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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Gonideini

genus Ptychorhynchus Simpson, 1900: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 54 cladograms in 28 publications
maximum number of terminals in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = 0, mean = 0
proportion of cladograms that support basic monophyly = 0


Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 15 10 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +-------------------- Sinosolenaia           
  |                                            
  |              +----- Gonidea                
--+  +-----------+                             
  |  |           |  +-- Leguminaia             
  |  |           +--+                          
  +--+              +-- Microcondylaea         
     |                                         
     |  +-------------- Pseudopostolata        
     |  |                                      
     +--+           +-- Parvasolenaia          
        |  +--------+                          
        |  |        +-- Koreosolenaia          
        +--+                                   
           |  +-------- Postolata              
           +--+                                
              |  +----- Ptychorhynchus_pfisteri **
              +--+                             
                 |  +-- Obovalis               
                 +--+                          
                    +-- Cosmopseudodon         

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 4 --
  +-------- Inversidens            
  |                                
--+     +-- Parvasolenaia          
  |  +--+                          
  |  |  +-- Ptychorhynchus_pfisteri **
  +--+                             
     |  +-- Microcondylaea         
     +--+                          
        +-- Sinosolenaia           

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 3 3 --
  +-------- Microcondylaea         
--+                                
  |  +----- Sinosolenaia           
  +--+                             
     |  +-- Ptychorhynchus_pfisteri **
     +--+                          
        +-- Parvasolenaia          

Liu et al., 2024 [Publ 4380], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 11 9 6 --
  +-------------- Sinosolenaia_carinata  
  |                                      
  |        +----- Inversidens            
--+  +-----+                             
  |  |     |  +-- Gonidea                
  |  |     +--+                          
  +--+        +-- Leguminaia             
     |                                   
     |  +-------- Sinosolenaia_oleivora  
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Obovalis               
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 68 2 2 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Obovalis               

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 16 2 1 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia           

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 3 3 --
  +-- Obovalis               
--+                          
  +-- Ptychorhynchus_pfisteri **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 111 8 5 --
  +-- Sinosolenaia_iridinea    
  |                            
  +-- Sinosolenaia_recognita__1
  |                            
  +-- Sinosolenaia_carinata    
  |                            
  +-- Leguminaia               
  |                            
  +-- Microcondylaea           
--+                            
  +-- Gonidea                  
  |                            
  +-- Ptychorhynchus_pfisteri **  
  |                            
  +-- Parvasolenaia            
  |                            
  +-- Sinosolenaia_recognita__2
  |                            
  +-- Sinosolenaia_oleivora    

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 4 3 --
     +-- Parvasolenaia          
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 9 6 --
  +-------- Sinosolenaia_carinata  
  |                                
  |  +----- Gonidea                
  +--+                             
  |  |  +-- Leguminaia             
  |  +--+                          
  |     +-- Microcondylaea         
--+                                
  |  +----- Ptychorhynchus_pfisteri **
  +--+                             
  |  |  +-- Parvasolenaia          
  |  +--+                          
  |     +-- Koreosolenaia          
  |                                
  |  +----- Sinosolenaia_recognita 
  +--+                             
     |  +-- Sinosolenaia_iridinea  
     +--+                          
        +-- Sinosolenaia_oleivora  

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 2 2 --
  +-- Obovalis               
--+                          
  +-- Ptychorhynchus_pfisteri **

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 2 1 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia           

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 68 2 2 --
  +----- Inversidens            
--+                             
  |  +-- Obovalis               
  +--+                          
     +-- Ptychorhynchus_pfisteri **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
1 1 0 1 36 8 7 --
  +----------------- Inversidens            
  |                                         
--+  +-------------- Sinosolenaia           
  |  |                                      
  +--+           +-- Gonidea                
     |  +--------+                          
     |  |        +-- Microcondylaea         
     +--+                                   
        |  +-------- Koreosolenaia          
        +--+                                
           |  +----- Parvasolenaia          
           +--+                             
              |  +-- Postolata              
              +--+                          
                 +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 7 6 --
           +-- Ptychorhynchus_pfisteri **
  +--------+                          
  |        +-- Parvasolenaia          
--+                                   
  |  +-------- Inversidens            
  +--+                                
     |  +----- Gonidea                
     +--+                             
        |  +-- Microcondylaea         
        +--+                          
           +-- Leguminaia             

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 5 5 --
  +----------- Sinosolenaia           
  |                                   
--+        +-- Ptychorhynchus_pfisteri **
  |  +-----+                          
  |  |     +-- Parvasolenaia          
  +--+                                
     |  +----- Gonidea                
     +--+                             
        |  +-- Microcondylaea         
        +--+                          
           +-- Leguminaia             

