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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Nodularia Conrad, 1853: cladomics
     14 valid species

Cladomics Summary

number of species = 14

analyzed in 106 cladograms in 66 publications
maximum number of terminals in an analysis = 9, mean = 2.24

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 55
maximum number of included terminals = 9, mean = 3.38
proportion of cladograms that support basic monophyly = 0.98


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
9 9 8 10 45 20 6 0.0

*** Top Monophyly Cladogram ***

  +----------------------------- Aculamprotula          
  |                                                     
--+  +-------------------------- Unio                   
  |  |                                                  
  +--+  +----------------------- Pseudocuneopsis        
     |  |                                               
     |  |                 +----- Schistodesmus          
     +--+  +--------------+                             
        |  |              |  +-- Tchangsinaia           
        |  |              +--+                          
        |  |                 +-- Cuneopsis              
        +--+                                            
           |              +----- Nodularia_dorri **        
           |  +-----------+                             
           |  |           |  +-- Nodularia_hanensis **     
           |  |           +--+                          
           +--+              +-- Nodularia_micheloti **    
              |                                         
              |  +-------------- Nodularia_breviconcha **  
              |  |                                      
              +--+  +----------- Nodularia_huana **        
                 |  |                                   
                 +--+  +-------- Nodularia_fusiformans **  
                    |  |                                
                    +--+     +-- Nodularia_douglasiae__1 **
                       |  +--+                          
                       |  |  +-- Nodularia_dualobtusa **   
                       +--+                             
                          |  +-- Nodularia_douglasiae__2 **
                          +--+                          
                             +-- Nodularia_nuxpersicae **  

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --

*** Top Sister-Group Cladogram ***

  +----------------- Aculamprotula        
  |                                       
  |           +----- Middendorffinaia     
--+  +--------+                           
  |  |        |  +-- Nodularia_douglasiae ** 
  |  |        +--+                        
  |  |           +-- Nodularia_breviconcha **
  +--+                                    
     |  +----------- Cuneopsis            
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio                 

Wu et al., 2024 [Publ 4379], fig. 4_6

iT tT bi riS oT roT roS iD
9 9 8 10 13 5 2 1.0
  +-------------------- Schistodesmus          
  |                                            
  |              +----- Nodularia_dorri **        
--+  +-----------+                             
  |  |           |  +-- Nodularia_micheloti **    
  |  |           +--+                          
  +--+              +-- Nodularia_hanensis **     
     |                                         
     |  +-------------- Nodularia_breviconcha **  
     |  |                                      
     +--+  +----------- Nodularia_huana **        
        |  |                                   
        +--+  +-------- Nodularia_fusiformans **  
           |  |                                
           +--+     +-- Nodularia_douglasiae__1 **
              |  +--+                          
              |  |  +-- Nodularia_dualobtusa **   
              +--+                             
                 |  +-- Nodularia_nuxpersicae **  
                 +--+                          
                    +-- Nodularia_douglasiae__2 **

Wu et al., 2024 [Publ 4379], fig. 2

iT tT bi riS oT roT roS iD
9 9 8 10 3 3 3 3.0
                 +-- Cuneopsis              
                 |                          
  +--------------+-- Tchangsinaia           
  |              |                          
  |              +-- Pseudocuneopsis        
  |                                         
--+           +----- Nodularia_dorri **        
  |  +--------+                             
  |  |        |  +-- Nodularia_hanensis **     
  |  |        +--+                          
  |  |           +-- Nodularia_micheloti **    
  +--+                                      
     |  +----------- Nodularia_breviconcha **  
     |  |                                   
     |  |        +-- Nodularia_fusiformans **  
     +--+  +-----+                          
        |  |     +-- Nodularia_huana **        
        |  |                                
        +--+     +-- Nodularia_douglasiae__1 **
           |  +--+                          
           |  |  +-- Nodularia_dualobtusa **   
           +--+                             
              |  +-- Nodularia_nuxpersicae **  
              +--+                          
                 +-- Nodularia_douglasiae__2 **

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iT tT bi riS oT roT roS iD
7 7 5 7 3 1 1 7.0
  +-------------------- Middendorffinaia     
--+                                          
  |  +----------------- Nodularia_breviconcha **
  +--+                                       
     |  +-------------- Nodularia_fusiformans **
     +--+                                    
        |  +----------- Nodularia_huana **      
        +--+                                 
           |  +-------- Nodularia_nuxpersicae **
           +--+                              
              |  +----- Nodularia_nipponensis **
              +--+                           
                 |  +-- Nodularia_douglasiae ** 
                 +--+                        
                    +-- Nodularia_dualobtusa ** 

Choi et al., 2020 [Publ 4169], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 7 3 0 0 7.0
  +----------- Nodularia_breviconcha **
  |                                 
  |        +-- Nodularia_fusiformans **
--+  +-----+                        
  |  |     +-- Nodularia_huana **      
  |  |                              
  +--+     +-- Nodularia_dualobtusa ** 
     |  +--+                        
     |  |  +-- Nodularia_douglasiae ** 
     +--+                           
        |  +-- Nodularia_nipponensis **
        +--+                        
           +-- Nodularia_nuxpersicae **

