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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

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This page documents data for Burch (1975), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Burch, J.B. 1975. Freshwater Unionacean Clams (Mollusca: Pelecypoda) of North America. 204 pp.

Attributed Species Names

Unio hippopaeus Lea, 1852
‘Lea’ of Authors.
Lea, 1852, Synopsis Fam. Naiades: 21 [as ‘hippopæus’].
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 261 [as ‘hippopæus’].
Adams & Adams, 1857, Gen. Rec. Moll. 2(31-32): 497 [as ‘hippopæus’].
Reeve, 1864, Conch. Icon. 16: pl. 11, sp. 40 [as ‘hippopæus’].
Lea, 1870, Synopsis Fam. Naiades: 31 [as ‘hippopæus’].
B.H. Wright, 1888, Check List N. Amer. Unionidae etc.: [3], [as ‘hippopæus’].
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 767 [as ‘hippopæa’].
Ortmann, 1909, Nautilus 22(10): 101 [as ‘Q. hippopæa’].
Ortmann, 1909, Ann. Carnegie Mus. 5: 203 [as ‘Quadrula undulata hippopæa’].
Simpson, 1914, Descr. Cat. Naiades: 816 [as ‘hippopæa’].
Frierson, 1927, Check List N. Amer. Naiades: 61.
Haas, 1969, Das Tierreich 88: 287.
Clarke, 1973, Malacologia 13(1-2): 35.
Burch, 1975, Freshw. Unio. North Amer.: 165.
Strayer & Jirka, 1997, Pearly Mussels NY (26): 98.
He & Zhuang, 2013, Freshwater Bivalves China: 27.
(error for Unio hippopoeus Lea, 1845)

Unio sowerbyanus Küster, 1852
‘Lea’ of Authors.
Küster, 1852, Conchyl. Cab. 9(2): 62.
Agassiz, 1852, Arch. Naturg. 18(1): 43.
Reeve, 1864, Conch. Icon. 16: pl. 14, sp. 58.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 520.
Simpson, 1914, Descr. Cat. Naiades: 17.
Frierson, 1927, Check List N. Amer. Naiades: 94.
Haas, 1969, Das Tierreich 88: 483.
Burch, 1975, Freshw. Unio. North Amer.: 173.
R.I. Johnson, 1978, Bull. MCZ 148: 280.
Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 91.
(error for Unio sowerbianus Lea, 1834)

Unio hanleyanus Conrad, 1853
‘Lea’ of Authors.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 250.
Sowerby, 1866, Conch. Icon. 16: pl. 46, sp. 249.
B.H. Wright, 1888, Check List N. Amer. Unionidae etc.: [3].
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 759.
Simpson, 1914, Descr. Cat. Naiades: 780.
Frierson, 1927, Check List N. Amer. Naiades: 42, sp. 175.
Haas, 1969, Das Tierreich 88: 256.
Burch, 1975, Freshw. Unio. North Amer.: 13.
Turgeon et al., 1988, Am. Fisher. Soc. Spec. Publ. (16): 32.
Williams et al., 1993, Fisheries 18(9): 13.
Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 184.
Lydeard et al., 1999, Gulf Mex. Sci. 17(2): 129.
(error for Unio hanleyianus Lea, 1852)

Unio hazelhurstianus B.H. Wright, 1888
‘Lea’ of Authors.
B.H. Wright, 1888, Check List N. Amer. Unionidae etc.: [3].
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 719 [as ‘Unio (sec. Elliptio) hazelhurstianus’].
Simpson, 1914, Descr. Cat. Naiades: 648.
Frierson, 1927, Check List N. Amer. Naiades: 29 [as ‘Elliptio (Elliptio) hazelhurstianus’].
Haas, 1969, Das Tierreich 88: 229 [as ‘Elliptio (Elliptio) hazelhurstianus’].
R.I. Johnson, 1970, Bull. MCZ 140(6): 333.
Burch, 1975, Freshw. Unio. North Amer.: 164.
(error for Unio hazlehurstianus Lea, 1858)

