopsis
| |
| | +-- Nodularia
| +--+
+--+ +-- Schistodesmus
|
| +-- Tchangsinaia
| |
+--+-- Inversiunio
|
+-- Pseudocuneopsis
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
37 |
4 |
4 |
-- |
+-- Aculamprotula_tientsinensis **
--+
+-- Aculamprotula_tortuosa **
Chen et al., 2020 [Publ 4170], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
32 |
1 |
1 |
-- |
+-- Aculamprotula_coreana **
--+
+-- Aculamprotula_tortuosa **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
34 |
4 |
4 |
-- |
+-- Aculamprotula_tientsinensis **
+--------+
| +-- Aculamprotula_tortuosa **
--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Liu et al., 2023 [Publ 4310], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
22 |
12 |
7 |
-- |
+----------------- Unio
|
--+ +-- Aculamprotula_scripta **
| +-----------+
| | +-- Aculamprotula_tientsinensis **
+--+
| +----------- Cuneopsis
| |
+--+ +-- Nodularia
| +-----+
| | +-- Schistodesmus
+--+
| +----- Tchangsinaia
+--+
| +-- Inversiunio
+--+
+-- Pseudocuneopsis
Huang et al., 2019 [Publ 3843], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
22 |
4 |
2 |
-- |
+----- Schistodesmus
--+
| +-- Aculamprotula_tortuosa **
+--+
+-- Unio
Sonowal et al., 2021 [Publ 4221], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
2 |
1 |
-- |
-- Aculamprotula_tientsinensis
Dai et al., 2024 [Publ 4386], fig. 5-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
10 |
6 |
-- |
-- Aculamprotula_scripta
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
79 |
10 |
5 |
-- |
+-------------- Unio
--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Sonowal et al., 2021 [Publ 4221], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
-- Aculamprotula_tientsinensis
Araujo et al., 2017 [Publ 3801], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
2 |
2 |
-- |
+-- Schistodesmus
--+
+-- Aculamprotula_tortuosa **
Xue et al., 2016 [Publ 3659], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
3 |
3 |
-- |
+-------- Aculamprotula_tortuosa **
--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Unio
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
4 |
1 |
-- |
+-- Aculamprotula_tortuosa **
--+
+-- Unio
Guerra et al., 2017 [Publ 3772], fig. 2b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
1 |
1 |
-- |
-- Aculamprotula_tortuosa
Wang et al., 2018 [Publ 3865], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
31 |
6 |
3 |
-- |
+-------- Aculamprotula_tientsinensis **
--+
| +----- Unio
+--+
| +-- Tchangsinaia
+--+
+-- Nodularia
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
2 |
1 |
-- |
+-- Aculamprotula_tortuosa **
--+
+-- Unio
Wu et al., 2022 [Publ 4238], fig. 2e
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
11 |
6 |
-- |
+----------------- Aculamprotula_tortuosa **
--+
| +-------------- Unio
+--+
| +----------- Pseudocuneopsis
+--+
| +-------- Nodularia
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis
+--+
+-- Schistodesmus
Guerra et al., 2019 [Publ 4168], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
4 |
2 |
-- |
+----- Aculamprotula_tortuosa **
--+
| +-- Schistodesmus
+--+
+-- Unio
Rosenberg et al., 1997 [Publ 2720], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
1 |
1 |
-- |
-- Aculamprotula_tortuosa
Ouyang et al., 2011 [Publ 3408], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
2 |
2 |
-- |
-- Aculamprotula_tortuosa
Guerra et al., 2017 [Publ 3772], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
1 |
1 |
-- |
+-- Aculamprotula_tortuosa **
--+
+-- Unio
Wang et al., 2018 [Publ 3846], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
31 |
7 |
4 |
-- |
+----------- Aculamprotula_tientsinensis **
--+
| +-------- Unio
+--+
| +----- Nodularia
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis
Sonowal et al., 2021 [Publ 4221], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
1 |
-- |
-- Aculamprotula_tientsinensis
Burzynski et al., 2017 [Publ 3774], fig. 1-top
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
2 |
1 |
-- |
+-- Aculamprotula_tortuosa **