Choi et al., 2020 [Publ 4169], fig. 2a

iT tT bi riS oT roT roS iD
7 7 6 8 5 5 4 6.0
              +-------- Schistodesmus          
  +-----------+                                
  |           |  +----- Cuneopsis              
  |           +--+                             
  |              |  +-- Unio                   
--+              +--+                          
  |                 +-- Tchangsinaia           
  |                                            
  |  +----------------- Nodularia_breviconcha **  
  |  |                                         
  +--+  +-------------- Nodularia_huana **        
     |  |                                      
     +--+  +----------- Nodularia_fusiformans **  
        |  |                                   
        +--+        +-- Nodularia_nipponensis **  
           |  +-----+                          
           |  |     +-- Nodularia_nuxpersicae **  
           +--+                                
              |  +----- Nodularia_dualobtusa__2 **
              +--+                             
                 |  +-- Nodularia_douglasiae **   
                 +--+                          
                    +-- Nodularia_dualobtusa__1 **

Sayenko et al., 2021 [Publ 4189], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 6 7 7 3 6.0
  +----------------------- Unio                 
--+                                             
  |  +-------------------- Inversiunio          
  +--+                                          
     |  +----------------- Middendorffinaia     
     +--+                                       
        |  +-------------- Nodularia_breviconcha **
        +--+                                    
           |  +----------- Nodularia_huana **      
           +--+                                 
              |  +-------- Nodularia_nuxpersicae **
              +--+                              
                 |  +----- Nodularia_nipponensis **
                 +--+                           
                    |  +-- Nodularia_douglasiae ** 
                    +--+                        
                       +-- Nodularia_dualobtusa ** 

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
6 6 4 6 114 20 7 6.0
  +-------------------------- Aculamprotula        
  |                                                
--+  +----------------------- Middendorffinaia     
  |  |                                             
  +--+  +-------------------- Unio                 
     |  |                                          
     |  |              +----- Schistodesmus        
     +--+              |                           
        |  +-----------+----- Cuneopsis            
        |  |           |                           
        |  |           |  +-- Tchangsinaia         
        +--+           +--+                        
           |              +-- Inversiunio          
           |                                       
           |  +-------------- Nodularia_breviconcha **
           +--+                                    
              |  +----------- Nodularia_huana **      
              +--+                                 
                 |  +-------- Nodularia_nuxpersicae **
                 +--+                              
                    |  +----- Nodularia_nipponensis **
                    +--+                           
                       |  +-- Nodularia_douglasiae ** 
                       +--+                        
                          +-- Nodularia_dualobtusa ** 

Dai et al., 2024 [Publ 4386], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 8 5 5 7.0
  +----------------- Middendorffinaia     
  |                                       
--+  +-------------- Inversiunio          
  |  |                                    
  +--+  +----------- Cuneopsis            
     |  |                                 
     +--+  +-------- Nodularia_breviconcha **
        |  |                              
        +--+     +-- Nodularia_douglasiae ** 
           |  +--+                        
           |  |  +-- Nodularia_nuxpersicae **
           +--+                           
              |  +-- Nodularia_fusiformans **
              +--+                        
                 +-- Nodularia_huana **      

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
5 5 3 6 16 6 6 10.0
  +----------------- Aculamprotula          
  |                                         
--+  +-------------- Unio                   
  |  |                                      
  |  |  +----------- Middendorffinaia       
  +--+  |                                   
     |  |     +----- Cuneopsis              
     |  |  +--+                             
     +--+  |  |  +-- Inversiunio            
        |  |  +--+                          
        |  |     +-- Schistodesmus          
        |  |                                
        +--+     +-- Nodularia_nuxpersicae **  
           |  +--+                          
           |  |  +-- Nodularia_douglasiae__1 **
           |  |                             
           |  |  +-- Nodularia_huana **        
           +--+--+                          
              |  +-- Nodularia_breviconcha **  
              |                             
              |  +-- Nodularia_douglasiae__2 **
              +--+                          
                 +-- Nodularia_dualobtusa **   

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
5 5 4 6 16 6 6 9.0
  +----------------- Unio                   
  |                                         
--+  +-------------- Middendorffinaia       
  |  |                                      
  |  |  +----------- Cuneopsis              
  +--+  |                                   
     |  |        +-- Inversiunio            
     |  +--------+                          
     |  |        +-- Schistodesmus          
     +--+                                   
        |        +-- Nodularia_huana **        
        |  +-----+                          
        |  |     +-- Nodularia_breviconcha **  
        |  |                                
        +--+     +-- Nodularia_nuxpersicae **  
           |  +--+                          
           |  |  +-- Nodularia_douglasiae__1 **
           +--+                             
              |  +-- Nodularia_douglasiae__2 **
              +--+                          
                 +-- Nodularia_dualobtusa **   

