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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini | Leaunio

species Leaunio lienosus (Conrad, 1834): nominal species
     17 specimens available.

Unio lienosus Conrad, 1834
Conrad, 1834, Amer. J. Sci. 25(2): 339, pl. 1, fig. 4.
Conrad, 1836, Monogr. Fam. Unionidae 1(7): 60, pl. 34, fig. 2.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 251.
Type(s): HOLOTYPE ANSP_9747, Alabama R., Ala.

Unio lienosusConrad (1834), Ferussac (1835), Conrad (1836), Hanley (1843), Conrad (1853), Küster (1853), Sowerby (1866), Sowerby (1868), B.H. Wright (1888)
Margarita (Unio) lienosusLea (1836), Lea (1838)
Margaron (Unio) lienosusLea (1852), Lea (1870)
Unio (Hyridella) lienosusAdams & Adams (1857)
Unio luteolusSowerby (1867)
Lampsilis (sec. Eurynia) lienosusSimpson (1900)
Eurynia (Micromya) lienosaOrtmann (1912), Ortmann (1916), Utterback (1916)
Lampsilis (Eurynia) lienosaSimpson (1914)
Micromya lienosaOrtmann (1925)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956), Williams et al. (1993)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Gordon et al. (1979), R.I. Johnson (1980), Turgeon et al. (1988), ..., Keogh & Simons (2019), Fukata & Iigo (2020), Inoue et al. (2020), Hauck et al. (2023)
Villosa lienosa lienosaOesch (1984)
Leaunio lienosus lienosusWatters (2018)
V. lienosaMarshall et al. (2018)
Leaunio lienosusPfeiffer et al. (2019), Burlakova et al. (2019), Rogers et al. (2020), Graf & Cummings (2021), ..., Dai et al. (2023), Wu et al. (2024), Dai et al. (2024), Wu et al. (2024)

Unio caliginosus Lea, 1845
Lea, 1845, Proc. Amer. Phil. Soc. 4: 165.
Lea, 1848, Trans. Amer. Phil. Soc. 10 [O. 4]: 79 [53], pl. 7, fig. 21.
Type(s): FIGURED SYNTYPE USNM_85108, Red R., Alexandria, Louisiana.

Unio caliginosusLea (1845), Conrad (1853), Küster (1861), B.H. Wright (1888), Singley (1893), Call (1895), Pilsbry & Rhoads (1896)
Margaron (Unio) caliginosusLea (1852)
Unio (Hyridella) caliginosusAdams & Adams (1857)
Margaron (Unio) calliginosusLea (1870)
Lampsilis (sec. Eurynia) lienosusSimpson (1900)
Lampsilis (Eurynia) lienosaSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Villosa lienosa lienosaOesch (1984)
Leaunio lienosus caliginosusWatters (2018)

Unio linguaeformis Lea, 1860
Lea, 1860, Proc. Acad. Nat. Sci. 12: 305 [as ‘linguæformis’].
Lea, 1860, Jour. Acad. Nat. Sci. 4 [O. 8]: 345, pl. 56, fig. 170 [as ‘linguæformis’].
Type(s): HOLOTYPE USNM_85259, Columbus, Ga.

Unio linguaeformisLea (1860)
Margaron (Unio) linguæformisLea (1870)
Unio linguæformisB.H. Wright (1888)
Lampsilis (sec. Eurynia) nebulosusSimpson (1900)
Lampsilis (Eurynia) nebulosaSimpson (1914)
Eurynia nebulosusOrtmann (1918)
Lampsilis (Ligumia) nebulosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia nebulosaHaas (1969)
Villosa lienosaBurch (1975), Brim Box & Williams (2000), Williams et al. (2008), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio unicostatus B.H. Wright, 1899
B.H. Wright, 1899, Nautilus 13: 79.
Type(s): USNM_159966, Spring Creek, Decatur Co., Ga.

Unio unicostatusB.H. Wright (1899)
Lampsilis (sec. Eurynia) lienosus var. unicostatusSimpson (1900)
Lampsilis (Eurynia) lienosa var. unicostataSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000), Williams et al. (2008), Watters et al. (2009), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus unicostatusWatters (2018)

Unio prattii Lea, 1858
Lea, 1858, Proc. Acad. Nat. Sci. 10: 166.
Lea, 1859, Jour. Acad. Nat. Sci. 4 [O. 7]: 206 [24], pl. 24, figs. 88-88a.
Type(s): POSSIBLE SYNTYPES USNM_85145, Roswell, Cobb Co., Georgia.