--+
+-- Unio
Guerra et al., 2019 [Publ 4168], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
4 |
2 |
-- |
+----- Schistodesmus
--+
| +-- Aculamprotula_tortuosa **
+--+
+-- Unio
Kim et al., 2021 [Publ 4197], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
10 |
4 |
-- |
-- Aculamprotula_tientsinensis
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
4 |
1 |
-- |
+-- Aculamprotula_tortuosa **
--+
+-- Unio
Chen et al., 2023 [Publ 4302], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
16 |
9 |
-- |
+----------------- Aculamprotula_fibrosa **
|
| +-- Middendorffinaia
--+ +-----------+
| | +-- Nodularia
| |
+--+ +----------- Cuneopsis
| |
| | +-- Diaurora
+--+--------+
| +-- Schistodesmus
|
| +-------- Pseudocuneopsis
+--+
| +----- Inversiunio
+--+
| +-- Tchangsinaia
+--+
+-- Unio
Bolotov et al., 2017 [Publ 3761], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
138 |
9 |
5 |
-- |
+-------------- Unio
--+
| +----------- Aculamprotula_tortuosa **
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Tchangsinaia
+--+
+-- Schistodesmus
Fukata & Iigo, 2019 [Publ 4108], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
11 |
6 |
-- |
-- Aculamprotula_tientsinensis
Dai et al., 2023 [Publ 4285], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
52 |
6 |
6 |
-- |
+-------------- Aculamprotula_tientsinensis **
|
--+ +----------- Unio
| |
+--+ +-- Pseudocuneopsis
| +-----+
| | +-- Nodularia
+--+
| +----- Schistodesmus
+--+
| +-- Cuneopsis
+--+
+-- Tchangsinaia
Huang et al., 2018 [Publ 3828], fig. 6-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
3 |
1 |
-- |
+-- Aculamprotula_tortuosa **
--+
+-- Unio
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
75 |
9 |
5 |
-- |
+----------------- Unio
--+
| +-------------- Aculamprotula_tortuosa **
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_heudei
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Unio
--+
+-- Aculamprotula_tortuosa **
Huang et al., 2018 [Publ 3828], fig. 3_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
28 |
1 |
1 |
-- |
+-- Unio
--+
+-- Aculamprotula_tientsinensis **
Dai et al., 2024 [Publ 4386], fig. 5-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
10 |
6 |
-- |
+----------------- Aculamprotula_scripta **
--+
| +-------------- Unio
+--+
| +----------- Middendorffinaia
+--+
| +-------- Nodularia
+--+
| +----- Cuneopsis
+--+
| +-- Inversiunio
+--+
+-- Schistodesmus
Bolotov et al., 2017 [Publ 3761], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
138 |
9 |
5 |
-- |
+----------- Unio
|
--+ +-------- Aculamprotula_tortuosa **
| |
| +-------- Schistodesmus
+--+
+-------- Nodularia
|
| +----- Cuneopsis_heudei
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Bolotov et al., 2017 [Publ 3764], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
124 |
9 |
5 |
-- |
+-- Unio_crassus
|
+-- Unio_pictorum
|
+-- Unio_tumidus
|
+-- Aculamprotula_tortuosa **
|
+-- Cuneopsis_heudei
--+
+-- Tchangsinaia
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae
|
+-- Nodularia_nuxpersicae
|
+-- Schistodesmus
Sano et al., 2020 [Publ 4138], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
6 |
4 |
-- |
-- Aculamprotula_tortuosa
Dai et al., 2024 [Publ 4386], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
9 |
5 |
-- |
-- Aculamprotula_scripta
Guerra et al., 2019 [Publ 4168], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
40 |
4 |
2 |
-- |
+----- Aculamprotula_tortuosa **
--+
| +-- Schistodesmus
+--+
+-- Unio
Guerra et al., 2019 [Publ 4168], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
40 |
4 |
2 |
-- |
-- Aculamprotula_tortuosa
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
75 |
9 |
5 |
-- |
+----------------- Unio
--+
| +-------------- Aculamprotula_tortuosa **
+--+
| +----------- Nodularia
+--+
| +-------- Schistodesmus
+--+
| +----- Cuneopsis_heudei
+--+
| +-- Tchangsinaia
+--+
+-- Cuneopsis_rufescens
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Unio
--+
+-- Aculamprotula_tortuosa **
|