Mabuchi et al., 2021 [Publ 4233], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 1 1 1 10.0
  +-------- Tchangsinaia         
  |                              
--+     +-- Nodularia **            
  |  +--+                        
  |  |  +-- Nodularia_nipponensis **
  +--+                           
     |  +-- Nodularia_fusiformans **
     +--+                        
        +-- Nodularia_breviconcha **

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 13 7 4 8.0
  +-------------- Unio                 
  |                                    
--+           +-- Schistodesmus        
  |  +--------+                        
  |  |        +-- Cuneopsis            
  +--+                                 
     |  +-------- Nodularia_breviconcha **
     +--+                              
        |  +----- Nodularia_nipponensis **
        +--+                           
           |  +-- Nodularia_dualobtusa ** 
           +--+                        
              +-- Nodularia_douglasiae ** 

Klishko et al., 2018 [Publ 3945], fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 4 4 1 8.0
  +----------- Unio                 
--+                                 
  |  +-------- Nodularia_breviconcha **
  +--+                              
     |  +----- Nodularia_huana **      
     +--+                           
        |  +-- Nodularia_douglasiae ** 
        +--+                        
           +-- Nodularia_nipponensis **

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
4 4 3 5 36 16 6 6.0
  +----------------------- Aculamprotula          
  |                                               
--+  +-------------------- Unio                   
  |  |                                            
  +--+  +----------------- Pseudocuneopsis        
     |  |                                         
     |  |           +----- Schistodesmus          
     +--+  +--------+                             
        |  |        |  +-- Tchangsinaia           
        |  |        +--+                          
        +--+           +-- Cuneopsis              
           |                                      
           |  +----------- Nodularia_breviconcha **  
           +--+                                   
              |  +-------- Nodularia_fusiformans **  
              +--+                                
                 |  +----- Nodularia_douglasiae__2 **
                 +--+                             
                    |  +-- Nodularia_nuxpersicae **  
                    +--+                          
                       +-- Nodularia_douglasiae__1 **

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
4 4 3 6 11 7 3 8.0
  +----------------- Unio                   
  |                                         
--+  +-------------- Schistodesmus          
  |  |                                      
  |  +-------------- Cuneopsis              
  +--+                                      
     |  +----------- Nodularia_breviconcha **  
     |  |                                   
     +--+  +-------- Nodularia_douglasiae__3 **
        |  |                                
        +--+  +----- Nodularia_douglasiae__2 **
           |  |                             
           +--+  +-- Nodularia_douglasiae__1 **
              |  |                          
              +--+-- Nodularia_nuxpersicae **  
                 |                          
                 +-- Nodularia_nipponensis **  

Choi et al., 2020 [Publ 4169], fig. 3a

iT tT bi riS oT roT roS iD
4 4 3 8 2 2 2 8.0
              +-- Cuneopsis               
  +-----------+                           
  |           +-- Unio                    
--+                                       
  |  +----------- Nodularia_breviconcha **   
  |  |                                    
  +--+  +-------- Nodularia_nipponensis__3 **
     |  |                                 
     +--+  +----- Nodularia_douglasiae__3 ** 
        |  |                              
        |  |  +-- Nodularia_dualobtusa **    
        +--+  |                           
           |  +-- Nodularia_nipponensis__1 **
           |  |                           
           +--+-- Nodularia_nipponensis__2 **
              |                           
              +-- Nodularia_douglasiae__1 ** 
              |                           
              +-- Nodularia_douglasiae__2 ** 

Wu et al., 2024 [Publ 4388], fig. 4

iT tT bi riS oT roT roS iD
4 4 3 5 5 0 0 6.0
  +-------- Nodularia_breviconcha **  
  |                                
--+     +-- Nodularia_douglasiae__1 **
  |  +--+                          
  |  |  +-- Nodularia_dualobtusa **   
  +--+                             
     |  +-- Nodularia_douglasiae__2 **
     +--+                          
        +-- Nodularia_nuxpersicae **  

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
3 3 2 4 30 5 3 9.0
  +----------- Tchangsinaia            
--+                                    
  |  +-------- Nodularia_nipponensis__1 **
  +--+                                 
     |  +----- Nodularia_breviconcha **   
     +--+                              
        |  +-- Nodularia_nipponensis__2 **
        +--+                           
           +-- Nodularia_douglasiae **    

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 193 14 5 9.0
  +-------------- Aculamprotula        
  |                                    
--+  +----------- Unio                 
  |  |                                 
  |  |     +----- Nodularia_douglasiae ** 
  +--+  +--+                           
     |  |  |  +-- Nodularia_huana **      
     |  |  +--+                        
     +--+     +-- Nodularia_nuxpersicae **
        |                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Tchangsinaia         
           +--+                        
              +-- Cuneopsis            

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 4 7.0
  +----------- Aculamprotula        
  |                                 
  |     +----- Nodularia_huana **      
--+  +--+                           
  |  |  |  +-- Nodularia_nuxpersicae **
  |  |  +--+                        
  +--+     +-- Nodularia_douglasiae ** 
     |                              
     |  +----- Tchangsinaia         
     +--+                           
        |  +-- Schistodesmus        
        +--+                        
           +-- Cuneopsis            

