MUSSELp
 
MUSSELpdb
 
 
Taxonomy
 
 
Geography
 
 
 

Page last updated
Fri 05 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Chen et al. (2025), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/fmuotwaolcb/db.php?ty=publ&id=4448

Search the database pages:

Publication information

Chen, H., H.-Q. Xiang, Y.-T. Dai, Z.-G. Chen, Y.-M. He et al. 2025. A new genus and two new species of freshwater mussels (Bivalvia, Unionidae) from Sichuan, China: Overlooked cryptic endemism in the upper Yangtze River Basin. Zoosystematics and Evolution 101(4): 1459-1470.

Click here for online access.

Attributed Genus Names

Tongchuania Chen et al., 2025
Chen Xiang, He, Huang & Wu in Chen et al., 2025, Zoosyst. Evol. 101(4): 1464.
Type species: Tongchuania mingiana Chen et al., 2025 (orig. desig.) = Tongchuania mingiana

Attributed Species Names

Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
Chen et al., 2025, Zoosyst. Evol. 101(4): Tables S2-S3.
Dai et al., 2025, Zoosyst. Evol. 101(2): Tables 1-2.
(misidentified reference to Unio crassus Retzius, 1788)

Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
Dai et al., 2024, Zoosyst. Evol. 100(4): Table 2.
Chen et al., 2025, Zoosyst. Evol. 101(4): Table S3.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1.
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 2.
(misidentified reference to Unio hainesianus Lea, 1856)

Cuneopsis heudei Lopes-Lima et al., 2017
‘Heude’ of Authors.
Liu et al., 2017, PLoS One 12(12 ), e0189737: Table S4.
Lopes-Lima et al., 2017, MPE 106: Table 2.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Wu et al., 2018, PLoS One 13(8), e0200956: 5.
Huang et al., 2019, MPE 130: Tables S1, S4. [in part]
Lopes-Lima et al., 2020, MPE 146: Table 1. [in part]
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1. [in part]
Wu et al., 2022, Zool. Scripta 51: Table S1. [in part]
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, ZooKeys 1166: Table 1.
Liu et al., 2023, ZooKeys 1179: Table 1.
Chen et al., 2025, Zoosyst. Evol. 101(4): Table S2. [in part]
Dai et al., 2025, Zoosyst. Evol. 101(2): Table 1. [in part]
(misidentified reference to Unio heudei Heude, 1874)

Nodularia douglasiae Choi et al., 2020
‘Griffith & Pidgeon’ of Authors.
Cha et al., 2018, mtDNA B 3(2): 946. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3488. [in part]
Choi et al., 2020, Sci. Repts. 10: 5 [as ‘Nodularia douglasiae Clade A’]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 57, Table S2. [in part]
Mabuchi et al., 2021, mtDNA B 6(4): 1536. [in part]
Kim et al., 2021, mtDNA B 6(1): 80. [in part]
Lopes-Lima et al., 2024, MPE 195: Table S2. [in part]
Dai et al., 2024, Invert. Syst. 38(6): fig. 3, Table S1. [in part]
Wu et al., 2024, Invert. Syst. 38, IS24014: Table S1. [in part]
Chen et al., 2025, Zoosyst. Evol. 101(4): Table S2. [in part]
(misidentified reference to Unio douglasiae Griffith & Pidgeon, 1833)

Tongchuania mingiana Chen et al., 2025
Chen, Xiang, He, Huang & Wu in Chen et al., 2025, Zoosyst. Evol. 101(4): 1464, figs. 4-5.

Pseudocuneopsis perflora Chen et al., 2025
Chen, Xiang, He, Huang & Wu in Chen et al., 2025, Zoosyst. Evol. 101(4): 1464, fig. 6.
Type(s): HOLOTYPE (ORIG. DESIG.) NCUPP_250521, Bahe River, Dazhou City, Sichuan Province, China, 31.0978°N, 107.1856°E.