Unio nasutilus Simpson, 1900
‘Lea’ of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 735.
Simpson, 1914, Descr. Cat. Naiades: 691.
R.I. Johnson, 1970, Bull. MCZ 140(6): 334.
Burch, 1975, Freshw. Unio. North Amer.: 167.
Bogan, 2002, Workbook Key Freshw. Biv. NC: Table 5.
Perkins et al., 2017, Conserv. Genet. 18: 752.
Inoue et al., 2018, Invert. Syst. 32: Table S2.
(error for Unio nasutidus Lea, 1863)

Unio averillii Simpson, 1900
“Wright” of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 560.
Simpson, 1914, Descr. Cat. Naiades: 138.
Frierson, 1927, Check List N. Amer. Naiades: 76.
R.I. Johnson, 1970, Bull. MCZ 140(6): 373.
R.I. Johnson, 1972, Bull. Fla. State Mus. 16(4): 237.
Burch, 1975, Freshw. Unio. North Amer.: 159.
(error for Unio averellii B.H. Wright, 1888)

Unio cuspitatus R.I. Johnson, 1970
‘Lea’ of Authors.
R.I. Johnson, 1970, Bull. MCZ 140(6): 327.
R.I. Johnson, 1972, Bull. Fla. State Mus. 16(4): 206.
Burch, 1975, Freshw. Unio. North Amer.: 162.
(error for Unio cuspidatus Lea, 1872)

Pleurobema chattanogaense Burch, 1975
‘Lea’ Burch, 1975, Freshw. Unio. North Amer.: 12.
(error for Unio chattanoogaensis Lea, 1858)

Taxonomic Opinions

Actinonaias carinata carinata (Barnes, 1823)
+ Unio crassus Say, 1817
+ Unio ellipticus Barnes, 1823

Actinonaias carinata gibba (Simpson, 1900)

Actinonaias ellipsiformis (Conrad, 1836)

Actinonaias pectorosa (Conrad, 1834)
+ Unio perdix Lea, 1834
+ Unio biangularis Lea, 1840

Alasmidonta (Pegias) fabula (Lea, 1838)

Alasmidonta (s.s.) arcula (Lea, 1838)

Alasmidonta (s.s.) calceola (Lea, 1828)

Alasmidonta (s.s.) heterodon (Lea, 1829)

Alasmidonta (s.s.) marginata Say, 1819
+ Decurambis atropurpureum Rafinesque, 1831
+ Decurambis scriptum Rafinesque, 1831
+ Alasmidonta (Decurambis) marginata susquehannae Ortmann, 1919

Alasmidonta (s.s.) mccordi Athearn, 1964

Alasmidonta (s.s.) raveneliana (Lea, 1834)

Alasmidonta (s.s.) triangulata (Lea, 1858)
+ Strophitus wrightianus Walker, 1901

Alasmidonta (s.s.) undulata (Say, 1816)
+ Unio glabratus Sowerby & Sowerby, 1821
+ Unio hians Valenciennes, 1827
+ Alasmodonta sculptilis Say, 1829
+ Uniopsis mytiloides Swainson, 1840
+ Uniopsis radiata Swainson, 1840
+ Unio swainsoni Sowerby, 1868

Alasmidonta (s.s.) varicosa (Lamarck, 1819)
+ Unio rugulosus Wood, 1828
+ Alasmodon corrugata De Kay, 1843

Alasmidonta (s.s.) wrightiana (Walker, 1901)

Amblema antrosa Rafinesque, 1820

Amblema neisleri (Lea, 1858)

Amblema plicata (Say, 1817)
+ Unio peruviana Lamarck, 1819
+ Unio rariplicata Lamarck, 1819
+ Amblema costata Rafinesque, 1820
+ Unio perplicatus Conrad, 1841
+ Unio hippopoeus Lea, 1845
+ Unio elliottii Lea, 1857

