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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

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Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Anemina Haas, 1969: cladomics
     4 valid species

Cladomics Summary

number of species = 4

analyzed in 72 cladograms in 49 publications
maximum number of terminals in an analysis = 2, mean = 1.49

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 35
maximum number of included terminals = 2, mean = 2.0
proportion of cladograms that support basic monophyly = 1.0


Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 10 43 7 2 --

*** Top Monophyly Cladogram ***

  +-------------- Anemina_arcaeformis__2 **
  |                                     
--+  +----------- Anemina_arcaeformis__5 **
  |  |                                  
  |  |  +-------- Anemina_arcaeformis__3 **
  +--+  |                               
     |  +-------- Anemina_arcaeformis__7 **
     |  |                               
     +--+  +----- Anemina_arcaeformis__4 **
        |  |                            
        |  +----- Anemina_arcaeformis__6 **
        |  |                            
        +--+----- Anemina_arcaeformis__8 **
           |                            
           +----- Anemina_arcaeformis__9 **
           |                            
           |  +-- Anemina_arcaeformis__1 **
           +--+                         
              +-- Anemina_euscaphys **     

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 30 26 6 --

*** Top Sister-Group Cladogram ***

  +----------- Cristaria          
  |                               
  |     +----- Amuranodonta       
--+  +--+                         
  |  |  |  +-- Anemina_arcaeformis **
  |  |  +--+                      
  +--+     +-- Buldowskia         
     |                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Beringiana         
        +--+                      
           +-- Sinanodonta        

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 5 3 --
        +-- Beringiana         
  +-----+                      
  |     +-- Sinanodonta        
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 22 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 30 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 37 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 67 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 10 8 3 --
  +----------- Sinanodonta        
--+                               
  |  +-------- Amuranodonta       
  +--+                            
     |  +----- Buldowskia         
     +--+                         
        |  +-- Anemina_arcaeformis **
        +--+                      
           +-- Anemina_euscaphys **  

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 37 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 48 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 5 3 --
  +-------- Cristaria          
  |                            
--+     +-- Anemina_arcaeformis **
  |  +--+                      
  |  |  +-- Anemina_euscaphys **  
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 43 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 52 5 3 --
  +-------- Cristaria          
  |                            
--+     +-- Anemina_arcaeformis **
  |  +--+                      
  |  |  +-- Anemina_euscaphys **  
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 30 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 55 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 39 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 2 37 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 67 10 5 --
  +----- Buldowskia         
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Anemina_euscaphys **  

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 15 0 0 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Anemina_euscaphys **  

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 30 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 67 10 5 --
  +----- Buldowskia         
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Anemina_euscaphys **  

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 5 3 --
  +-------- Cristaria          
  |                            
--+     +-- Anemina_arcaeformis **
  |  +--+                      
  |  |  +-- Anemina_euscaphys **  
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 43 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Riccardi et al., 2019 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 18 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 41 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 2 0 2 94 5 3 --
  +-------- Cristaria          
  |                            
--+     +-- Anemina_arcaeformis **
  |  +--+                      
  |  |  +-- Anemina_euscaphys **  
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 31 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 2 72 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
2 2 0 2 47 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 45 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 2 72 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_euscaphys **  
     +--+                      
        +-- Anemina_arcaeformis **

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 67 10 5 --
  +----- Buldowskia         
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Anemina_euscaphys **  

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 14 3 2 --
  +----- Sinanodonta        
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Anemina_euscaphys **  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 52 3 2 --
  +-------- Cristaria          
--+                            
  |  +----- Sinanodonta        
  +--+                         
     |  +-- Anemina_arcaeformis **
     +--+                      
        +-- Anemina_euscaphys **  

Feng et al., 2016 [Publ 3800], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 13 2 1 --
  +----- Sinanodonta        
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Anemina_euscaphys **  

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 2 2 --
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta        

Bespalaya et al., 2018 [Publ 3811], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 9 7 1 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta        

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 3 2 --
  +----- Anemina_arcaeformis **
--+                         
  |  +-- Cristaria          
  +--+                      
     +-- Sinanodonta        

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 1 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta        

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 --
  +----- Cristaria        
--+                       
  |  +-- Sinanodonta      
  +--+                    
     +-- Anemina_euscaphys **