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 2 1 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia           

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Ptychorhynchus_pfisteri

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 44 13 9 --
  +-------------- Sinosolenaia           
  |                                      
  |        +----- Gonidea                
--+  +-----+                             
  |  |     |  +-- Leguminaia             
  |  |     +--+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |        +-- Koreosolenaia          
     |  +-----+                          
     |  |     +-- Parvasolenaia          
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Ptychorhynchus_pfisteri **
           +--+                          
              +-- Obovalis               

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 36 8 7 --
  +-------------- Sinosolenaia           
  |                                      
--+           +-- Gonidea                
  |  +--------+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |  +-------- Koreosolenaia          
     +--+                                
        |  +----- Parvasolenaia          
        +--+                             
           |  +-- Postolata              
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 41 6 2 --
  +----- Parvasolenaia          
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Sinosolenaia           

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 3 2 --
  +----- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 4 3 --
  +-------- Sinosolenaia_carinata  
  |                                
--+  +----- Gonidea                
  |  |                             
  +--+----- Sinosolenaia_oleivora  
     |                             
     |  +-- Parvasolenaia          
     +--+                          
        +-- Ptychorhynchus_pfisteri **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 8 6 --
  +----------- Sinosolenaia           
  |                                   
  |     +----- Gonidea                
--+  +--+                             
  |  |  |  +-- Leguminaia             
  |  |  +--+                          
  +--+     +-- Microcondylaea         
     |                                
     |  +----- Ptychorhynchus_pfisteri **
     +--+                             
        |  +-- Parvasolenaia          
        +--+                          
           +-- Koreosolenaia          

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 36 8 7 --
  +-------------- Sinosolenaia           
  |                                      
  |           +-- Gonidea                
--+  +--------+                          
  |  |        +-- Microcondylaea         
  |  |                                   
  +--+        +-- Koreosolenaia          
     |  +-----+                          
     |  |     +-- Parvasolenaia          
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Obovalis               
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 7 6 --
        +----- Gonidea                
  +-----+                             
  |     |  +-- Leguminaia             
  |     +--+                          
--+        +-- Microcondylaea         
  |                                   
  |  +-------- Inversidens            
  +--+                                
     |  +----- Sinosolenaia           
     +--+                             
        |  +-- Parvasolenaia          
        +--+                          
           +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 4 3 --
     +-- Parvasolenaia          
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 4 3 --
     +-- Parvasolenaia          
  +--+                          
  |  +-- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 2 1 --
  +----- Sinosolenaia_oleivora  
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Sinosolenaia_carinata  

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 3 3 --
  +-------- Sinosolenaia           
  |                                
--+  +----- Inversidens            
  |  |                             
  +--+  +-- Obovalis_omiensis__1   
     |  |                          
     +--+-- Obovalis_omiensis__2   
        |                          
        +-- Ptychorhynchus_pfisteri **

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 5 3 --
  +-------- Microcondylaea         
--+                                
  |  +----- Sinosolenaia           
  +--+                             
     |  +-- Parvasolenaia          
     +--+                          
        +-- Ptychorhynchus_pfisteri **

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 4 --
  +-------- Inversidens            
  |                                
--+     +-- Parvasolenaia          
  |  +--+                          
  |  |  +-- Ptychorhynchus_pfisteri **
  +--+                             
     |  +-- Microcondylaea         
     +--+                          
        +-- Sinosolenaia           

Wu et al., 2024 [Publ 4388], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 8 4 3 --
     +-- Postolata              
  +--+                          
  |  +-- Obovalis               
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Cosmopseudodon         

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 2 1 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia           

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 5 4 --
        +-- Microcondylaea         
  +-----+                          
  |     +-- Sinosolenaia           
--+                                
  |  +----- Parvasolenaia          
  +--+                             
     |  +-- Postolata              
     +--+                          
        +-- Ptychorhynchus_pfisteri **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 11 8 --
     +-------- Sinosolenaia           
  +--+                                
  |  |  +----- Gonidea                
  |  +--+                             
  |     |  +-- Leguminaia             
--+     +--+                          
  |        +-- Microcondylaea         
  |                                   
  |  +-------- Inversidens            
  |  |                                
  +--+     +-- Obovalis               
     |  +--+                          
     |  |  +-- Ptychorhynchus_pfisteri **
     +--+                             
        |  +-- Parvasolenaia          
        +--+                          
           +-- Koreosolenaia          

