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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Beringiana Starobogatov in Zatravkin, 1983: cladomics
     4 valid species

Cladomics Summary

number of species = 4

analyzed in 36 cladograms in 25 publications
maximum number of terminals in an analysis = 4, mean = 2.06

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 16
maximum number of included terminals = 4, mean = 3.38
proportion of cladograms that support basic monophyly = 0.94


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iT tT bi riS oT roT roS iD
1 3 0 20 11 4 2 0.0

*** Top Monophyly Cladogram ***

  +-------------- Cristaria               
  |                                       
  |  +----------- Sinanodonta_tumens      
  |  |                                    
  |  |        +-- Sinanodonta_lauta__5    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta__6    
  |  |        |                           
  |  |        +-- Beringiana_japonica__15 ** 
  |  |        |                           
  |  |        +-- Beringiana_japonica__16 ** 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica__17 ** 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica__18 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__19 ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica__20 ** 
     |  |                                 
     |  |  +----- Beringiana_japonica__1 **  
     |  |  |                              
     |  |  +----- Beringiana_japonica__2 **  
     |  |  |                              
     +--+  +----- Beringiana_japonica__3 **  
        |  |                              
        |  +----- Beringiana_japonica__4 **  
        |  |                              
        |  +----- Beringiana_japonica__5 **  
        |  |                              
        |  +----- Sinanodonta_lauta__1    
        |  |                              
        |  +----- Beringiana_japonica__6 **  
        |  |                              
        |  +----- Sinanodonta_lauta__2    
        |  |                              
        |  +----- Sinanodonta_calipygos__1
        |  |                              
        +--+----- Beringiana_japonica__7 **  
           |                              
           +----- Sinanodonta_lauta__4    
           |                              
           +----- Beringiana_japonica__8 **  
           |                              
           +----- Beringiana_japonica__9 **  
           |                              
           +----- Beringiana_japonica__10 ** 
           |                              
           +----- Beringiana_japonica__11 ** 
           |                              
           +----- Beringiana_japonica__12 ** 
           |                              
           +----- Beringiana_japonica__13 ** 
           |                              
           +----- Beringiana_japonica__14 ** 
           |                              
           |  +-- Sinanodonta_lauta__3    
           +--+                           
              +-- Sinanodonta_calipygos__2

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 27 23 6 28.0

*** Top Sister-Group Cladogram ***

  +-------------------- Cristaria             
  |                                           
  |              +----- Amuranodonta          
--+  +-----------+                            
  |  |           |  +-- Anemina               
  |  |           +--+                         
  +--+              +-- Buldowskia            
     |                                        
     |  +-------------- Pletholophus          
     +--+                                     
        |  +----------- Sinanodonta           
        +--+                                  
           |  +-------- Beringiana_japonica **   
           +--+                               
              |  +----- Beringiana_beringiana ** 
              +--+                            
                 |  +-- Beringiana_fukuharai **  
                 +--+                         
                    +-- Beringiana_gosannensis **

Bulakhova et al., 2023 [Publ 4325], fig. 5

iT tT bi riS oT roT roS iD
4 4 2 4 0 0 0 28.0
     +-- Beringiana_beringiana ** 
  +--+                         
  |  +-- Beringiana_japonica **   
--+                            
  |  +-- Beringiana_gosannensis **
  +--+                         
     +-- Beringiana_fukuharai **  

Sano et al., 2022 [Publ 4311], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 8 8 4 28.0
           +----- Pletholophus          
  +--------+                            
  |        |  +-- Cristaria             
  |        +--+                         
--+           +-- Buldowskia            
  |                                     
  |  +----------- Sinanodonta           
  +--+                                  
     |  +-------- Beringiana_fukuharai **  
     +--+                               
        |  +----- Beringiana_japonica **   
        +--+                            
           |  +-- Beringiana_gosannensis **
           +--+                         
              +-- Beringiana_beringiana ** 

