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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae

tribe ACUTICOSTINI Starobogatov, 1967: cladomics
     1 genus

Cladomics Summary

number of species = 5
number of subtaxa (rank: genus) = 1

analyzed in 50 cladograms in 31 publications
maximum number of terminals in an analysis = 2, mean = 1.04
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 2
maximum number of included terminals = 2, mean = 2.0
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0


Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 68 4 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----- Acuticosta **   
  |                   
  +----- Lepidodesmini
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolariini
  +--+                
     +-- Anodontini   

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 1 43 29 4 --
  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticosta **   
     |  |                      
     +--+     +-- Lepidodesmini
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolariini
           +--+                
              +-- Unionini     

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 21 13 3 --
     +----- Schistodesmus
  +--+                   
  |  |  +-- Lanceolariini
  |  +--+                
--+     +-- Acuticosta **   
  |                      
  |  +----- Aculamprotula
  +--+                   
     |  +-- Tchangsinaia 
     +--+                
        +-- Nodularia    

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 35 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 16 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 39 25 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini   

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 25 4 --
  +----------- Acuticosta **   
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Lepidodesmini
        +--+                
           +-- Anodontini   

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 8 3 --
  +----------- Anodontini          
  |                                
--+        +-- Nodularia           
  |  +-----+                       
  |  |     +-- Cuneopsis           
  +--+                             
     |  +----- Lanceolaria_grayii  
     |  |                          
     +--+----- Acuticosta **          
        |                          
        |  +-- Lanceolaria_gladiola
        +--+                       
           +-- Aculamprotula       

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 46 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 14 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini   

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 20 3 --
  +-------- Acuticosta **   
--+                      
  |  +----- Lanceolariini
  +--+                   
     |  +-- Anodontini   
     +--+                
        +-- Unionini     

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
1 1 0 1 38 38 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini   

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 32 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 12 2 --
  +----- Anodontini
--+                
  |  +-- Acuticosta **
  +--+             
     +-- Unionini  

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 43 4 --
  +----- Acuticosta **           
  |                           
  +----- Alasmidonta          
  |                           
  +----- Anodonta_anatina     
  |                           
  +----- Anemina              
  |                           
  +----- Anodonta_cygnea      
  |                           
  +----- Cristaria            
  |                           
  +----- Anodonta_nuttalliana 
  |                           
  +----- Lasmigona            
  |                           
  +----- Lepidodesmini        
  |                           
  +----- Pseudanodonta        
  |                           
  +----- Pyganodon            
  |                           
--+----- Simpsonaias          
  |                           
  +----- Strophitus           
  |                           
  +----- Schistodesmus        
  |                           
  +----- Utterbackia          
  |                           
  +----- Lanceolariini        
  |                           
  +----- Unio                 
  |                           
  +----- Aculamprotula        
  |                           
  |  +-- Pletholophus         
  +--+                        
  |  +-- Sinanodonta          
  |                           
  |  +-- Cuneopsis_heudei     
  |  |                        
  |  +-- Cuneopsis_celtiformis
  |  |                        
  +--+-- Tchangsinaia         
     |                        
     +-- Cuneopsis_rufescens  
     |                        
     +-- Nodularia            

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 27 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 41 27 4 --
  +-------------------------------- Anemina              
  |                                                      
--+  +----------------------------- Acuticosta **           
  |  |                                                   
  +--+  +-------------------------- Sinanodonta          
     |  |                                                
     +--+  +----------------------- Aculamprotula        
        |  |                                             
        +--+                    +-- Cristaria            
           |  +-----------------+                        
           |  |                 +-- Lanceolariini        
           +--+                                          
              |  +----------------- Lepidodesmini        
              +--+                                       
                 |  +-------------- Schistodesmus        
                 +--+                                    
                    |  +----------- Nodularia            
                    +--+                                 
                       |  +-------- Cuneopsis_rufescens  
                       +--+                              
                          |  +----- Tchangsinaia         
                          +--+                           
                             |  +-- Cuneopsis_heudei     
                             +--+                        
                                +-- Cuneopsis_celtiformis

