|
Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae tribe LEPIDODESMINI Huang & Wu in Huang et al. 2019: cladomics
1 genus
Cladomics Summary
number of species = 3
number of subtaxa (rank: genus) = 1
analyzed in 52 cladograms in 29 publications
maximum number of terminals in an analysis = 2, mean = 1.1
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 2, mean = 2.0
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
119 |
68 |
4 |
-- |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+----- Acuticostini
|
+----- Lepidodesma **
--+
+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
94 |
45 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
90 |
45 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2023 [Publ 4339], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
17 |
9 |
1 |
-- |
+-- Lepidodesma **
--+
+-- Anodontini
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
90 |
45 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
94 |
45 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
73 |
35 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
52 |
16 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2024 [Publ 4386], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
25 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
90 |
25 |
4 |
-- |
+----------- Acuticostini
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Anodontini
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
14 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
23 |
23 |
3 |
-- |
+-------- Anodontini
--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Unionini
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
21 |
3 |
-- |
+-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2022 [Publ 4238], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
38 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
32 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
197 |
43 |
4 |
-- |
+----- Acuticostini
|
+----- Alasmidonta
|
+----- Anodonta_anatina
|
+----- Anemina
|
+----- Anodonta_cygnea
|
+----- Cristaria
|
+----- Anodonta_nuttalliana
|
+----- Lasmigona
|
+----- Lepidodesma **
|
+----- Pseudanodonta
|
+----- Pyganodon
|
--+----- Simpsonaias
|
+----- Strophitus
|
+----- Schistodesmus
|
+----- Utterbackia
|
+----- Lanceolariini
|
+----- Unio
|
+----- Aculamprotula
|
| +-- Pletholophus
+--+
| +-- Sinanodonta
|
| +-- Cuneopsis_heudei
| |
| +-- Cuneopsis_celtiformis
| |
+--+-- Tchangsinaia
|
+-- Cuneopsis_rufescens
|
+-- Nodularia
Huang et al., 2019 [Publ 3843], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
41 |
27 |
4 |
-- |
+-------------------------------- Anemina
|
--+ +----------------------------- Acuticostini
| |
+--+ +-------------------------- Sinanodonta
| |
+--+ +----------------------- Aculamprotula
| |
+--+ +-- Cristaria
| +-----------------+
| | +-- Lanceolariini
+--+
| +----------------- Lepidodesma **
+--+
| +-------------- Schistodesmus
+--+
| +----------- Nodularia
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Tchangsinaia
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
49 |
27 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
13 |
4 |
-- |
+----------- Anodontini
|
| +----- Acuticostini
--+ +--+
| | | +-- Nodularia
| | +--+
+--+ +-- Tchangsinaia
|
| +----- Lanceolariini
+--+
| +-- Schistodesmus
+--+
+-- Lepidodesma **
Dai et al., 2024 [Publ 4386], fig. 5-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
15 |
4 |
-- |
+----------------- Acuticostini
|
+----------------- Cristariina
|
+----------------- Anodontina
--+
| +-------------- Lanceolariini
| |
| | +-- Aculamprotula
| +-----------+
+--+ +-- Lepidodesma **
|
| +----------- Unio
| |
+--+ +-------- Middendorffinaia
| |
+--+ +----- Cuneopsis
| |
+--+----- Nodularia
|
| +-- Inversiunio
+--+
+-- Schistodesmus
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
22 |
3 |
-- |
+-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2024 [Publ 4379], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
53 |
51 |
3 |
-- |
+-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
13 |
4 |
-- |
+-------- Strophitus
|
+-------- Schistodesmus
|
+-------- Lepidodesma **
|
+-------- Acuticostini
|
+-------- Cristariina
--+
+-------- Lanceolariini
|
| +-- Nodularia
+-----+
| +-- Tchangsinaia
|
| +----- Utterbackia
| |
+--+----- Utterbackiana
|
| +-- Alasmidonta
+--+
+-- Lasmigona
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
6 |
2 |
-- |
+-------- Anodontini
--+
| +----- Aculamprotula
+--+
| +-- Lepidodesma **
+--+
+-- Unio
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
197 |
43 |
4 |
-- |
+-------- Acuticostini
|
--+ +----- Lepidodesma **
| |
+--+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Dai et al., 2024 [Publ 4386], fig. 5-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
15 |
4 |
-- |
+----------- Anodontina
|
--+ +-- Acuticostini
| +-----+
| | +-- Cristariina
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
195 |
42 |
4 |
-- |
+----------- Acuticostini
--+
| +-------- Unionini
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
26 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
40 |
26 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Sano et al., 2020 [Publ 4138], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