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 13 9 6 9.0
  +----------------- Unio                 
  |                                       
--+  +-------------- Pseudocuneopsis      
  |  |                                    
  +--+           +-- Schistodesmus        
     |  +--------+                        
     |  |        +-- Cuneopsis            
     +--+                                 
        |  +-------- Tchangsinaia         
        +--+                              
           |  +----- Nodularia_nipponensis **
           +--+                           
              |  +-- Nodularia_douglasiae ** 
              +--+                        
                 +-- Nodularia_dualobtusa ** 

Klishko et al., 2019 [Publ 3886], fig. 9

iT tT bi riS oT roT roS iD
3 3 1 3 8 5 2 7.0
  +----------- Unio                 
--+                                 
  |  +-------- Middendorffinaia     
  +--+                              
     |  +----- Nodularia_breviconcha **
     +--+                           
        |  +-- Nodularia_douglasiae ** 
        +--+                        
           +-- Nodularia_huana **      

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 51 6 6 9.0
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Pseudocuneopsis      
     |  |                                 
     |  |     +----- Nodularia_breviconcha **
     +--+  +--+                           
        |  |  |  +-- Nodularia_douglasiae ** 
        |  |  +--+                        
        +--+     +-- Nodularia_nipponensis **
           |                              
           |  +----- Tchangsinaia         
           +--+                           
              |  +-- Cuneopsis            
              +--+                        
                 +-- Schistodesmus        

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 195 15 5 7.0
  +-------- Cuneopsis_heudei     
  |                              
  +-------- Cuneopsis_celtiformis
  |                              
--+-------- Tchangsinaia         
  |                              
  +-------- Cuneopsis_rufescens  
  |                              
  |  +----- Nodularia_huana **      
  +--+                           
     |  +-- Nodularia_douglasiae ** 
     +--+                        
        +-- Nodularia_nuxpersicae **

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 39 11 4 9.0
  +-------------- Schistodesmus        
  |                                    
  |        +----- Nodularia_douglasiae ** 
--+  +-----+                           
  |  |     |  +-- Nodularia_huana **      
  |  |     +--+                        
  +--+        +-- Nodularia_nuxpersicae **
     |                                 
     |  +-------- Cuneopsis_rufescens  
     +--+                              
        |  +----- Tchangsinaia         
        +--+                           
           |  +-- Cuneopsis_heudei     
           +--+                        
              +-- Cuneopsis_celtiformis

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 195 15 5 9.0
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Schistodesmus        
     |  |                                 
     +--+        +-- Tchangsinaia         
        |  +-----+                        
        |  |     +-- Cuneopsis            
        +--+                              
           |  +----- Nodularia_douglasiae ** 
           +--+                           
              |  +-- Nodularia_huana **      
              +--+                        
                 +-- Nodularia_nuxpersicae **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 20 6 --
  +-------------- Aculamprotula        
  |                                    
--+  +----------- Unio                 
  |  |                                 
  +--+  +-------- Pseudocuneopsis      
     |  |                              
     |  |     +-- Nodularia_breviconcha **
     +--+-----+                        
        |     +-- Nodularia_douglasiae ** 
        |                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Tchangsinaia         
           +--+                        
              +-- Cuneopsis            

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 37 21 7 --
  +-------------------- Aculamprotula        
  |                                          
--+  +----------------- Unio                 
  |  |                                       
  +--+  +-------------- Pseudocuneopsis      
     |  |                                    
     +--+           +-- Nodularia_breviconcha **
        |  +--------+                        
        |  |        +-- Nodularia_douglasiae ** 
        +--+                                 
           |  +-------- Schistodesmus        
           +--+                              
              |  +----- Cuneopsis            
              +--+                           
                 |  +-- Tchangsinaia         
                 +--+                        
                    +-- Inversiunio          

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 2 72 13 6 --
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Pseudocuneopsis      
     |  |                                 
     +--+        +-- Nodularia_douglasiae ** 
        |  +-----+                        
        |  |     +-- Nodularia_breviconcha **
        +--+                              
           |  +----- Schistodesmus        
           +--+                           
              |  +-- Cuneopsis            
              +--+                        
                 +-- Tchangsinaia         

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 4 67 11 5 --
  +-------------------- Aculamprotula           
  |                                             
--+  +----------------- Unio                    
  |  |                                          
  +--+  +-------------- Inversiunio             
     |  |                                       
     +--+           +-- Schistodesmus           
        |  +--------+                           
        |  |        +-- Tchangsinaia            
        +--+                                    
           |  +-------- Nodularia_nipponensis__1 **
           +--+                                 
              |  +----- Nodularia_nipponensis__2 **
              +--+                              
                 |  +-- Nodularia_nipponensis__3 **
                 +--+                           
                    +-- Nodularia_douglasiae **    

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 3 32 5 4 --
  +----- Cuneopsis               
  |                              
--+  +-- Nodularia_nipponensis__1 **
  |  |                           
  +--+-- Nodularia_douglasiae **    
     |                           
     +-- Nodularia_nipponensis__2 **