Unio prattiiLea (1858), B.H. Wright (1888)
Margaron (Unio) prattiiLea (1870)
Lampsilis (sec. Eurynia) prattiiSimpson (1900)
Lampsilis (Eurynia) prattiiSimpson (1914)
Unio prattii, var.de Gregorio (1914)
Lampsilis (Ligumia) prattiiFrierson (1927)
Ligumia prattiiHaas (1969)
Villosa lienosaParmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio saxeus Conrad, 1838
Conrad, 1838, Monogr. Fam. Unionidae 2(11): back cover.
Conrad, 1840, Monogr. Fam. Unionidae 2(12): 109, pl. 60, fig. 1.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 257.

Unio saxeusConrad (1838), Conrad (1840), Conrad (1853), B.H. Wright (1888)
Margaron (Unio) saxeusLea (1852), Lea (1870)
Unio (Hyridella) saxeusAdams & Adams (1857)
Lampsilis (sec. Eurynia) lienosusSimpson (1900)
Lampsilis (Eurynia) lienosaSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Villosa lienosa lienosaOesch (1984)
Leaunio lienosus lienosusWatters (2018)

Unio haleianus Lea, 1842
Lea, 1842, Trans. Amer. Phil. Soc. 8 [O. 3]: 247 [85], pl. 27, fig. 63.
Lea, 1843, Proc. Amer. Phil. Soc. 2: 224.
Type(s): HOLOTYPE USNM_85306, Alexandria, La.

Unio haleianusLea (1842), Conrad (1853), Reeve (1865), B.H. Wright (1888)
Margaron (Unio) haleianusLea (1852), Lea (1870)
Unio (Hyridella) haleianusAdams & Adams (1857)
Unio texasensisSingley (1893)
Unio luteolusCall (1895)
Lampsilis (Carunculina) haleianusSimpson (1900)
Lampsilis (Carunculina) haleianaSimpson (1914)
Carunculina parva-haleianaFrierson (1927)
Carunculina parva parvaHaas (1969)
Villosa lienosaBurch (1975), Vidrine (1993)
Toxolasma parvusHowells et al. (1996), R.I. Johnson (1999)
Toxolasma texasensisParmalee & Bogan (1998)

Unio gouldii Lea, 1845
Lea, 1845, Proc. Amer. Phil. Soc. 4: 165.
Lea, 1848, Trans. Amer. Phil. Soc. 10 [O. 4]: 76 [50], pl. 6, fig. 16.
Type(s): HOLOTYPE (MONOTYPY) USNM_85160, Tuscaloosa, Alabama.

Unio gouldiiLea (1845), Conrad (1853), B.H. Wright (1888)
Margaron (Unio) gouldiiLea (1852), Lea (1870)
Unio (Hyridella) gouldiiAdams & Adams (1857)
Lampsilis (Proptera) gouldiiSimpson (1900), Simpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Leaunio lienosus lienosusWatters (2018)

Unio nigerrimus Lea, 1852
Lea, 1852, Trans. Amer. Phil. Soc. 10 [O. 5]: 268, pl. 18, fig. 23.
Lea, 1852, Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Type(s): HOLOTYPE USNM_85307, Alexandria, Louisiana.

Unio nigerrimusLea (1852), Conrad (1853), B.H. Wright (1888), Singley (1893)
Margaron (Unio) nigerrimusLea (1852), Lea (1870)
Unio (Hyridella) nigerrimusAdams & Adams (1857)
Unio subrostratusCall (1895)
Lampsilis (sec. Eurynia) nigerrimusSimpson (1900)
Lampsilis (Eurynia) nigerrimaSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Vidrine (1993), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Leaunio lienosus caliginosusWatters (2018)

Unio proximus Lea, 1852
Lea, 1852, Trans. Amer. Phil. Soc. 10 [O. 5]: 271 [27], pl. 20, fig. 27.
Lea, 1852, Proc. Amer. Phil. Soc. 5: 252. [nomen nudum]
Type(s): HOLOTYPE USNM_85131, Georgia.