Anodonta beringiana Middendorff, 1851
+ Anodonta youkanensis Lea, 1867

Anodonta californiensis Lea, 1852

Anodonta cataracta Say, 1817
+ Anodonta marginata Say, 1817
+ Anodonta teres Conrad, 1834
+ Anodonta newtonensis Lea, 1838
+ Anodonta excurvata De Kay, 1843
+ Anodonta housatonica Gould, 1848
+ Anodonta virgulata Lea, 1857
+ Anodonta lacustris Lea, 1857
+ Anodonta dariensis Lea, 1858
+ Anodonta gesnerii Lea, 1858
+ Anodonta hallenbeckii Lea, 1858
+ Anodonta tryonii Lea, 1862
+ Anodonta williamsii Lea, 1862
+ Anodonta dolearis Lea, 1863
+ Anodonta doliaris Lea, 1863

Anodonta couperiana Lea, 1842
+ Anodonta cowperiana Lea, 1840
+ Anodonta dunlapiana Lea, 1842

Anodonta dejecta Lewis, 1875
+ Anodonta mearnsiana Simpson, 1893

Anodonta dignota [sic] Rafinesque, 1831

Anodonta gibbosa Say, 1824

Anodonta grandis corpulenta Cooper, 1834

Anodonta grandis grandis Say, 1829
+ Anodonta plana Lea, 1834
+ Anodonta gigantea Lea, 1838
+ Anodonta ovata Lea, 1838
+ Anodonta salmonia Lea, 1838
+ Anodonta footiana Lea, 1840
+ Anodonta nilssonii Küster, 1842
+ Anodonta sulcata Küster, 1842
+ Anodonta marryattana Conrad, 1853
+ Anodonta lewisii Lea, 1857
+ Anodonta leonensis Lea, 1862
+ Anodonta dallasiana Lea, 1863
+ Anodon imbricata J.G. Anthony, 1865
+ Anodon opalina J.G. Anthony, 1865
+ Anodon subinflata J.G. Anthony, 1865
+ Anodon subangulata J.G. Anthony, 1865
+ Anodon mcnielii J.G. Anthony, 1866
+ Anodon inornata J.G. Anthony, 1866
+ Anodon ovata Sowerby, 1869
+ Anodonta houghtonensis DeCamp, 1881

Anodonta grandis simpsoniana Lea, 1861
+ Anodonta kennicottii Lea, 1861

Anodonta imbecilis [sic] Say, 1829
+ Anodonta incerta Lea, 1834
+ Anodon horda Gould, 1855
+ Anodonta henryana Lea, 1857

Anodonta implicata Say, 1829

Anodonta inflata Rafinesque, 1831

Anodonta kennerlyi Lea, 1860

Anodonta nuttalliana Lea, 1838
+ Anodonta wahlamatensis Lea, 1838
+ Anodonta rotundovata Trask, 1854
+ Anodonta triangularis Trask, 1854
+ Anodon rostratus Sowerby, 1867
+ Anodonta laosensis Fischer, 1891
+ Anodonta nuttalliana var. idahoensis Hemphill, 1891

Anodonta oregonensis Lea, 1838
+ Anodon cognata Gould, 1850

Anodonta peggyae Johnson, R.I., 1965

Anodonta suborbiculata Say, 1831

Anodontoides ferussacianus Lea, 1834
+ Anodonta buchanensis Lea, 1838
+ Anodonta subcylindracea Lea, 1838

Anodontoides radiatus (Conrad, 1834)
+ Margaritana elliottii Lea, 1858
+ Margaritana elliptica Lea, 1859
+ Anodonta showalterii Lea, 1860

Arcidens confragosa (Say, 1829)

Arkansia wheeleri Ortmann & Walker, 1912

Carunculina parva (Barnes, 1823)
+ Unio glans Lea, 1831
+ Unio paulus Lea, 1840
+ Unio moestus Lea, 1841
+ Unio minor Lea, 1843
+ Unio bairdianus Lea, 1857
+ Unio texasiensis Lea, 1857
+ Unio germanus Lea, 1861
+ Unio bealei Lea, 1862
+ Unio cromwellii Lea, 1865
+ Unio marginis Lea, 1865
+ Unio corvinus Lea, 1868
+ Unio corvunculus Lea, 1868
+ Unio cylindrellus Lea, 1868
+ Unio stearnsii B.H. Wright, 1888
+ Unio singleyanus Marsh, 1891
+ Lampsilis mearnsi Simpson, 1900
+ Carunculina parva cahni F.C. Baker, 1927