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 2 --
-- Anemina_arcaeformis

Dai et al., 2024 [Publ 4385], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 30 26 6 --
        +----- Amuranodonta       
  +-----+                         
  |     |  +-- Anemina_arcaeformis **
  |     +--+                      
--+        +-- Buldowskia         
  |                               
  |  +-------- Cristaria          
  +--+                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Beringiana         
        +--+                      
           +-- Sinanodonta        

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 8 3 --
-- Anemina_arcaeformis

Bauer et al., 2021 [Publ 4365], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 7 7 1 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta        

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 41 7 2 --
-- Anemina_arcaeformis

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 0 0 --
-- Anemina_arcaeformis

Dai et al., 2024 [Publ 4385], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 30 26 6 --
        +----- Amuranodonta       
  +-----+                         
  |     |  +-- Anemina_arcaeformis **
  |     +--+                      
--+        +-- Buldowskia         
  |                               
  |  +-------- Cristaria          
  +--+                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Beringiana         
        +--+                      
           +-- Sinanodonta        

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 8 3 --
  +-------- Anemina_arcaeformis **
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta        

Froufe et al., 2017 [Publ 3802], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 11 9 3 --
  +-------- Anemina_arcaeformis **
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 0 0 --
-- Anemina_arcaeformis

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 8 3 --
  +-------- Anemina_arcaeformis **
--+                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta        

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 4 3 --
  +-------- Cristaria          
--+                            
  |  +----- Anemina_arcaeformis **
  +--+                         
     |  +-- Pletholophus       
     +--+                      
        +-- Sinanodonta        

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 2 2 --
-- Anemina_arcaeformis

Kondakov et al., 2018 [Publ 3914], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 7 7 1 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta        

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 9 5 --
-- Anemina_arcaeformis

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 2 2 --
  +----- Cristaria          
--+                         
  |  +-- Sinanodonta        
  +--+                      
     +-- Anemina_arcaeformis **

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 18 3 2 --
  +----- Anemina_arcaeformis **
--+                         
  |  +-- Cristaria          
  +--+                      
     +-- Sinanodonta        

Riccardi et al., 2019 [Publ 4089], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Sinanodonta        
--+                      
  +-- Anemina_arcaeformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 23 6 --
  +-------- Cristaria          
  |                            
  |  +----- Anemina_arcaeformis **
  +--+                         
--+  |  +-- Amuranodonta       
  |  +--+                      
  |     +-- Buldowskia         
  |                            
  |  +----- Pletholophus       
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 3 2 --
  +----- Anemina_arcaeformis **
--+                         
  |  +-- Cristaria          
  +--+                      
     +-- Sinanodonta        

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 3 1 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta        

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 5 2 --
  +-- Anemina_arcaeformis **
  |                      
--+-- Cristaria          
  |                      
  +-- Sinanodonta        

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 5 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Amuranodonta       

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 5 --
  +----------- Cristaria          
  |                               
--+        +-- Anemina_arcaeformis **
  |  +-----+                      
  |  |     +-- Amuranodonta       
  +--+                            
     |  +----- Pletholophus       
     +--+                         
        |  +-- Sinanodonta        
        +--+                      
           +-- Beringiana         

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Anemina_arcaeformis

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 2 2 --
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta        

Vikhrev et al., 2017 [Publ 3810], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 9 7 1 --
  +-- Anemina_arcaeformis **
--+                      
  +-- Sinanodonta        

Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 5 2 2 --
  +----- Cristaria          
--+                         
  |  +-- Anemina_arcaeformis **
  +--+                      
     +-- Sinanodonta        

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 8 5 --
     +----- Amuranodonta       
  +--+                         
  |  |  +-- Anemina_arcaeformis **
  |  +--+                      
--+     +-- Buldowskia         
  |                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 2 --
-- Anemina_euscaphys

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 12 5 --
     +----- Anemina_arcaeformis **
  +--+                         
  |  |  +-- Amuranodonta       
  |  +--+                      
--+     +-- Buldowskia         
  |                            
  |  +----- Cristaria          
  +--+                         
     |  +-- Beringiana         
     +--+                      
        +-- Sinanodonta        

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