| Page last updated
Sun 07 Sep 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Lopes-Lima et al. (2021), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work. The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=publ&id=4217
Search the database pages:
|
|
Publication information
Lopes-Lima, M., M.E. Gürlek, Ü. Kebapçi, H. ŞerefliŞan, T. Yanik et al. 2021. Diversity, biogeography, evolutionary relationships, and conservation of Eastern Mediterranean freshwater mussels (Bivalvia: Unionidae). Molecular Phylogenetics and Evolution 163. 107261 (23 pp.).
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 1
fig. 11
fig. 14
fig. 2
fig. 6
Attributed Species Names
Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
Chen et al., 2025, Zoosyst. Evol. 101(4): Tables S2-S3.
Dai et al., 2025, Zoosyst. Evol. 101(2): Tables 1-2.
(misidentified reference to Unio crassus Retzius, 1788)
|
Lamellidens corrianus Lopes-Lima et al., 2021
Lopes-Lima et al., 2021, MPE 163: Table S1.
(misidentified reference to Unio corrianus Lea, 1834)
|
Anodonta seddoni Lopes-Lima et al., 2021
Gürlek, Kebapçı & Lopes-Lima in Lopes-Lima et al., 2021, MPE 163: 19, Table 2, fig. 5.
|
Leguminaia anatolica Lopes-Lima et al., 2021
Gürlek, Kebapçı and Lopes-Lima in Lopes-Lima et al., 2021, MPE 163: 19, Table 2, fig. 5.
|
Unio trachaea Lopes-Lima et al., 2021
‘Rolle & Kobelt in Rolle & Kobelt’ Lopes-Lima et al., 2021, MPE 163: Supplementary Appendex 1, p. 10.
(error for Unio tracheae Kobelt & Rolle, 1895)
|
Unio bruguierianus Lopes-Lima et al., 2021
‘Bourguignat’ of Authors.
Lopes-Lima et al., 2021, MPE 163: Table S1. [in part]
(misidentified reference to Unio bruguierianus Bourguignat, 1853)
|
Unio bruguierianus Lopes-Lima et al., 2021
‘Bourguignat’ of Authors.
Lopes-Lima et al., 2021, MPE 163: Table S1. [in part]
(misidentified reference to Unio bruguierianus Bourguignat, 1853)
|
Taxonomic Opinions
Amblema plicata
Anodonta anatina (Linnaeus, 1758)
+ Anodonta taurica Bourguignat, 1880
+ Anodonta apollonica Bourguignat, 1880
+ Anodonta gallandi Bourguignat, 1880
+ Anodonta cyrea Drouët, 1881
+ Anodonta byzantina Drouët, 1881
+ Anodonta ectina Bourguignat, 1881
+ Anodonta gaudioni Drouët, 1881
+ Anodonta georgiana Drouët, 1881
+ Anodonta lenkoranensis Drouët, 1881
+ Anodonta mahometana Bourguignat, 1881
+ Anodonta maritzana Bourguignat, 1881
+ Anodonta pseudodopsis Locard, 1883
+ Anodonta brandti Drouët, 1888
+ Anodonta cilicica Kobelt & Rolle, 1895
+ Anodonta cygnea var. armenica Ovtschinnikov, 1935
Anodonta cygnea (Linnaeus, 1758)
+ Anodonta gravida Drouët, 1879
Anodonta exulcerata
Anodonta seddoni sp. nov.
Anodonta vescoiana Bourguignat, 1856
+ Anodonta schlaeflii Mousson, 1874
+ Anodonta vescoiana var. mesopotamica Mousson, 1874
+ Anodonta (Euphrata) bahlikiana Pallary, 1939
Anodontoides denigratus
Coelatura aegyptiaca
Gonidea angulata (Lea, 1838)
+ Anodonta randalli Trask, 1854
+ Anodonta randalli Trask, 1854
Lamellidens corrianus
Lamprotula leaii
Lampsilis ovata
Leguminaia anatolica sp. nov.