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --
  +----------------- Aculamprotula        
  |                                       
  |           +----- Middendorffinaia     
--+  +--------+                           
  |  |        |  +-- Nodularia_douglasiae ** 
  |  |        +--+                        
  |  |           +-- Nodularia_breviconcha **
  +--+                                    
     |  +----------- Cuneopsis            
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio                 

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 2 78 9 5 --
  +-------------- Unio                 
  |                                    
--+  +----------- Aculamprotula        
  |  |                                 
  +--+        +-- Nodularia_nuxpersicae **
     |  +-----+                        
     |  |     +-- Nodularia_douglasiae ** 
     +--+                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Tchangsinaia         
           +--+                        
              +-- Cuneopsis            

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iT tT bi riS oT roT roS iD
2 2 0 2 10 0 0 --
  +-- Nodularia_breviconcha **
--+                        
  +-- Nodularia_douglasiae ** 

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

iT tT bi riS oT roT roS iD
2 2 0 2 6 4 1 --
  +----- Inversiunio          
--+                           
  |  +-- Nodularia_breviconcha **
  +--+                        
     +-- Nodularia_douglasiae ** 

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
2 2 0 2 74 8 5 --
  +----------------- Unio                 
  |                                       
--+  +-------------- Aculamprotula        
  |  |                                    
  +--+           +-- Nodularia_douglasiae ** 
     |  +--------+                        
     |  |        +-- Nodularia_nuxpersicae **
     +--+                                 
        |  +-------- Schistodesmus        
        +--+                              
           |  +----- Cuneopsis_heudei     
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Cuneopsis_rufescens  

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 18 6 --
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Pseudocuneopsis      
     |  |                                 
     +--+        +-- Nodularia_douglasiae ** 
        |  +-----+                        
        |  |     +-- Nodularia_breviconcha **
        +--+                              
           |  +----- Schistodesmus        
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Cuneopsis            

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 2 0 2 94 20 6 --
  +-------------------- Aculamprotula               
  |                                                 
--+  +----------------- Unio                        
  |  |                                              
  +--+  +-------------- Pseudocuneopsis             
     |  |                                           
     |  |           +-- Nodularia_breviconcha **       
     +--+  +--------+                               
        |  |        +-- Nodularia_nipponensis **       
        |  |                                        
        +--+        +-- Schistodesmus_lampreyanus__1
           |  +-----+                               
           |  |     +-- Cuneopsis                   
           +--+                                     
              |  +----- Tchangsinaia                
              +--+                                  
                 |  +-- Schistodesmus_lampreyanus__2
                 +--+                               
                    +-- Schistodesmus_spinosus      

Froufe et al., 2016 [Publ 3738], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 12 12 1 --
                             +-- Nodularia_douglasiae ** 
  +--------------------------+                        
  |                          +-- Nodularia_breviconcha **
  |                                                   
--+                          +-- Unio_gibbus          
  |  +-----------------------+                        
  |  |                       +-- Unio_durieui         
  +--+                                                
     |  +----------------------- Unio_tumidus         
     +--+                                             
        |  +-------------------- Unio_tumidiformis    
        +--+                                          
           |  +----------------- Unio_delphinus       
           +--+                                       
              |  +-------------- Unio_foucauldianus   
              +--+                                    
                 |  +----------- Unio_ravoisieri      
                 +--+                                 
                    |  +-------- Unio_mancus          
                    +--+                              
                       |  +----- Unio_elongatulus     
                       +--+                           
                          |  +-- Unio_pictorum        
                          +--+                        
                             +-- Unio_caffer          

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 4 67 11 5 --
  +----------------- Aculamprotula           
  |                                          
--+  +-------------- Unio                    
  |  |                                       
  |  |        +----- Inversiunio             
  +--+  +-----+                              
     |  |     |  +-- Schistodesmus           
     |  |     +--+                           
     +--+        +-- Tchangsinaia            
        |                                    
        |  +-------- Nodularia_nipponensis__3 **
        +--+                                 
           |  +----- Nodularia_nipponensis__1 **
           +--+                              
              |  +-- Nodularia_douglasiae **    
              +--+                           
                 +-- Nodularia_nipponensis__2 **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 137 8 5 --
  +-------------- Unio                 
  |                                    
--+  +----------- Aculamprotula        
  |  |                                 
  +--+        +-- Nodularia_douglasiae ** 
     |  +-----+                        
     |  |     +-- Nodularia_nuxpersicae **
     +--+                              
        |  +----- Cuneopsis            
        +--+                           
           |  +-- Tchangsinaia         
           +--+                        
              +-- Schistodesmus        

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 3 67 11 5 --
  +----------------- Aculamprotula           
  |                                          
--+  +-------------- Unio                    
  |  |                                       
  +--+  +----------- Inversiunio             
     |  |                                    
     +--+        +-- Tchangsinaia            
        |  +-----+                           
        |  |     +-- Schistodesmus           
        +--+                                 
           |  +----- Nodularia_nipponensis__1 **
           +--+                              
              |  +-- Nodularia_nipponensis__2 **
              +--+                           
                 +-- Nodularia_douglasiae **    

