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Page last updated
Sun 07 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validgen_cld&id=376

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Gonideini

genus Leguminaia Conrad, 1865: cladomics
     3 valid species

Cladomics Summary

number of species = 3

analyzed in 34 cladograms in 25 publications
maximum number of terminals in an analysis = 3, mean = 1.24

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 6
maximum number of included terminals = 3, mean = 2.33
proportion of cladograms that support basic monophyly = 0.83

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Lopes-Lima et al., 2021 [Publ 4217], fig. 14

iT tT bi riS oT roT roS iD
3 3 2 7 2 2 2 0.0

*** Top Monophyly Cladogram *** 

           +-- Gonidea                
  +--------+                          
  |        +-- Microcondylaea         
  |                                   
--+  +-------- Leguminaia_anatolica **   
  |  |                                
  |  |  +----- Leguminaia_wheatleyi__2 **
  +--+  |                             
     |  +----- Leguminaia_wheatleyi__3 **
     |  |                             
     +--+----- Leguminaia_wheatleyi__4 **
        |                             
        +----- Leguminaia_wheatleyi__5 **
        |                             
        |  +-- Leguminaia_saulcyi **     
        +--+                          
           +-- Leguminaia_wheatleyi__1 **

Dai et al., 2025 [Publ 4441], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 47 16 11 --

*** Top Sister-Group Cladogram ***

  +-------------------- Sinosolenaia        
  |                                         
  |              +----- Gonidea             
--+  +-----------+                          
  |  |           |  +-- Leguminaia_wheatleyi **
  |  |           +--+                       
  +--+              +-- Microcondylaea      
     |                                      
     |  +-------------- Pseudopostolata     
     |  |                                   
     +--+           +-- Koreosolenaia       
        |  +--------+                       
        |  |        +-- Parvasolenaia       
        +--+                                
           |  +-------- Postolata           
           |  |                             
           +--+     +-- Obovalis            
              |  +--+                       
              |  |  +-- Ptychorhynchus      
              +--+                          
                 |  +-- Guiunio             
                 +--+                       
                    +-- Cosmopseudodon

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 22 0 0 4.0 
  +----- Leguminaia_anatolica **
--+                          
  |  +-- Leguminaia_saulcyi **  
  +--+                       
     +-- Leguminaia_wheatleyi **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
2 13 0 2 123 5 3 15.0 
  +-- Chamberlainia         
  |                         
  +-- Sinohyriopsis         
  |                         
  +-- Lamprotula_caveata    
  |                         
  +-- Lamprotula_leaii      
  |                         
  +-- Potomida              
  |                         
  +-- Pronodularia          
  |                         
--+-- Gonidea               
  |                         
  +-- Leguminaia_wheatleyi **  
  |                         
  +-- Leguminaia **            
  |                         
  +-- Microcondylaea        
  |                         
  +-- Sinosolenaia_carinata 
  |                         
  +-- Sinosolenaia_oleivora 
  |                         
  +-- Sinosolenaia_recognita

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 137 5 3 -- 
  +----------- Sinosolenaia        
--+                                
  |  +-------- Gonidea             
  +--+                             
     |  +----- Microcondylaea      
     +--+                          
        |  +-- Leguminaia_wheatleyi **
        +--+                       
           +-- Leguminaia **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 2 137 5 3 -- 
  +----------- Sinosolenaia_carinata 
  |                                  
  |        +-- Sinosolenaia_oleivora 
--+--------+                         
  |        +-- Sinosolenaia_recognita
  |                                  
  |  +-------- Gonidea               
  +--+                               
     |  +----- Microcondylaea        
     +--+                            
        |  +-- Leguminaia_wheatleyi **  
        +--+                         
           +-- Leguminaia **