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 13 4 --
  +----------- Anodontini   
  |                         
  |     +----- Acuticosta **   
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesmini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 46 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 15 4 --
  +----------------- Acuticosta **      
  |                                  
  +----------------- Cristariina     
  |                                  
  +----------------- Anodontina      
--+                                  
  |  +-------------- Lanceolariini   
  |  |                               
  |  |           +-- Aculamprotula   
  |  +-----------+                   
  +--+           +-- Lepidodesmini   
     |                               
     |  +----------- Unio            
     |  |                            
     +--+  +-------- Middendorffinaia
        |  |                         
        +--+  +----- Cuneopsis       
           |  |                      
           +--+----- Nodularia       
              |                      
              |  +-- Inversiunio     
              +--+                   
                 +-- Schistodesmus   

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 13 4 --
  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesmini
  |                      
  +-------- Acuticosta **   
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolariini
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 43 4 --
  +-------- Acuticosta **   
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 15 4 --
  +----------- Anodontina   
  |                         
--+        +-- Acuticosta **   
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Unionini     

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 42 4 --
  +----------- Acuticosta **   
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Anodontini   

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 26 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 26 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini   

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 21 4 --
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesmini          
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticosta **             
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 16 4 --
  +----------------- Anodontini   
  |                               
  |              +-- Acuticosta **   
--+  +-----------+                
  |  |           +-- Lanceolariini
  |  |                            
  +--+           +-- Lepidodesmini
     |  +--------+                
     |  |        +-- Aculamprotula
     +--+                         
        |  +-------- Unio         
        +--+                      
           |  +----- Schistodesmus
           +--+                   
              |  +-- Nodularia    
              +--+                
                 +-- Tchangsinaia 

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 20 4 --
  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticosta **   
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesmini
           +--+                
              +-- Lanceolariini

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 17 3 --
  +-------- Unionini     
--+                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Lanceolariini

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 3 1 --
  +-- Acuticosta **
--+             
  +-- Unionini  

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 46 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 25 4 --
  +-------- Acuticosta **   
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 15 3 --
  +-------------- Aculamprotula
  |                            
--+           +-- Acuticosta **   
  |  +--------+                
  |  |        +-- Lanceolariini
  +--+                         
     |  +-------- Unio         
     +--+                      
        |  +----- Nodularia    
        +--+                   
           |  +-- Schistodesmus
           +--+                
              +-- Cuneopsis    

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 46 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 32 4 --
  +-------- Lepidodesmini
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Lanceolariini

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 21 3 --
  +-------- Acuticosta **            
  |                               
--+  +----- Lanceolaria_grayii    
  |  |                            
  |  +----- Lanceolaria_gladiola  
  +--+                            
     +----- Lanceolaria_lanceolata
     |                            
     |  +-- Anodontini            
     +--+                         
        +-- Unionini              

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 30 4 --
           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola  
     |  |                            
     |  +----- Acuticosta **            
     |  |                            
     |  +----- Lepidodesmini         
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina            

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 33 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 30 4 --
  +-------- Acuticosta **   
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 13 3 --
-- Acuticosta_chinensis

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 9 2 --
  +-------- Aculamprotula
--+                      
  |  +----- Acuticosta **   
  +--+                   
     |  +-- Nodularia    
     +--+                
        +-- Cuneopsis    

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 21 3 --
  +----------- Acuticosta **            
  |                                  
--+        +-- Lanceolaria_lanceolata
  |  +-----+                         
  |  |     +-- Lanceolaria_grayii    
  +--+                               
     |  +----- Lanceolaria_gladiola  
     +--+                            
        |  +-- Anodontini            
        +--+                         
           +-- Unionini              

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 26 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 26 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini   

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 33 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Lanceolariini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 27 4 --
  +----------- Lepidodesmini
--+                         
  |  +-------- Lanceolariini
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticosta **   
        +--+                
           +-- Anodontini   

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 11 3 --
  +-------- Anodontini   
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Lanceolariini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 32 4 --
  +-------- Unionini     
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Lanceolariini
  +--+                   
     |  +-- Acuticosta **   
     +--+                
        +-- Anodontini   

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 15 2 --
           +----- Aculamprotula  
  +--------+                     
  |        |  +-- Acuticosta **     
  |        +--+                  
--+           +-- Lanceolariini  
  |                              
  |  +----------- Unio           
  |  |                           
  +--+  +-------- Pseudocuneopsis
     |  |                        
     +--+     +-- Schistodesmus  
        |  +--+                  
        |  |  +-- Cuneopsis      
        +--+                     
           |  +-- Tchangsinaia   
           +--+                  
              +-- Nodularia      

Page updated: 08:34:10 Wed 06 Nov 2024

 
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