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 67 13 6 --
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Pseudocuneopsis      
     |  |                                 
     +--+        +-- Nodularia_douglasiae ** 
        |  +-----+                        
        |  |     +-- Nodularia_breviconcha **
        +--+                              
           |  +----- Schistodesmus        
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Cuneopsis            

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 20 6 --
  +-------------- Aculamprotula        
  |                                    
--+  +----------- Unio                 
  |  |                                 
  |  |     +----- Pseudocuneopsis      
  +--+  +--+                           
     |  |  |  +-- Nodularia_breviconcha **
     |  |  +--+                        
     +--+     +-- Nodularia_nipponensis **
        |                              
        |  +----- Tchangsinaia         
        +--+                           
           |  +-- Schistodesmus        
           +--+                        
              +-- Cuneopsis            

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 3 81 3 1 --
  +-------- Unio                   
--+                                
  |  +----- Nodularia_douglasiae__1 **
  +--+                             
     |  +-- Nodularia_nuxpersicae **  
     +--+                          
        +-- Nodularia_douglasiae__2 **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 2 137 8 5 --
  +----------- Unio                 
  |                                 
  |  +-------- Aculamprotula        
--+  |                              
  |  +-------- Schistodesmus        
  |  |                              
  +--+     +-- Nodularia_douglasiae ** 
     +-----+                        
     |     +-- Nodularia_nuxpersicae **
     |                              
     |  +----- Cuneopsis_heudei     
     +--+                           
        |  +-- Tchangsinaia         
        +--+                        
           +-- Cuneopsis_rufescens  

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 7 5 1 --
  +----- Aculamprotula        
--+                           
  |  +-- Nodularia_breviconcha **
  +--+                        
     +-- Nodularia_douglasiae ** 

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 2 72 13 6 --
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Pseudocuneopsis      
     |  |                                 
     +--+        +-- Nodularia_douglasiae ** 
        |  +-----+                        
        |  |     +-- Nodularia_breviconcha **
        +--+                              
           |  +----- Tchangsinaia         
           +--+                           
              |  +-- Schistodesmus        
              +--+                        
                 +-- Cuneopsis            

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 50 15 6 --
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+  +----------- Pseudocuneopsis      
     |  |                                 
     +--+        +-- Nodularia_breviconcha **
        |  +-----+                        
        |  |     +-- Nodularia_douglasiae ** 
        +--+                              
           |  +----- Schistodesmus        
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Cuneopsis            

Klishko et al., 2017 [Publ 3946], fig. 8

iT tT bi riS oT roT roS iD
2 2 0 2 4 4 1 --
  +----- Unio                 
--+                           
  |  +-- Nodularia_douglasiae ** 
  +--+                        
     +-- Nodularia_breviconcha **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 20 6 --
  +-------------- Aculamprotula        
  |                                    
--+  +----------- Unio                 
  |  |                                 
  +--+  +-------- Pseudocuneopsis      
     |  |                              
     |  |     +-- Nodularia_breviconcha **
     +--+-----+                        
        |     +-- Nodularia_douglasiae ** 
        |                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Tchangsinaia         
           +--+                        
              +-- Cuneopsis            

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
2 10 0 2 123 8 5 16.0
  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Aculamprotula        
  |                        
  +-- Cuneopsis_heudei     
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens  
  |                        
  +-- Nodularia_douglasiae ** 
  |                        
  +-- Nodularia_nuxpersicae **
  |                        
  +-- Schistodesmus        

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
2 2 0 2 74 8 5 --
  +----------------- Unio                 
  |                                       
--+  +-------------- Aculamprotula        
  |  |                                    
  +--+           +-- Nodularia_douglasiae ** 
     |  +--------+                        
     |  |        +-- Nodularia_nuxpersicae **
     +--+                                 
        |  +-------- Schistodesmus        
        +--+                              
           |  +----- Cuneopsis_heudei     
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Cuneopsis_rufescens  

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 16 3 2 --
  +----- Aculamprotula       
--+                          
  |  +-- Nodularia_douglasiae **
  +--+                       
     +-- Tchangsinaia        

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 3 --
  +-------- Aculamprotula       
--+                             
  |  +----- Unio                
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Nodularia_douglasiae **

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 9 4 --
  +----------- Aculamprotula
--+                         
  |  +-------- Unio         
  +--+                      
     |  +----- Schistodesmus
     +--+                   
        |  +-- Nodularia **    
        +--+                
           +-- Tchangsinaia 

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 42 8 3 --
  +-------- Aculamprotula       
--+                             
  |  +----- Unio                
  +--+                          
     |  +-- Tchangsinaia        
     +--+                       
        +-- Nodularia_douglasiae **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Unio                
--+                          
  |  +-- Nodularia_douglasiae **
  +--+                       
     +-- Tchangsinaia        