Liu et al., 2024 [Publ 4380], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 10 8 6 -- 
  +-------------- Sinosolenaia_carinata
  |                                    
  |     +-------- Inversidens          
--+  +--+                              
  |  |  |  +----- Gonidea              
  |  |  +--+                           
  |  |     |  +-- Leguminaia_anatolica ** 
  +--+     +--+                        
     |        +-- Leguminaia_saulcyi **   
     |                                 
     |  +-------- Sinosolenaia_oleivora
     +--+                              
        |  +----- Postolata            
        +--+                           
           |  +-- Obovalis             
           +--+                        
              +-- Ptychorhynchus

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 198 9 6 -- 
  +-------- Sinosolenaia_carinata 
  |                               
  |  +----- Gonidea               
  +--+                            
  |  |  +-- Leguminaia_wheatleyi **  
  |  +--+                         
  |     +-- Microcondylaea        
--+                               
  |  +----- Ptychorhynchus        
  +--+                            
  |  |  +-- Parvasolenaia         
  |  +--+                         
  |     +-- Koreosolenaia         
  |                               
  |  +----- Sinosolenaia_recognita
  +--+                            
     |  +-- Sinosolenaia_iridinea 
     +--+                         
        +-- Sinosolenaia_oleivora

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Leguminaia_wheatleyi **
  +--+                       
     +-- Gonidea

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 5 5 -- 
  +----------- Sinosolenaia        
  |                                
--+        +-- Ptychorhynchus      
  |  +-----+                       
  |  |     +-- Parvasolenaia       
  +--+                             
     |  +----- Gonidea             
     +--+                          
        |  +-- Microcondylaea      
        +--+                       
           +-- Leguminaia_wheatleyi **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 196 8 6 -- 
  +----------- Sinosolenaia        
  |                                
  |     +----- Gonidea             
--+  +--+                          
  |  |  |  +-- Leguminaia_wheatleyi **
  |  |  +--+                       
  +--+     +-- Microcondylaea      
     |                             
     |  +----- Ptychorhynchus      
     +--+                          
        |  +-- Parvasolenaia       
        +--+                       
           +-- Koreosolenaia

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 7 6 -- 
        +----- Sinosolenaia        
  +-----+                          
  |     |  +-- Ptychorhynchus      
  |     +--+                       
--+        +-- Parvasolenaia       
  |                                
  |  +-------- Inversidens         
  +--+                             
     |  +----- Gonidea             
     +--+                          
        |  +-- Microcondylaea      
        +--+                       
           +-- Leguminaia_wheatleyi **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 198 9 6 -- 
  +----------- Sinosolenaia        
  |                                
  |     +----- Gonidea             
--+  +--+                          
  |  |  |  +-- Leguminaia_wheatleyi **
  |  |  +--+                       
  +--+     +-- Microcondylaea      
     |                             
     |     +-- Ptychorhynchus      
     |     |                       
     +-----+-- Parvasolenaia       
           |                       
           +-- Koreosolenaia

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 48 2 2 -- 
-- Leguminaia_wheatleyi

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 117 8 5 -- 
-- Leguminaia_wheatleyi

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Gonidea             
  +--+                       
     +-- Leguminaia_wheatleyi **

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 7 6 -- 
           +-- Ptychorhynchus      
  +--------+                       
  |        +-- Parvasolenaia       
--+                                
  |  +-------- Inversidens         
  +--+                             
     |  +----- Gonidea             
     +--+                          
        |  +-- Microcondylaea      
        +--+                       
           +-- Leguminaia_wheatleyi **

Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 15 10 -- 
  +-------------------- Sinosolenaia        
  |                                         
  |              +----- Gonidea             
--+  +-----------+                          
  |  |           |  +-- Leguminaia_wheatleyi **
  |  |           +--+                       
  +--+              +-- Microcondylaea      
     |                                      
     |  +-------------- Pseudopostolata     
     |  |                                   
     +--+           +-- Parvasolenaia       
        |  +--------+                       
        |  |        +-- Koreosolenaia       
        +--+                                
           |  +-------- Postolata           
           +--+                             
              |  +----- Ptychorhynchus      
              +--+                          
                 |  +-- Obovalis            
                 +--+                       
                    +-- Cosmopseudodon