Taxonomic Opinions

Aculamprotula coreana (Martens, 1886)

Aculamprotula polysticta (Heude, 1877)

Aculamprotula scripta (Heude, 1875)

Aculamprotula tientsinensis (Crosse & Debeaux, 1863)

Aculamprotula tortuosa (Lea, 1865)

Acuticosta chinensis (Lea, 1868)

Amblema plicata (Say, 1817)

Anemina arcaeformis (Heude, 1877)

Anodonta anatina (Linnaeus, 1758)

Anodonta cygnea (Linnaeus, 1758)

Anodontites trapesialis (Lamarck, 1819)

Chamberlainia hainesiana (Lea, 1856)

Cristaria plicata (Leach, 1814)

Cumberlandia monodonta (Say, 1829)

Cuneopsis celtiformis (Heude, 1874)

Cuneopsis demangei Haas, 1929

Cuneopsis heudei (Heude, 1874)

Cuneopsis kiangsiensis Tchang & Li, 1965

Cuneopsis rufescens (Heude, 1874)

Cuneopsis szechenyii (Neumayr, 1899)

Diaurora aurorea

Echyridella menziesii (Gray, 1843)

Elliptio complanata (Lightfoot, 1786)

Gibbosula rochechouartii (Heude, 1875)

Hyriopsis bialata Simpson, 1900

Lamprotula caveata (Heude, 1877)

Lamprotula leaii (Griffith & Pidgeon, 1833)

Lampsilis cardium Rafinesque, 1820

Lampsilis ornata (Conrad, 1835)

Lampsilis powellii (Lea, 1852)

Lampsilis siliquoidea (Barnes, 1823)

Lanceolaria gladiola (Heude, 1877)

Lanceolaria grayii (J.E. Gray, 1833)

Lanceolaria lanceolata (I. Lea, 1856)

Lasmigona compressa (Lea, 1829)

Leaunio lienosus (Conrad, 1834)

Lens contradens (Lea, 1838)

Lepidodesma languilati (Heude, 1874)

Leptodea leptodon (Rafinesque, 1820)

Margaritifera dahurica (Middendorff, 1850)

Margaritifera falcata (Gould, 1850)

Margaritifera margaritifera (Linnaeus, 1758)

Microcondylaea bonellii (Férussac, 1827)

Middendorffinaia mongolica

Mutela dubia (Gmelin, 1791)

Nodularia breviconcha Lee, Kim, Bogan & Kondo, 2020

Nodularia douglasiae (Gray, 1833)

Nodularia nipponensis (von Martens, 1877)

Parvasolenaia rivularis (Heude, 1877)

Physunio superbus (Lea, 1843)

Pilsbryoconcha exilis (Lea, 1838)

Pleurobema oviforme (Conrad, 1834)

Popenaias popeii (Lea, 1857)

Potamilus alatus (Say, 1817)

Potomida littoralis (Cuvier, 1798)

Pronodularia japanensis (Lea, 1859)

Pseudocuneopsis capitata (Heude, 1874)

Pseudocuneopsis perflora H. Chen H.-Q. Xiang, Y.-M. He, X.-C. Huang & X.-P. Wu, sp. nov.

Pseudocuneopsis sichuanensis Huang, Dai & Wu, 2022

Pseudocuneopsis wuana Liu & Wu, 2023

Pseudocuneopsis yangshuoensis Wu & Liu, 2023

Pseudocuneopsis yemaoi Dai, Chen, Peng, Ouyang, Huang & Wu, 2024

Pseudodon vondembuschianus (Lea, 1840)

Pseudunio marocanus (Pallary, 1918)

Ptychorhynchus pfisteri (Heude, 1874)

Pyganodon grandis (Say, 1829)

Quadrula quadrula (Rafinesque, 1820)

Rectidens sumatrensis (Dunker, 1852)

Schistodesmus lampreyanus (Baird & Adams, 1867)

Schistodesmus spinosus Simpson, 1900
+ Lamprotula gottschei He et al., 2016

Sinanodonta lucida (Heude, 1877)

Sinanodonta tumens (Haas, 1910)

Sinanodonta woodiana (Lea, 1834)

Sinohyriopsis cumingii (Lea, 1852)

Sinohyriopsis schlegelii (Martens, 1861)

Sinosolenaia carinata (Heude, 1877)

Sinosolenaia oleivora (Heude, 1877)

Tchangsinaia piscicula (Heude, 1874)

Tongchuania mingiana H. Chen H.-Q. Xiang, Y.-M. He, X.-C. Huang & X.-P. Wu, gen. et. sp. nov.

Toxolasma parvum (Barnes, 1823)

Unio crassus Philipsson, 1788

Unio delphinus Spengler, 1793

Unio pictorum (Linnaeus, 1758)

Unio tumidus Philipsson, 1788

Uniomerus tetralasmus (Say, 1831)

Utterbackia imbecillis (Say, 1829)

Utterbackia peninsularis Bogan & Hoeh, 1995

Venustaconcha ellipsiformis (Conrad, 1836)

Page updated: 12:47:53 Fri 05 Sep 2025

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."