Leguminaia cf. saulcyi (Bourguignat, 1852)
+ Unio michonii Bourguignat, 1852
+ Unio tripolitanus Bourguignat, 1852
+ Pseudodon chantrei Locard, 1883
+ Leguminaia (sec. Leguminaia) locardi Simpson, 1900
+ Leguminaia tripolitana var. kebirensis Pallary, 1939
Leguminaia cf. wheatleyi (Lea, 1862)
+ Monocondyloea mardinensis Lea, 1865
+ Leguminaia bourguignati Locard, 1883
+ Leguminaia wheatleyi var. euphratica Kobelt, 1912
+ Leguminaia (mardinensis var.) naegelei Kobelt, 1913
+ Leguminaia graeteri Kobelt, 1913
+ Leguminaia nisibina Kobelt, 1913
+ Microdondylaea haasii Kobelt, 1914
+ Leguminaia mardinensis var. truncata Pallary, 1939
Microcondylaea bonellii
Middendorffinaia mongolica
Nodularia douglasiae
Potomida acarnanica
Potomida littoralis (Cuvier, 1798)
+ Mysca (Potomida) corrugata Swainson, 1840
Potomida semirugata (Lamarck, 1819)
+ Unio delesserti Bourguignat, 1852
+ Unio rothi Bourguignat, 1863
+ Unio emesaensis Lea, 1865
+ Unio simonis Tristram, 1865
+ Unio ferussacianus Lea, 1868
+ Unio galilaei Locard, 1883
+ Unio luynesi Locard, 1883
+ Unio rhomboidopsis Locard, 1883
+ Unio timius Locard, 1883
+ Unio tinctus Drouët, 1893
+ Unio rollei Kobelt & Rolle, 1895
+ Unio lycicus Kobelt & Rolle, 1895
+ Unio wagneri Kobelt & Rolle, 1895
+ Unio tracheae Kobelt & Rolle, 1895
+ Unio (Rhombunio ?) abrus Kobelt, 1913
+ Unio graeteri Kobelt, 1913
+ Unio blanchianus Kobelt, 1913
+ Unio (Rhombunio) babensis Kobelt, 1913
+ Unio deschampsi Kobelt, 1913
+ Unio (cilicicus) var. langloisi Kobelt, 1913
+ Unio (Rhombunio) naegelei Kobelt, 1913
+ Unio beroeus Kobelt, 1913
+ Unio (Rhombunio) halepensis Kobelt, 1913
+ Unio (Rhombunio) halepensis var. cazioti Kobelt, 1913
+ Unio corbiculiformis Kobelt, 1913
+ Unio chinnerethensis Preston, 1913
+ Unio salamboana Pallary, 1929
Pronodularia japanensis
Pseudanodonta complanata
Pseudodontopsis euphratica Bourguignat, 1852
+ Unio opperti Bourguignat, 1856
+ Unio churchillianus Bourguignat, 1857
+ Monocondyloea rhomboidea Lea, 1859
+ Pseudodon pachyolenus Bourguignat, 1880
+ Pseudodontopsis babylonica Kobelt, 1913
+ Pseudodontopsis piestius Kobelt, 1913
Pseudunio homsensis
Pyganodon cataracta
Sinanodonta woodiana
Unio bruguierianus Bourguignat, 1853
+ Unio orientalis Bourguignat, 1852
+ Unio prusii Bourguignat, 1856
+ Unio vescoi Bourguignat, 1856
+ Unio schwerzenbachi Bourguignat, 1856
+ Unio kotschyi Küster, 1862
+ Unio turcicus Küster, 1862
+ Unio kullethensis Lea, 1865
+ Unio mardinensis Lea, 1865
+ Unio orphaensis Lea, 1865
+ Unio vicarius Westerlund & Blanc, 1879
+ Unio rothi var. komarowi O. Boettger, 1880
+ Unio araxenus Drouët, 1881
+ Unio heldreichi O. Boettger, 1886
+ Unio thiesseae Drouët, 1892
+ Unio (sec. Lymnium) pseudonymus Simpson, 1900
+ Unio desectus f. pursacensis Sturany, 1902
+ Unio armeniacus Kobelt, 1912
+ Unio modiola Preston, 1912
+ Unio subbatavus Kobelt, 1913
+ Unio specialis Kobelt, 1913
+ Unio ancyrensis Kobelt, 1913
+ Unio ancyrensis var. louisei Kobelt, 1913
+ Unio (orfaensis var. ?) ehrmanni Kobelt, 1913
+ Unio diarbekirianus Kobelt, 1913
+ Unio bitlisensis Kobelt, 1913
+ Unio ciconius Kobelt, 1913
+ Unio medicus Kobelt, 1913
+ Unio thospiensis Kobelt, 1915
+ Unio (Rhombunio) wenzii Kobelt, 1915
+ Unio crassus Araujo et al., 2018
Unio caffer
Unio cf. courtillieri
Unio cf. desectus
Unio crassus
Unio damascensis Lea, 1863
+ Unio orontesensis Lea, 1863
+ Unio syriacus Lea, 1863
Unio delicatus Lea, 1863
+ Unio anemprosthus Locard, 1883
+ Unio antiochianus Locard, 1883
+ Unio axiacus Locard, 1883
+ Unio chantrei Locard, 1883
+ Unio jauberti Locard, 1883
+ Unio subtigridis Locard, 1883
+ Unio alexandri Kobelt & Rolle, 1895
+ Unio cilicicus Kobelt & Rolle, 1895
+ Unio cilicicus var. jenemterensis Kobelt & Rolle, 1895
+ Unio cilicicus var. adanensis Kobelt & Rolle, 1895
+ Unio cilicicus var. subsaccatus Kobelt & Rolle, 1895