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 44 13 9 -- 
  +-------------- Sinosolenaia        
  |                                   
  |        +----- Gonidea             
--+  +-----+                          
  |  |     |  +-- Leguminaia_wheatleyi **
  |  |     +--+                       
  |  |        +-- Microcondylaea      
  +--+                                
     |        +-- Koreosolenaia       
     |  +-----+                       
     |  |     +-- Parvasolenaia       
     +--+                             
        |  +----- Postolata           
        +--+                          
           |  +-- Ptychorhynchus      
           +--+                       
              +-- Obovalis

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Gonidea             
  +--+                       
     +-- Leguminaia_wheatleyi **

Froufe et al., 2016 [Publ 3727], fig. 3_4

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 -- 
-- Leguminaia_wheatleyi

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 111 8 5 -- 
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia_wheatleyi **     
  |                            
  +-- Microcondylaea           
--+                            
  +-- Gonidea                  
  |                            
  +-- Ptychorhynchus           
  |                            
  +-- Parvasolenaia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 5 4 -- 
  +-------- Sinosolenaia        
--+                             
  |  +----- Gonidea             
  +--+                          
     |  +-- Microcondylaea      
     +--+                       
        +-- Leguminaia_wheatleyi **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Leguminaia_wheatleyi **
  +--+                       
     +-- Gonidea

Jeratthitikul et al., 2025 [Publ 4423], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 49 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Gonidea             
  +--+                       
     +-- Leguminaia_wheatleyi **

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 -- 
-- Leguminaia_wheatleyi

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 81 1 1 -- 
  +-- Gonidea             
--+                       
  +-- Leguminaia_wheatleyi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 120 11 8 -- 
     +-------- Sinosolenaia        
  +--+                             
  |  |  +----- Gonidea             
  |  +--+                          
  |     |  +-- Leguminaia_wheatleyi **
--+     +--+                       
  |        +-- Microcondylaea      
  |                                
  |  +-------- Inversidens         
  |  |                             
  +--+     +-- Obovalis            
     |  +--+                       
     |  |  +-- Ptychorhynchus      
     +--+                          
        |  +-- Parvasolenaia       
        +--+                       
           +-- Koreosolenaia

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 57 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Gonidea             
  +--+                       
     +-- Leguminaia_wheatleyi **

Kongim et al., 2023 [Publ 4290], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 2 2 -- 
  +----- Sinosolenaia        
--+                          
  |  +-- Leguminaia_wheatleyi **
  +--+                       
     +-- Gonidea

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 44 13 9 -- 
  +-------------- Sinosolenaia        
  |                                   
  |        +----- Gonidea             
--+  +-----+                          
  |  |     |  +-- Leguminaia_wheatleyi **
  |  |     +--+                       
  |  |        +-- Microcondylaea      
  +--+                                
     |        +-- Koreosolenaia       
     |  +-----+                       
     |  |     +-- Parvasolenaia       
     +--+                             
        |  +----- Postolata           
        +--+                          
           |  +-- Ptychorhynchus      
           +--+                       
              +-- Obovalis

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 7 6 -- 
        +----- Gonidea             
  +-----+                          
  |     |  +-- Leguminaia_wheatleyi **
  |     +--+                       
--+        +-- Microcondylaea      
  |                                
  |  +-------- Inversidens         
  +--+                             
     |  +----- Sinosolenaia        
     +--+                          
        |  +-- Parvasolenaia       
        +--+                       
           +-- Ptychorhynchus

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 82 2 2 -- 
-- Leguminaia_wheatleyi

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 26 1 1 -- 
-- Leguminaia_wheatleyi

Page updated: 08:17:44 Sun 07 Sep 2025
 
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