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 10 5 --
  +-------------- Aculamprotula       
--+                                   
  |  +----------- Unio                
  +--+                                
     |  +-------- Nodularia_douglasiae **
     +--+                             
        |  +----- Tchangsinaia        
        +--+                          
           |  +-- Cuneopsis           
           +--+                       
              +-- Schistodesmus       

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 24 2 1 --
-- Nodularia_douglasiae

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 --
  +-------------- Aculamprotula       
  |                                   
--+  +----------- Unio                
  |  |                                
  +--+        +-- Pseudocuneopsis     
     |  +-----+                       
     |  |     +-- Nodularia_douglasiae **
     +--+                             
        |  +----- Schistodesmus       
        +--+                          
           |  +-- Cuneopsis           
           +--+                       
              +-- Tchangsinaia        

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
1 1 0 1 16 11 6 --
  +----------------- Aculamprotula       
--+                                      
  |  +-------------- Unio                
  +--+                                   
     |  +----------- Pseudocuneopsis     
     +--+                                
        |  +-------- Nodularia_douglasiae **
        +--+                             
           |  +----- Tchangsinaia        
           +--+                          
              |  +-- Cuneopsis           
              +--+                       
                 +-- Schistodesmus       

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 7 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Nodularia_douglasiae **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis           

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 10 5 --
  +-------------- Aculamprotula       
--+                                   
  |  +----------- Unio                
  +--+                                
     |  +-------- Nodularia_douglasiae **
     +--+                             
        |  +----- Cuneopsis           
        +--+                          
           |  +-- Tchangsinaia        
           +--+                       
              +-- Schistodesmus       

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 13 6 --
  +----------------- Aculamprotula       
--+                                      
  |  +-------------- Unio                
  +--+                                   
     |  +----------- Pseudocuneopsis     
     +--+                                
        |  +-------- Nodularia_douglasiae **
        +--+                             
           |  +----- Tchangsinaia        
           +--+                          
              |  +-- Cuneopsis           
              +--+                       
                 +-- Schistodesmus       

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 9 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Nodularia_douglasiae **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Schistodesmus       

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 2 --
  +-- Nodularia_douglasiae **
--+                       
  +-- Cuneopsis           

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 1 --
  +-- Nodularia_douglasiae **
--+                       
  +-- Inversiunio         

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 11 5 --
  +-------------- Middendorffinaia    
--+                                   
  |  +----------- Aculamprotula       
  +--+                                
     |  +-------- Unio                
     +--+                             
        |  +----- Nodularia_douglasiae **
        +--+                          
           |  +-- Tchangsinaia        
           +--+                       
              +-- Cuneopsis           

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 13 5 --
  +----------- Aculamprotula       
  |                                
--+  +-------- Unio                
  |  |                             
  +--+     +-- Nodularia_douglasiae **
     |  +--+                       
     |  |  +-- Schistodesmus       
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Cuneopsis           

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 4 --
  +----------- Aculamprotula        
--+                                 
  |  +-------- Unio                 
  +--+                              
     |  +----- Schistodesmus        
     +--+                           
        |  +-- Nodularia_nipponensis **
        +--+                        
           +-- Tchangsinaia         

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iT tT bi riS oT roT roS iD
1 1 0 1 21 18 2 --
  +----- Unio                
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Nodularia_douglasiae **

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 12 6 --
  +----------------- Aculamprotula       
--+                                      
  |  +-------------- Unio                
  +--+                                   
     |  +----------- Pseudocuneopsis     
     +--+                                
        |  +-------- Nodularia_douglasiae **
        +--+                             
           |  +----- Tchangsinaia        
           +--+                          
              |  +-- Cuneopsis           
              +--+                       
                 +-- Schistodesmus       

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 8 3 --
  +----- Aculamprotula       
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Nodularia_douglasiae **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Unio                
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Nodularia_douglasiae **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 9 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Nodularia_douglasiae **
     +--+                          
        |  +-- Schistodesmus       
        +--+                       
           +-- Tchangsinaia        

Araujo et al., 2009 [Publ 2589], fig. 7A

iT tT bi riS oT roT roS iD
1 1 0 1 10 6 1 --
-- Nodularia_douglasiae

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 13 7 --
  +----------------- Unio                
  |                                      
--+  +-------------- Aculamprotula       
  |  |                                   
  +--+  +----------- Cuneopsis           
     |  |                                
     +--+        +-- Nodularia_douglasiae **
        |  +-----+                       
        |  |     +-- Schistodesmus       
        +--+                             
           |  +----- Tchangsinaia        
           +--+                          
              |  +-- Inversiunio         
              +--+                       
                 +-- Pseudocuneopsis     

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 10 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Nodularia_douglasiae **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Schistodesmus       

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 9 6 --
  +----------------- Aculamprotula       
--+                                      
  |  +-------------- Unio                
  +--+                                   
     |  +----------- Pseudocuneopsis     
     +--+                                
        |  +-------- Nodularia_douglasiae **
        +--+                             
           |  +----- Schistodesmus       
           +--+                          
              |  +-- Cuneopsis           
              +--+                       
                 +-- Tchangsinaia        