Carunculina pulla (Conrad, 1838)
+ Carunculina patrickae Bates, 1966

Cumberlandia monodonta (Say, 1829)

Cyclonaias tuberculata (Rafinesque, 1820)

Cyprogenia aberti (Conrad, 1850)

Cyprogenia irrorata (Lea, 1828)
+ Obovaria stegaria Rafinesque, 1820

Cyrtonaias tampicoensis (Lea, 1838)
+ Unio berlandierii Lea, 1857

Dromus dromus [sic] (Lea, 1834)

Dysnomia arcaeformis (Lea, 1831)

Dysnomia biemarginata (Lea, 1857)

Dysnomia brevidens (Lea, 1831)
+ Unio interruptus Conrad, 1834

Dysnomia capsaeformis (Lea, 1834)

Dysnomia flexuosa (Rafinesque, 1820)
+ Unio foliatus Hildreth, 1828

Dysnomia florentina (Lea, 1857)
+ Unio turgidulus Lea, 1858
+ Unio nux Küster, 1861
+ Unio sacculus Reeve, 1864

Dysnomia haysiana (Lea, 1834)
+ Unio sowerbianus Lea, 1834

Dysnomia lenior (Lea, 1840)

Dysnomia lewisi (Walker, 1910)

Dysnomia metastriata (Conrad, 1840)
+ Unio othcaloogensis Lea, 1857
+ Unio compactus Lea, 1859

Dysnomia modicella (Lea, 1859)

Dysnomia penita (Conrad, 1834)

Dysnomia personata (Say, 1829)
+ Unio pileus Lea, 1831
+ Unio capillaris Lea, 1834

Dysnomia stewardsoni (Lea, 1852)

Dysnomia sulcata (Lea, 1829)
+ Unio ridibundus Say, 1829
+ Unio gibbosus var. perobliquus Conrad, 1836
+ Unio pectitus Conrad, 1854
+ Truncilla (Scalenaria) sulcata var. delicata Simpson, 1900

Dysnomia torulosa propinqua (Lea, 1857)

Dysnomia torulosa torulosa (Rafinesque, 1820)
+ Amblema gibbosa Rafinesque, 1820
+ Unio perplexus Lea, 1831
+ Unio gibbosus Ferussac, 1835
+ Unio phillipsii Conrad, 1835
+ Unio rangianus Lea, 1838
+ Unio cincinnatiensis Lea, 1840
+ Unio obliquus Potiez & Michaud, 1844
+ Unio sampsonii Lea, 1861
+ Unio gubernaculum Reeve, 1865

Dysnomia triquetra (Rafinesque, 1820)
+ Unio cuneatus Swainson, 1823
+ Unio triangularis Barnes, 1823
+ Unio formosus Lea, 1831

Dysnomia turgidula (Lea, 1858)
+ Unio deviatus Reeve, 1864
+ Truncilla curtisii Utterback, 1916

Ellipsaria lineolata (Rafinesque, 1820)
+ Obliquaria (Plagiola) depressa Rafinesque, 1820
+ Obliquaria (Ellipsaria) ellipsaria Rafinesque, 1820
+ Unio securis Lea, 1829

Elliptio (Canthyria) spinosa (Lea, 1838)

Elliptio (s.s.) arctata (Conrad, 1834)
+ Unio strigosus Lea, 1840
+ Unio tortivus Lea, 1840
+ Unio lazarus Lea, 1852
+ Unio perstriatus Lea, 1852
+ Unio gracilentus Lea, 1857
+ Unio perlatus Lea, 1863

Elliptio (s.s.) beadleiana (Lea, 1861)

Elliptio (s.s.) chipolaensis (Walker, 1905)