Sano et al., 2022 [Publ 4311], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 14 14 4 28.0
           +----- Pletholophus          
  +--------+                            
  |        |  +-- Cristaria             
  |        +--+                         
--+           +-- Buldowskia            
  |                                     
  |  +----------- Sinanodonta           
  +--+                                  
     |  +-------- Beringiana_fukuharai **  
     +--+                               
        |  +----- Beringiana_gosannensis **
        +--+                            
           |  +-- Beringiana_japonica **   
           +--+                         
              +-- Beringiana_beringiana ** 

Dai et al., 2024 [Publ 4385], fig. 3-ML

iT tT bi riS oT roT roS iD
4 4 2 4 27 23 6 28.0
              +----- Amuranodonta          
  +-----------+                            
  |           |  +-- Anemina               
  |           +--+                         
--+              +-- Buldowskia            
  |                                        
  |  +-------------- Cristaria             
  |  |                                     
  +--+  +----------- Pletholophus          
     |  |                                  
     +--+  +-------- Sinanodonta           
        |  |                               
        +--+     +-- Beringiana_fukuharai **  
           |  +--+                         
           |  |  +-- Beringiana_gosannensis **
           +--+                            
              |  +-- Beringiana_beringiana ** 
              +--+                         
                 +-- Beringiana_japonica **   

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
4 4 2 4 116 20 6 28.0
  +----------------- Cristaria             
  |                                        
  |           +----- Anemina               
  +-----------+                            
--+           |  +-- Amuranodonta          
  |           +--+                         
  |              +-- Buldowskia            
  |                                        
  |  +-------------- Pletholophus          
  +--+                                     
     |  +----------- Sinanodonta           
     +--+                                  
        |  +-------- Beringiana_japonica **   
        +--+                               
           |  +----- Beringiana_beringiana ** 
           +--+                            
              |  +-- Beringiana_fukuharai **  
              +--+                         
                 +-- Beringiana_gosannensis **

Dai et al., 2024 [Publ 4385], fig. 3-BI

iT tT bi riS oT roT roS iD
4 4 2 4 27 23 6 28.0
                 +----- Amuranodonta          
  +--------------+                            
  |              |  +-- Anemina               
  |              +--+                         
--+                 +-- Buldowskia            
  |                                           
  |  +----------------- Cristaria             
  +--+                                        
     |  +-------------- Pletholophus          
     +--+                                     
        |  +----------- Sinanodonta           
        +--+                                  
           |  +-------- Beringiana_japonica **   
           +--+                               
              |  +----- Beringiana_beringiana ** 
              +--+                            
                 |  +-- Beringiana_fukuharai **  
                 +--+                         
                    +-- Beringiana_gosannensis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iT tT bi riS oT roT roS iD
4 4 2 4 4 2 1 28.0
  +----------- Sinanodonta           
--+                                  
  |  +-------- Beringiana_gosannensis **
  +--+                               
     |  +----- Beringiana_fukuharai **  
     +--+                            
        |  +-- Beringiana_beringiana ** 
        +--+                         
           +-- Beringiana_japonica **   

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 66 9 5 28.0
  +----------- Sinanodonta          
--+                                 
  |  +-------- Pletholophus         
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 30 6 4 28.0
           +-- Cristaria            
  +--------+                        
  |        +-- Buldowskia           
  |                                 
--+        +-- Pletholophus         
  |  +-----+                        
  |  |     +-- Sinanodonta          
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 66 9 5 28.0
  +----------- Pletholophus         
--+                                 
  |  +-------- Sinanodonta          
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 66 9 5 28.0
  +----------- Sinanodonta          
--+                                 
  |  +-------- Pletholophus         
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
3 22 5 5 31 7 5 24.0
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica **    
  |                             
  +----- Beringiana_fukuharai__1 **
  |                             
  +----- Beringiana_beringiana **  
  |                             
  +----- Beringiana_fukuharai__2 **
  |                             
  +----- Beringiana_fukuharai__3 **
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Sano et al., 2017 [Publ 3799], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 18 5 4 28.0
              +-- Cristaria            
  +-----------+                        
  |           +-- Buldowskia           
--+                                    
  |  +----------- Pletholophus         
  +--+                                 
     |  +-------- Sinanodonta          
     +--+                              
        |  +----- Beringiana_fukuharai ** 
        +--+                           
           |  +-- Beringiana_japonica **  
           +--+                        
              +-- Beringiana_beringiana **