+ Unio (cilicicus var. ?) berytensis Kobelt & Rolle, 1895
+ Unio kobelti Kobelt & Rolle, 1895
+ Unio kuweikensis Kobelt, 1913
+ Unio (cilicicus var.) anceyi Kobelt, 1913
+ Unio schwarzii Kobelt, 1915
+ Unio tigridis Araujo et al., 2018
Unio delphinus
Unio durieui
Unio elongatulus
Unio eucirrus Bourguignat, 1857
Unio foucauldianus
Unio gibbus
Unio hueti Bourguignat, 1855
Unio ionicus
Unio mancus
Unio pictorum (Linnaeus, 1758)
+ Unio gaudioni Drouët, 1881
+ Unio gentilis Hesse, 1911
+ Unio ascanius Kobelt, 1913
Unio ravoisieri
Unio sesirmensis Kobelt, 1913
Unio terminalis Bourguignat, 1852
+ Unio grelloisianus Bourguignat, 1856
+ Unio jordanicus Bourguignat, 1856
+ Unio lunulifer Bourguignat, 1856
+ Unio lorteti Locard, 1883
+ Unio pietri Locard, 1883
+ Unio ellipsoideus Locard, 1883
+ Unio genezarethanus Locard, 1883
+ Unio prosacrus Locard, 1883
+ Unio raymondi Locard, 1883
+ Unio tiberiadensis Locard, 1883
+ Unio tristrami Locard, 1883
+ Unio zabulonicus Locard, 1883
+ Unio herodes Kobelt & Rolle, 1895
+ Unio kisonis Kobelt & Rolle, 1895
+ Unio raymondopsis Kobelt, 1913
+ Unio tigridis Araujo et al., 2018
Unio tigridis Bourguignat, 1852
+ Unio mussolianus Küster, 1861
+ Unio bourguignatianus Lea, 1863
+ Unio dignatus Lea, 1863
+ Unio mosulensis Lea, 1863
+ Unio rasus Lea, 1863
+ Margaron (Unio) tigris Lea, 1870
+ Unio tigridis var. haruni Kobelt, 1912
+ Unio calliopsis Kobelt, 1913
+ Unio hyperamblius Kobelt, 1913
+ Unio siouffi Kobelt, 1913
+ Unio dignatus assuricus Kobelt, 1913
+ Unio dignatus ninusi Kobelt, 1913
+ Unio dignatus semiramidis Kobelt, 1913
+ Unio (Nemrodia) tammouziana Pallary, 1939
+ Unio (Nemrodia) tammouziana var. major Pallary, 1939
+ Unio (Nemrodia) tammouziana var. minor Pallary, 1939
+ Unio (Nemrodia) tigris [sic] var. cuneiformis Pallary, 1939
+ Unio (Nemrodia) tigris [sic] var. khabourensis Pallary, 1939
Unio tumidiformis
Unio tumidus Philipsson in Retzius, 1788
Utterbackia peggyae
|
Cladogram Figure 1
+----- Potomida_littoralis
+--+
| | +-- Potomida_semirugata
| +--+
+--------------------+ +-- Potomida_acarnanica
| |
| | +----- Leguminaia_anatolica
| +--+
| | +-- Leguminaia_saulcyi
| +--+
--+ +-- Leguminaia_wheatleyi
|
| +----- Anodonta_anatina
| +--------------------+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_seddonae
+--+
| +----------------------- Unio_tumidus
| |
| | +-------- Unio_damascensis
| | +-----------+
+--+ | | +----- Unio_sesirmensis
| | +--+
| | | +-- Unio_bruguierianus
| | +--+
+--+ +-- Unio_nanus
|
| +-- Unio_delicatus
| +--------------+
| | +-- Unio_terminalis
| |
+--+ +-- Unio_tigridis
| +-----------+
| | +-- Unio_hueti
| |
+--+ +----------- Unio_ravoisieri
| |
| | +-- Unio_pictorum
+--+ +-----+
| | +-- Unio_eucirrus
| |
+--+ +-- Unio_elongatulus
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_foucauldianus
+--+
+-- Unio_delphinus
Cladogram Figure 11
+-------- Anodonta_anatina
--+
| +----- Anodonta_exulcerata
+--+
| +-- Anodonta_seddonae
+--+
+-- Anodonta_cygnea
Cladogram Figure 14
+-- Gonidea_angulata
+--------+
| +-- Microcondylaea_bonellii
|
--+ +-------- Leguminaia_anatolica
| |
| | +----- Leguminaia_wheatleyi__2
+--+ |
| +----- Leguminaia_wheatleyi__3
| |
+--+----- Leguminaia_wheatleyi__4
|
+----- Leguminaia_wheatleyi__5
|
| +-- Leguminaia_saulcyi
+--+
+-- Leguminaia_wheatleyi__1
Cladogram Figure 2
+----------------- Unio_tumidus
|
--+ +-------------- Unio_bruguierianus
| |
| | +-- Unio_delicatus
+--+ +--------+
| | +-- Unio_terminalis
| |
+--+ +-- Unio_tigridis
| +-----+
| | +-- Unio_hueti
| |
+--+ +-- Unio_pictorum
| +--+
| | +-- Unio_eucirrus
+--+
| +-- Unio_elongatulus
+--+
+-- Unio_mancus
Cladogram Figure 6
+----------------- Unio_tumidus
--+
| +-------------- Unio_pictorum
+--+
| +----------- Unio_sesirmensis
+--+
| +-------- Unio_damascensis
+--+
| +----- Unio_crassus
+--+
| +-- Unio
+--+
+-- Unio_nanus
|
Page updated: 08:31:36 Sun 07 Sep 2025 |
|