Unio proximusLea (1852), Conrad (1853), Küster (1861), B.H. Wright (1888)
Margaron (Unio) proximusLea (1852), Lea (1870)
Unio (Hyridella) proximusAdams & Adams (1857)
Lampsilis (sec. Eurynia) ogeecheensisSimpson (1900)
Lampsilis (Eurynia) ogeecheensisSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa delumbisR.I. Johnson (1970), Burch (1975)
Villosa lienosaBrim Box & Williams (2000), Williams et al. (2008)
Leaunio lienosus lienosusWatters (2018)

Unio subellipsis Lea, 1856
Lea, 1856, Proc. Acad. Nat. Sci. 8: 262.
Lea, 1858, Jour. Acad. Nat. Sci. 4 [O. 6]: 62, pl. 10, fig. 44.
Type(s): HOLOTYPE USNM_85095, creeks near Columbus, Georgia.

Unio subellipsisLea (1856), B.H. Wright (1888)
Margaron (Unio) subellipsisLea (1870)
Lampsilis (sec. Eurynia) vibexSimpson (1900)
Lampsilis (Eurynia) vibexSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa vibexClench & Turner (1956), R.I. Johnson (1970), R.I. Johnson (1972), Burch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000)
Ligumia stagnalisHaas (1969)
Villosa lienosaWilliams et al. (2008), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio apicinus Lea, 1857
Lea, 1857, Proc. Acad. Nat. Sci. 9: 32.
Lea, 1858, Jour. Acad. Nat. Sci. 4 [O. 6]: 76, pl. 14, fig. 56.

Unio apicinusLea (1857), B.H. Wright (1888)
Margaron (Unio) apicinusLea (1870)
Lampsilis (sec. Eurynia) apicinusSimpson (1900)
Lampsilis (Eurynia) apicinaSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Villosa lienosaBurch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Leaunio lienosus lienosusWatters (2018)

Unio concestator Lea, 1857
Lea, 1857, Proc. Acad. Nat. Sci. 9: 31.
Lea, 1858, Jour. Acad. Nat. Sci. 4 [O. 6]: 66, pl. 12, fig. 48.
Type(s): HOLOTYPE USNM_85102, Columbus, Georgia.

Unio concestatorLea (1857), Küster (1861), Reeve (1865), Lea (1874), B.H. Wright (1888)
Margaron (Unio) concestatorLea (1870)
Lampsilis (sec. Eurynia) concestatorSimpson (1900)
Lampsilis (Eurynia) concestatorSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Howells et al. (1996), Parmalee & Bogan (1998), Brim Box & Williams (2000), Williams et al. (2008), Watters et al. (2009), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio fallax Lea, 1857
Lea, 1857, Proc. Acad. Nat. Sci. 9: 32.
Lea, 1858, Jour. Acad. Nat. Sci. 4 [O. 6]: 79, pl. 15, fig. 59.

Unio fallaxLea (1857), Küster (1861), Sowerby (1868), Lea (1874), B.H. Wright (1888)
Margaron (Unio) fallaxLea (1870)
Lampsilis (sec. Eurynia) concestatorSimpson (1900)
Lampsilis (Eurynia) concestatorSimpson (1914)
Villosa lienosusClench & Turner (1956)
Villosa lienosaBurch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000), Williams et al. (2008), Watters et al. (2009), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio intercedens Lea, 1857
Lea, 1857, Proc. Acad. Nat. Sci. 9: 32.
Lea, 1858, Jour. Acad. Nat. Sci. 4 [O. 6]: 77, pl. 15, fig. 57.
Type(s): HOLOTYPE USNM_85122, Columbus, Georgia.

Unio intercedensLea (1857), B.H. Wright (1888)
Margaron (Unio) intercedensLea (1870)
Lampsilis (sec. Eurynia) concestatorSimpson (1900)
Lampsilis (Eurynia) concestatorSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000), Williams et al. (2008), Watters et al. (2009), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio obfuscus Lea, 1857
Lea, 1857, Proc. Acad. Nat. Sci. 9: 172.
Lea, 1859, Jour. Acad. Nat. Sci. 4 [O. 7]: 197 [15], pl. 22, fig. 80.
Type(s): HOLOTYPE USNM_85089, Flint R., Macon Ga.

Unio obfuscusLea (1857), B.H. Wright (1888)
Margaron (Unio) obfuscusLea (1870)
Lampsilis (sec. Eurynia) graciliorSimpson (1900)
Lampsilis (Eurynia) graciliorSimpson (1914)
Lampsilis (Ligumia) graciliorFrierson (1927)
Ligumia graciliorHaas (1969)
Villosa vibexR.I. Johnson (1970), R.I. Johnson (1972), Burch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000)
Villosa lienosaWilliams et al. (2008), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio dispar Lea, 1860
Lea, 1860, Proc. Acad. Nat. Sci. 12: 305.
Lea, 1860, Jour. Acad. Nat. Sci. 4 [O. 8]: 327 [9], pl. 51, fig. 153.
Type(s): HOLOTYPE USNM_85101, Columbus, Georgia.