Elliptio (s.s.) congaraeus (Lea, 1831)
+ Unio fulvus Lea, 1834
+ Unio planilateris Conrad, 1838
+ Unio lecontianus Lea, 1838
+ Unio pusillus Lea, 1840
+ Unio rufusculus Lea, 1852
+ Unio sordidus Lea, 1852
+ Unio buxeus Lea, 1852
+ Unio forbesianus Lea, 1852
+ Unio moussonianus Lea, 1852
+ Unio gibbesianus Lea, 1852
+ Unio corvus Lea, 1859
+ Unio vestitus Lea, 1861
+ Unio dorsatus Lea, 1868
+ Unio strumosus Lea, 1872

Elliptio (s.s.) crassidens (Lamarck, 1819)
+ Unio (Elliptio) nigra Rafinesque, 1820
+ Unio cuneatus Barnes, 1823
+ Unio incrassatus Lea, 1840
+ Unio satillaensis Lea, 1858
+ Unio spissus Lea, 1859
+ Unio discus Sowerby, 1868
+ Unio lehmanii S.H. Wright, 1897
+ Unio danielsii B.H. Wright, 1899
+ Unio polymorphus B.H. Wright, 1899
+ Elliptio pachyodon Olson & Harbison, 1953

Elliptio (s.s.) dariensis (Lea, 1842)
+ Unio monroensis Lea, 1843
+ Unio websterii B.H. Wright, 1888
+ Unio hartwrightii B.H. Wright, 1896

Elliptio (s.s.) dilatata (Rafinesque, 1820)
+ Obliquaria (Ellipsaria) cuprea Rafinesque, 1820
+ Unio gibbosus Barnes, 1823
+ Elliptio dilatata var. sterkii Grier, 1918

Elliptio (s.s.) fraterna (Lea, 1852)
+ Unio anthonyi Lea, 1861

Elliptio (s.s.) hopetonensis (Lea, 1838)
+ Unio inusitatis Lea, 1859

Elliptio (s.s.) jayensis (Lea, 1838)
+ Unio aheneus Lea, 1843
+ Unio marshii B.H. Wright, 1888
+ Unio tryoni B.H. Wright, 1888
+ Unio waltoni B.H. Wright, 1888
+ Unio oscari B.H. Wright, 1892
+ Unio leonensis Simpson, 1892
+ Unio buxtoni B.H. Wright, 1897
+ Unio suttoni B.H. Wright, 1897

Elliptio (s.s.) lanceolata (Lea, 1828)
+ Unio angustatus Lea, 1831
+ Unio productus Conrad, 1836
+ Unio folliculatus Lea, 1838
+ Unio duttonianus Lea, 1841
+ Unio sagittiformis Lea, 1852
+ Unio emmonsii Lea, 1857
+ Unio extensus Lea, 1857
+ Unio maconensis Lea, 1857
+ Unio naviculoides Lea, 1857
+ Unio rostraeformis Lea, 1857
+ Unio rostriformis Lea, 1857
+ Unio hazlehurstianus Lea, 1858
+ Unio nasutidus Lea, 1863
+ Unio viridulus Lea, 1863
+ Unio nasutulus Lea, 1866
+ Unio attenuatus Lea, 1872
+ Unio exacutus Lea, 1872
+ Unio rostellum Lea, 1872
+ Unio subcylindraceus Lea, 1874
+ Unio arctior var. fisheropsis de Gregorio, 1914

Elliptio (s.s.) marsupiobesa Fuller, 1972

Elliptio (s.s.) mcmichaeli Clench & Turner, 1956

Elliptio (s.s.) nigella (Lea, 1853)
+ Unio denigratus Lea, 1857
+ Unio purpurellus Lea, 1857

Elliptio (s.s.) shepardiana (Lea, 1834)

Elliptio (s.s.) waccamawensis (Lea, 1863)

Elliptoideus sloatianus (Lea, 1840)
+ Unio atromarginatus Lea, 1840
+ Unio aratus Conrad, 1849
+ Unio plectophorus Conrad, 1849