Douda et al., 2024 [Publ 4364], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 12 11 1 --
  +----- Sinanodonta          
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Beringiana_japonica **  

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iT tT bi riS oT roT roS iD
2 2 0 2 12 10 1 --
              +-- Beringiana_japonica **  
  +-----------+                        
  |           +-- Beringiana_beringiana **
  |                                    
--+           +-- Sinanodonta_angula   
  |  +--------+                        
  |  |        +-- Sinanodonta_lucida   
  |  |                                 
  +--+  +-------- Sinanodonta_schrenkii
     |  |                              
     |  |     +-- Sinanodonta_calipygos
     |  +-----+                        
     +--+     +-- Sinanodonta_tumens   
        |                              
        |  +----- Sinanodonta_woodiana 
        |  |                           
        +--+----- Sinanodonta_jourdyi  
           |                           
           |  +-- Sinanodonta_elliptica
           +--+                        
              +-- Sinanodonta_lauta    

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 3 --
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta         

Froufe et al., 2017 [Publ 3802], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 11 9 3 --
  +-------- Anemina              
--+                              
  |  +----- Cristaria            
  +--+                           
     |  +-- Beringiana_beringiana **
     +--+                        
        +-- Sinanodonta          

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 12 5 --
     +----- Anemina              
  +--+                           
  |  |  +-- Amuranodonta         
  |  +--+                        
--+     +-- Buldowskia           
  |                              
  |  +----- Cristaria            
  +--+                           
     |  +-- Beringiana_beringiana **
     +--+                        
        +-- Sinanodonta          

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 6 3 --
     +-- Cristaria           
  +--+                       
  |  +-- Anemina             
--+                          
  |  +-- Beringiana_fukuharai **
  +--+                       
     +-- Sinanodonta         

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 5 --
  +----------- Cristaria            
  |                                 
--+        +-- Anemina              
  |  +-----+                        
  |  |     +-- Amuranodonta         
  +--+                              
     |  +----- Pletholophus         
     +--+                           
        |  +-- Sinanodonta          
        +--+                        
           +-- Beringiana_beringiana **

Klishko et al., 2016 [Publ 3733], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 2 --
  +----- Cristaria            
--+                           
  |  +-- Sinanodonta          
  +--+                        
     +-- Beringiana_beringiana **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 3 --
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta         

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
1 1 0 1 103 0 0 --
-- Beringiana_beringiana

Klishko et al., 2014 [Publ 3666], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 2 --
  +----- Cristaria            
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Sinanodonta          

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Beringiana_beringiana

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 5 --
  +----- Pletholophus         
--+                           
  |  +-- Sinanodonta          
  +--+                        
     +-- Beringiana_beringiana **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 6 3 --
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta         

Sayenko et al., 2017 [Publ 3760], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 1 --
  +-- Beringiana_beringiana **
--+                        
  +-- Sinanodonta          

Soroka, 2010 [Publ 3404], fig. 9a

iT tT bi riS oT roT roS iD
1 1 0 1 4 4 2 --
  +----- Buldowskia           
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Sinanodonta          

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 53 6 3 --
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta         

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 8 5 --
     +----- Amuranodonta         
  +--+                           
  |  |  +-- Anemina              
  |  +--+                        
--+     +-- Buldowskia           
  |                              
  |  +----- Cristaria            
  +--+                           
     |  +-- Beringiana_beringiana **
     +--+                        
        +-- Sinanodonta          

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 --
  +-- Beringiana_beringiana **
--+                        
  +-- Sinanodonta          

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 3 3 --
  +-------- Cristaria           
--+                             
  |  +----- Anemina             
  +--+                          
     |  +-- Beringiana_fukuharai **
     +--+                       
        +-- Sinanodonta         

Kano et al., 2019 [Publ 4194], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 9 2 2 --
  +----- Sinanodonta        
--+                         
  |  +-- Pletholophus       
  +--+                      
     +-- Beringiana_japonica **

Page updated: 14:57:28 Wed 04 Dec 2024

 
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