Dai et al., 2024 [Publ 4358], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 5 2 --
  +----- Pseudocuneopsis     
--+                          
  |  +-- Schistodesmus       
  +--+                       
     +-- Nodularia_douglasiae **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 10 5 --
  +-------------- Aculamprotula       
--+                                   
  |  +----------- Unio                
  +--+                                
     |  +-------- Nodularia_douglasiae **
     +--+                             
        |  +----- Cuneopsis           
        +--+                          
           |  +-- Tchangsinaia        
           +--+                       
              +-- Schistodesmus       

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 4 2 --
  +----- Aculamprotula       
--+                          
  |  +-- Unio                
  +--+                       
     +-- Nodularia_douglasiae **

Araujo et al., 2018 [Publ 3779], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 13 1 --
  +-- Nodularia_douglasiae **
--+                       
  +-- Unio                

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 6 2 --
  +-- Nodularia_douglasiae **
--+                       
  +-- Cuneopsis           

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iT tT bi riS oT roT roS iD
1 1 0 1 21 18 2 --
  +----- Unio                
--+                          
  |  +-- Tchangsinaia        
  +--+                       
     +-- Nodularia_dualobtusa **

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 6 4 --
  +-------------- Tchangsinaia        
--+                                   
  |  +----------- Cuneopsis_heudei    
  +--+                                
     |  +-------- Nodularia_douglasiae **
     +--+                             
        |  +----- Pseudocuneopsis     
        +--+                          
           |  +-- Cuneopsis_rufescens 
           +--+                       
              +-- Schistodesmus       

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 5 4 --
  +-------- Unio                 
--+                              
  |  +----- Schistodesmus        
  +--+                           
     |  +-- Nodularia_nipponensis **
     +--+                        
        +-- Tchangsinaia         

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 23 9 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Schistodesmus       
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Nodularia_douglasiae **

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 2 --
  +-- Nodularia_douglasiae **
--+                       
  +-- Tchangsinaia        

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 13 7 --
  +-------- Unio                
  |                             
  +-------- Aculamprotula       
  |                             
--+  +----- Cuneopsis           
  |  |                          
  |  |  +-- Nodularia_douglasiae **
  |  +--+                       
  +--+  +-- Schistodesmus       
     |                          
     |  +-- Tchangsinaia        
     |  |                       
     +--+-- Inversiunio         
        |                       
        +-- Pseudocuneopsis     

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 9 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Nodularia_douglasiae **
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Schistodesmus       

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 6 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Schistodesmus       
     +--+                          
        |  +-- Tchangsinaia        
        +--+                       
           +-- Nodularia_douglasiae **

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 12 7 --
  +----------------- Unio                
  |                                      
--+  +-------------- Aculamprotula       
  |  |                                   
  +--+  +----------- Cuneopsis           
     |  |                                
     +--+        +-- Nodularia_douglasiae **
        |  +-----+                       
        |  |     +-- Schistodesmus       
        +--+                             
           |  +----- Tchangsinaia        
           +--+                          
              |  +-- Inversiunio         
              +--+                       
                 +-- Pseudocuneopsis     

Sano et al., 2017 [Publ 3799], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 3 1 --
  +-- Nodularia_nipponensis **
--+                        
  +-- Inversiunio          

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 4 --
  +----------- Aculamprotula       
--+                                
  |  +-------- Unio                
  +--+                             
     |  +----- Schistodesmus       
     +--+                          
        |  +-- Nodularia_douglasiae **
        +--+                       
           +-- Tchangsinaia        

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
-- Nodularia_douglasiae

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 21 5 2 --
  +----- Aculamprotula       
--+                          
  |  +-- Unio                
  +--+                       
     +-- Nodularia_douglasiae **

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 6 2 --
  +----- Aculamprotula       
--+                          
  |  +-- Unio                
  +--+                       
     +-- Nodularia_douglasiae **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 11 6 --
  +-------------- Aculamprotula       
  |                                   
--+  +----------- Unio                
  |  |                                
  +--+        +-- Schistodesmus       
     |  +-----+                       
     |  |     +-- Cuneopsis           
     +--+                             
        |  +----- Pseudocuneopsis     
        +--+                          
           |  +-- Nodularia_douglasiae **
           +--+                       
              +-- Tchangsinaia        

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 4 2 --
  +----- Middendorffinaia    
--+                          
  |  +-- Nodularia_douglasiae **
  +--+                       
     +-- Unio                

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 2 --
  +-- Nodularia_douglasiae **
--+                       
  +-- Tchangsinaia        

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 3 3 --
  +-------- Aculamprotula       
--+                             
  |  +----- Schistodesmus       
  +--+                          
     |  +-- Nodularia_douglasiae **
     +--+                       
        +-- Unio                

Kano et al., 2019 [Publ 4194], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Inversiunio          
--+                        
  +-- Nodularia_nipponensis **

Page updated: 14:57:33 Wed 04 Dec 2024

 
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