21 |
4 |
-- |
+----- Lanceolaria_grayii__1
|
+----- Lanceolaria_grayii__2
|
+----- Cristaria_plicata__1
|
+----- Cristaria_plicata__2
|
+----- Sinanodonta_lauta
|
+----- Beringiana_japonica
|
+----- Beringiana_fukuharai__1
|
+----- Beringiana_beringiana
|
+----- Beringiana_fukuharai__2
|
+----- Beringiana_fukuharai__3
|
+----- Buldowskia_iwakawai__1
|
+----- Buldowskia_iwakawai__2
|
+----- Sinanodonta_tumens__1
--+
+----- Sinanodonta_tumens__2
|
+----- Pletholophus
|
+----- Anemina
|
+----- Lanceolaria_lanceolata
|
+----- Cristaria_plicata__3
|
+----- Lanceolaria_gladiola
|
+----- Sinanodonta_angula
|
+----- Lepidodesma **
|
| +-- Aculamprotula
+--+
| +-- Acuticostini
|
| +-- Cuneopsis
+--+
| +-- Nodularia
|
| +-- Schistodesmus
+--+
+-- Inversiunio
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
16 |
4 |
-- |
+----------------- Anodontini
|
| +-- Acuticostini
--+ +-----------+
| | +-- Lanceolariini
| |
+--+ +-- Lepidodesma **
| +--------+
| | +-- Aculamprotula
+--+
| +-------- Unio
+--+
| +----- Schistodesmus
+--+
| +-- Nodularia
+--+
+-- Tchangsinaia
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
35 |
3 |
-- |
+-- Buldowskia
+--------------+
| +-- Anemina
|
--+ +----- Pletholophus
| +--------+
| | | +-- Beringiana
| | +--+
+--+ +-- Sinanodonta
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Unionini
Sano et al., 2017 [Publ 3799], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
32 |
20 |
4 |
-- |
+-------------- Anodontini
|
| +-- Tchangsinaia
--+ +--------+
| | +-- Nodularia
| |
+--+ +-- Schistodesmus
| +-----+
| | +-- Acuticostini
+--+
| +----- Inversiunio
+--+
| +-- Lepidodesma **
+--+
+-- Lanceolariini
Huang et al., 2019 [Publ 3843], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
30 |
4 |
-- |
+-------------- Anemina
|
--+ +----------- Sinanodonta
| |
+--+ +-------- Acuticostini
| |
+--+ +-- Lepidodesma **
| +--+
| | +-- Cristaria
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
35 |
3 |
-- |
+-- Buldowskia
+--------------+
| +-- Anemina
|
--+ +----- Sinanodonta
| +--------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lanceolariini
+--+
| +-- Lepidodesma **
+--+
+-- Unionini
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
22 |
3 |
-- |
+-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
90 |
25 |
4 |
-- |
+-------- Acuticostini
|
--+ +-- Lepidodesma **
| +--+
| | +-- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Kim et al., 2021 [Publ 4197], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
15 |
3 |
-- |
-- Lepidodesma_languilati
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
52 |
32 |
4 |
-- |
+-------- Lepidodesma **
|
--+-------- Unionini
|
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
8 |
2 |
-- |
+----- Anodontini
--+
| +-- Lepidodesma **
+--+
+-- Unionini
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
30 |
4 |
-- |
+-- Anemina
+--------+
| +-- Amuranodonta
|
| +----- Pletholophus
--+ +--+
| | | +-- Sinanodonta
| | +--+
| | +-- Beringiana
| |
+--+ +----- Lanceolaria_gladiola
| |
| +----- Acuticostini
| |
| +----- Lepidodesma **
| |
+--+----- Lanceolaria_oxyrhyncha
|
+----- Cristaria
|
+----- Unionini
|
| +-- Alasmidontina
+--+
+-- Anodontina
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
73 |
33 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2024 [Publ 4386], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
11 |
2 |
-- |
+-------------- Unio
|
| +----- Aculamprotula
--+ +-----+
| | | +-- Anodontini
| | +--+
+--+ +-- Lepidodesma **
|
| +-------- Middendorffinaia
+--+
| +----- Inversiunio
+--+
| +-- Cuneopsis
+--+
+-- Nodularia
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
30 |
4 |
-- |
+-------- Acuticostini
|
--+ +----- Lepidodesma **
| |
+--+----- Unionini
|
| +-- Lanceolariini
+--+
+-- Anodontini
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
48 |
22 |
3 |
-- |
+-------- Lepidodesma **
--+
| +----- Unionini
+--+
| +-- Lanceolariini
+--+
+-- Anodontini
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
26 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
51 |
26 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
73 |
33 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Unionini
+--+
| +----- Anodontini
+--+
| +-- Acuticostini
+--+
+-- Lanceolariini
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
14 |
2 |
-- |
+-------- Anodontini
--+
| +----- Schistodesmus
+--+
| +-- Lepidodesma **
+--+
+-- Aculamprotula
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
35 |
3 |
-- |
+-- Buldowskia
+--------------+
| +-- Anemina
|
--+ +----- Sinanodonta
| +--------+
| | | +-- Pletholophus
| | +--+
+--+ +-- Beringiana
|
| +----------- Cristaria
| |
+--+ +-- Anodontina
| +-----+
| | +-- Alasmidontina
+--+
| +----- Lepidodesma **
+--+
| +-- Lanceolariini
+--+
+-- Unionini
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
53 |
27 |
4 |
-- |
+----------- Lepidodesma **
--+
| +-------- Lanceolariini
+--+
| +----- Unionini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
55 |
32 |
4 |
-- |
+-------- Unionini
|
--+ +-- Lepidodesma **
| +--+
| | +-- Lanceolariini
+--+
| +-- Acuticostini
+--+
+-- Anodontini
| 