Fusconaia askewi (Marsh, 1896)

Fusconaia barnesiana (Lea, 1838)
+ Unio bigbyensis Lea, 1841
+ Unio tumescens Lea, 1845
+ Unio estabrookianus Lea, 1845
+ Unio pudicus Lea, 1860
+ Unio raveneliensis Reeve, 1864
+ Unio lyonii Lea, 1865
+ Unio fassinans Lea, 1868
+ Unio meredithii Lea, 1870
+ Unio validus Lea, 1871
+ Unio crudus Lea, 1871
+ Unio radiosus Lea, 1871
+ Unio tellicoensis Lea, 1872
+ Unio lenticularis Lea, 1872
+ Pleurobema fassinans var. rhomboidea Simpson, 1900

Fusconaia cor (Conrad, 1834)
+ Unio edgarianus Lea, 1840
+ Unio obuncus Lea, 1871
+ Unio andersonensis Lea, 1872
+ Fusconaia cor var. analoga Ortmann, 1918

Fusconaia cuneolus (Lea, 1840)
+ Unio clinchensis Lea, 1867
+ Unio tuscumbiensis Lea, 1871
+ Unio appressus Lea, 1871
+ Unio flavidus Lea, 1872

Fusconaia ebenus (Lea, 1831)

Fusconaia escambia Clench & Turner, 1956

Fusconaia flava (Rafinesque, 1820)
+ Unio rubiginosus Lea, 1829
+ Unio hebetatus Conrad, 1854
+ Unio rubidus Lea, 1861
+ Fusconaia flava var. parvula Grier, 1918

Fusconaia lananensis (Frierson, 1900)

Fusconaia masoni (Conrad, 1834)
+ Unio pumilus Lea, 1838
+ Unio merus Lea, 1852
+ Unio striatulus Lea, 1857
+ Unio castus Lea, 1860
+ Unio brimleyi S.H. Wright, 1897

Fusconaia ozarkensis (Call, 1887)
+ Pleurobema argentea pannosa Simpson, 1900
+ Pleurobema brevis subelliptica Simpson, 1900
+ Pleurobema utterbackii Utterback, 1915

Fusconaia subrotunda (Lea, 1831)
+ Unio brevialis Crouch, 1827
+ Unio kirtlandianus Lea, 1834
+ Unio personatus Conrad, 1834
+ Unio politus Conrad, 1837
+ Unio lesueurianus Lea, 1840
+ Unio pilaris Lea, 1840
+ Unio globatus Lea, 1871
+ Unio bursapastoris B.H. Wright, 1896
+ Quadrula flexuosa Simpson, 1900
+ Quadrula andrewsii Marsh, 1902
+ Quadrula beauchampii Marsh, 1902
+ Fusconaja [sic] subrotunda leucogona Ortmann, 1913
+ Fusconaia subtrotunda Ortmann, 1925

Fusconaia succissa (Lea, 1852)
+ Unio cacao Lea, 1859
+ Quadrula (sec. Fusconaia) wrightii Simpson, 1914

Fusconaia undata (Barnes, 1823)
+ Unio trigonus Lea, 1831
+ Fusconaia selecta Wheeler, 1914
+ Fusconaia undata wagneri F.C. Baker, 1928

Glebula rotundata (Lamarck, 1819)
+ Unio suborbiculata Lamarck, 1819
+ Unio glebulus Say, 1831
+ Unio subglobosus Lea, 1834
+ Unio grandensis Conrad, 1855
+ Unio granadensis Simpson, 1900

Gonidea angulata (Lea, 1838)
+ Anodon feminalis Gould, 1850
+ Anodonta randalli Trask, 1854
+ Anodon biangulata Sowerby, 1869

Hemistena lata (Rafinesque, 1820)
+ Unio dehiscens Say, 1829
+ Unio oriens Lea, 1831
+ Odatelia radiata Rafinesque, 1832

Lampsilis altilis (Conrad, 1834)