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 117 8 5 --
-- Ptychorhynchus_pfisteri

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 44 6 2 --
  +----- Parvasolenaia_rivularis   
--+                                
  |  +-- Parvasolenaia_triangularis
  +--+                             
     +-- Ptychorhynchus_pfisteri **   

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 4 3 --
     +-- Ptychorhynchus_pfisteri **
  +--+                          
  |  +-- Parvasolenaia          
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 5 3 --
     +-- Ptychorhynchus_pfisteri **
  +--+                          
  |  +-- Microcondylaea         
--+                             
  |  +-- Parvasolenaia          
  +--+                          
     +-- Sinosolenaia           

Wu et al., 2024 [Publ 4388], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 37 9 8 --
  +-------------------- Sinosolenaia           
  |                                            
--+                 +-- Gonidea                
  |  +--------------+                          
  |  |              +-- Microcondylaea         
  +--+                                         
     |  +-------------- Koreosolenaia          
     +--+                                      
        |  +----------- Parvasolenaia          
        +--+                                   
           |  +-------- Postolata              
           +--+                                
              |  +----- Cosmopseudodon         
              +--+                             
                 |  +-- Ptychorhynchus_pfisteri **
                 +--+                          
                    +-- Obovalis               

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 2 1 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia           

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 36 8 7 --
  +-------------- Sinosolenaia           
  |                                      
--+           +-- Gonidea                
  |  +--------+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |  +-------- Parvasolenaia          
     +--+                                
        |  +----- Koreosolenaia          
        +--+                             
           |  +-- Postolata              
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 7 6 --
        +----- Sinosolenaia           
  +-----+                             
  |     |  +-- Ptychorhynchus_pfisteri **
  |     +--+                          
--+        +-- Parvasolenaia          
  |                                   
  |  +-------- Inversidens            
  +--+                                
     |  +----- Gonidea                
     +--+                             
        |  +-- Microcondylaea         
        +--+                          
           +-- Leguminaia             

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 3 2 --
  +----- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 2 1 --
  +-- Ptychorhynchus_pfisteri **
--+                          
  +-- Sinosolenaia           

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 3 2 --
  +----- Sinosolenaia           
--+                             
  |  +-- Parvasolenaia          
  +--+                          
     +-- Ptychorhynchus_pfisteri **

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Ptychorhynchus_pfisteri

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 3 2 --
  +----- Ptychorhynchus_pfisteri **
--+                             
  |  +-- Microcondylaea         
  +--+                          
     +-- Sinosolenaia           

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 44 13 9 --
  +-------------- Sinosolenaia           
  |                                      
  |        +----- Gonidea                
--+  +-----+                             
  |  |     |  +-- Leguminaia             
  |  |     +--+                          
  |  |        +-- Microcondylaea         
  +--+                                   
     |        +-- Koreosolenaia          
     |  +-----+                          
     |  |     +-- Parvasolenaia          
     +--+                                
        |  +----- Postolata              
        +--+                             
           |  +-- Ptychorhynchus_pfisteri **
           +--+                          
              +-- Obovalis               

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 9 6 --
  +----------- Sinosolenaia           
  |                                   
  |     +----- Gonidea                
--+  +--+                             
  |  |  |  +-- Leguminaia             
  |  |  +--+                          
  +--+     +-- Microcondylaea         
     |                                
     |     +-- Ptychorhynchus_pfisteri **
     |     |                          
     +-----+-- Parvasolenaia          
           |                          
           +-- Koreosolenaia          

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
1 1 0 1 36 8 7 --
  +-------------- Sinosolenaia           
  |                                      
--+  +----------- Gonidea                
  |  |                                   
  +--+----------- Microcondylaea         
     |                                   
     |  +-------- Koreosolenaia          
     +--+                                
        |  +----- Parvasolenaia          
        +--+                             
           |  +-- Postolata              
           +--+                          
              +-- Ptychorhynchus_pfisteri **

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 8 5 --
           +-- Microcondylaea         
  +--------+                          
  |        +-- Sinosolenaia           
--+                                   
  |  +-------- Parvasolenaia          
  +--+                                
     |  +----- Ptychorhynchus_pfisteri **
     +--+                             
        |  +-- Postolata              
        +--+                          
           +-- Cosmopseudodon         

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 --
  +----- Sinosolenaia           
--+                             
  |  +-- Ptychorhynchus_pfisteri **
  +--+                          
     +-- Parvasolenaia          

Page updated: 14:56:59 Wed 04 Dec 2024

 
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