Unio disparLea (1860), B.H. Wright (1888)
Margaron (Unio) disparLea (1870)
Lampsilis (sec. Eurynia) disparSimpson (1900)
Lampsilis (Eurynia) disparSimpson (1914)
Lampsilis (Ligumia) vibex-disparFrierson (1927)
Villosa vibexClench & Turner (1956), R.I. Johnson (1970), R.I. Johnson (1972), Burch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000)
Ligumia vibexHaas (1969)
Villosa lienosaWilliams et al. (2008), Williams et al. (2011), Williams et al. (2014)
Leaunio lienosus lienosusWatters (2018)

Unio contiguus Lea, 1861
Lea, 1861, Proc. Acad. Nat. Sci. 13: 392.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 9]: 199 [21], pl. 28, fig. 268.
Type(s): HOLOTYPE USNM_85120, Stewarts Mill Dam, Union Co., N.C.

Unio contiguusLea (1861), B.H. Wright (1888)
Margaron (Unio) contiguusLea (1870)
Lampsilis (sec. Eurynia) concestatorSimpson (1900)
Lampsilis (Eurynia) concestatorSimpson (1914)
Lampsilis (Ligumia) delumbisFrierson (1927)
Ligumia delumbisHaas (1969)
Villosa delumbisR.I. Johnson (1970), Burch (1975)
Villosa lienosaParmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Leaunio lienosus lienosusWatters (2018)

Unio bicaelatus Reeve, 1865
Reeve, 1865, Conch. Icon. 16: pl. 26, sp. 130 [as ‘bi-cælatus’].

Unio bicaelatusReeve (1865), Paetel (1890)
Unio bi-cælatusReeve (1865)
Lampsilis (sec. Eurynia) concestatorSimpson (1900)
Lampsilis (Eurynia) concestatorSimpson (1914)
Nodularia laevirostrisFrierson (1927)
Villosa lienosaParmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Leaunio lienosus lienosusWatters (2018)

Unio fuligo Reeve, 1865
Reeve, 1865, Conch. Icon. 16: pl. 30, sp. 159.

Unio fuligoReeve (1865)
Lampsilis (sec. Eurynia) nigerrimusSimpson (1900)
Lampsilis (Eurynia) nigerrimaSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosaParmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Leaunio lienosus lienosusWatters (2018)

Unio fontanus Conrad, 1866
Conrad, 1866, Amer. J. Conch. 2: 279, pl. 15, fig. 3.
Type(s): HOLOTYPE ANSP_56458, Vicksburg, Miss.

Unio fontanusConrad (1866), B.H. Wright (1888)
Lampsilis (sec. Eurynia) lienosusSimpson (1900)
Lampsilis (Eurynia) lienosaSimpson (1914)
Lampsilis (Ligumia) lienosaFrierson (1927)
Villosa lienosusClench & Turner (1956)
Ligumia lienosaHaas (1969)
Villosa lienosaBurch (1975), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Villosa lienosa lienosaOesch (1984)
Leaunio lienosus caliginosusWatters (2018)

Unio bicoelatus Lea, 1870
‘Reeve’ Lea, 1870, Synopsis Fam. Naiades: 63 [as ‘bicœlatus’].
(error for Unio bicaelatus Reeve, 1865)

Unio bicoelatusLea (1870)

Unio calliginosus Lea, 1870
‘Lea’ Lea, 1870, Synopsis Fam. Naiades: 45.
(error for Unio caliginosus Lea, 1845)

Unio calliginosusLea (1870)

Unio haleanus Singley, 1893
Singley, 1893, Geol. Surv. Texas (1892): 321.
(error for Unio haleianus Lea, 1842)

Unio haleanusSingley (1893)

Leaunio lienosus aquilonius Watters, 2018
Watters, 2018, Visaya, Suppl. (10): 37, pl. 19.

Leaunio lienosus aquiloniusWatters (2018)

Leaunio lienosus ouachitaensis Watters, 2018
Watters, 2018, Visaya, Suppl. (10): 40, pl. 21.

Leaunio lienosus ouachitaensisWatters (2018)

Taxonomic History (since 1900)

Lampsilis (sec. Eurynia) lienosusSimpson (1900)
Lampsilis (Carunculina) haleianusSimpson (1900)
Lampsilis (Proptera) gouldiiSimpson (1900), Simpson (1914)
Lampsilis (sec. Eurynia) nigerrimusSimpson (1900)
Lampsilis (sec. Eurynia) ogeecheensisSimpson (1900)
Lampsilis (sec. Eurynia) vibexSimpson (1900)
Lampsilis (sec. Eurynia) apicinusSimpson (1900)
Lampsilis (sec. Eurynia) concestatorSimpson (1900)
Lampsilis (sec. Eurynia) graciliorSimpson (1900)
Lampsilis (sec. Eurynia) prattiiSimpson (1900)
Lampsilis (sec. Eurynia) disparSimpson (1900)
Lampsilis (sec. Eurynia) nebulosusSimpson (1900)
Lampsilis (sec. Eurynia) lienosus var. unicostatusSimpson (1900)
Eurynia (Micromya) lienosaOrtmann (1912), Ortmann (1916), Utterback (1916)
Lampsilis (Eurynia) lienosaSimpson (1914)
Lampsilis (Carunculina) haleianaSimpson (1914)
Lampsilis (Eurynia) nigerrimaSimpson (1914)
Lampsilis (Eurynia) ogeecheensisSimpson (1914)
Lampsilis (Eurynia) vibexSimpson (1914)
Lampsilis (Eurynia) apicinaSimpson (1914)
Lampsilis (Eurynia) concestatorSimpson (1914)
Lampsilis (Eurynia) graciliorSimpson (1914)
Lampsilis (Eurynia) prattiiSimpson (1914)
Unio prattii, var.de Gregorio (1914)
Lampsilis (Eurynia) disparSimpson (1914)
Lampsilis (Eurynia) nebulosaSimpson (1914)
Lampsilis (Eurynia) lienosa var. unicostataSimpson (1914)
Eurynia nebulosusOrtmann (1918)
Micromya lienosaOrtmann (1925)
Lampsilis (Ligumia) lienosaFrierson (1927)
Carunculina parva-haleianaFrierson (1927)
Lampsilis (Ligumia) graciliorFrierson (1927)
Lampsilis (Ligumia) prattiiFrierson (1927)
Lampsilis (Ligumia) vibex-disparFrierson (1927)
Lampsilis (Ligumia) nebulosaFrierson (1927)
Lampsilis (Ligumia) delumbisFrierson (1927)
Nodularia laevirostrisFrierson (1927)
Villosa lienosusClench & Turner (1956), Williams et al. (1993)
Villosa vibexClench & Turner (1956), R.I. Johnson (1970), R.I. Johnson (1972), Burch (1975), Parmalee & Bogan (1998), Brim Box & Williams (2000)
Ligumia lienosaHaas (1969)
Carunculina parva parvaHaas (1969)
Ligumia stagnalisHaas (1969)
Ligumia graciliorHaas (1969)
Ligumia prattiiHaas (1969)
Ligumia vibexHaas (1969)
Ligumia nebulosaHaas (1969)
Ligumia delumbisHaas (1969)
Villosa delumbisR.I. Johnson (1970), Burch (1975)
Villosa lienosaBurch (1975), Gordon et al. (1979), R.I. Johnson (1980), Turgeon et al. (1988), ..., Keogh & Simons (2019), Fukata & Iigo (2020), Inoue et al. (2020), Hauck et al. (2023)
Villosa lienosa lienosaOesch (1984)
Toxolasma parvusHowells et al. (1996), R.I. Johnson (1999)
Toxolasma texasensisParmalee & Bogan (1998)
Leaunio lienosus lienosusWatters (2018)
V. lienosaMarshall et al. (2018)
Leaunio lienosus caliginosusWatters (2018)
Leaunio lienosus unicostatusWatters (2018)
Leaunio lienosus aquiloniusWatters (2018)
Leaunio lienosus ouachitaensisWatters (2018)
Leaunio lienosusPfeiffer et al. (2019), Burlakova et al. (2019), Rogers et al. (2020), Graf & Cummings (2021), ..., Dai et al. (2023), Wu et al. (2024), Dai et al. (2024), Wu et al. (2024)

Page updated: 08:45:42 Wed 06 Nov 2024

 
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