Lampsilis australis Simpson, 1900

Lampsilis binominatus (Simpson, 1900)
+ Unio lineatus Lea, 1840

Lampsilis bracteata (Gould, 1855)

Lampsilis cariosa (Say, 1816)
+ Lampsilis pallida Rafinesque, 1820
+ Unio crocatus Lea, 1841
+ Unio oratus Conrad, 1849

Lampsilis claibornensis (Lea, 1838)
+ Unio obtusus Lea, 1840
+ Unio pallescens Lea, 1845
+ Unio contrarius Conrad, 1849

Lampsilis dolabraeformis (Lea, 1838)

Lampsilis excavatus (Lea, 1857)
+ Unio ornatus Sowerby, 1866

Lampsilis fasciola Rafinesque, 1820
+ Unio multiradiatus Lea, 1829

Lampsilis haddletoni Athearn, 1964

Lampsilis hydiana (Lea, 1838)

Lampsilis ochracea (Say, 1816)

Lampsilis orbiculata (Hildreth, 1828)

Lampsilis ovata (Say, 1816)
+ Lampsilis cardium Rafinesque, 1820
+ Unio ventricosus Barnes, 1823
+ Unio occidens Lea, 1829
+ Unio subovatus Lea, 1831
+ Unio fasciolus Ferussac, 1835
+ Unio lenis Conrad, 1838
+ Unio canadensis Lea, 1857
+ Unio dolabraeformis Sowerby, 1867
+ Unio latissimus Sowerby, 1868
+ Lampsilis ventricosa cohongoronta Ortmann, 1912
+ Lampsilis (s.s.) ventricosa var. lurida Simpson, 1914
+ Lampsilis ventricosa perglobosa F.C. Baker, 1928
+ Lampsilis ventricosa winnebagoensis F.C. Baker, 1928

Lampsilis perpasta (Lea, 1861)

Lampsilis radiata radiata (Gmelin, 1791)
+ Unio luteola Lamarck, 1819
+ Unio lineata Bory de St. Vincent, 1827
+ Unio oblongatus Wood, 1828
+ Unio tenebrosus Conrad, 1834
+ Unio melinus Conrad, 1838
+ Unio boydianus Lea, 1840
+ Unio rosaceus De Kay, 1843
+ Unio elongata Goodrich, 1859
+ Unio obliquiradiatus Reeve, 1865
+ Unio conspicuus Lea, 1872
+ Lampsilis radiata oneidensis F.C. Baker, 1916
+ Unio virginea Frierson, 1927

Lampsilis radiata siliquoidea (Barnes, 1823)
+ Unio inflatus Barnes, 1823
+ Unio borealis Gray, 1882
+ Unio superiorensis Marsh, 1897

Lampsilis radiata splendida (Lea, 1838)
+ Unio regularis Sowerby, 1866

Lampsilis straminea (Conrad, 1834)

Lampsilis streckeri Frierson, 1927

Lampsilis subangulata (Lea, 1840)
+ Unio kirklandianus S.H. Wright, 1897

Lampsilis teres (Rafinesque, 1820)
+ Unio anodontoides Lea, 1831
+ Unio floridensis Lea, 1852
+ Lampsilis fallaciosus Smith, 1899

Lasmigona complanata (Barnes, 1823)
+ Unio katherinae Lea, 1838

Lasmigona compressa (Lea, 1829)
+ Unio compressus var. plebius C.B. Adams, 1842

Lasmigona costata (Rafinesque, 1820)
+ Alasmodonta rugosa Barnes, 1823
+ Lasmigona costata var. ereganensis Grier, 1918
+ Lasmigona costata nuda F.C. Baker, 1928
+ Lasmigona costata pepinensis F.C. Baker, 1928

Lasmigona holstonia (Lea, 1838)

Lasmigona subviridis (Conrad, 1835)
+ Unio tappanianus Lea, 1838
+ Unio hyalinus Lea, 1845
+ Unio pygmaeus Lea, 1852
+ Unio decoratus Lea, 1852
+ Unio charlottensis Lea, 1863
+ Unio pertenuis Lea, 1863
+ Unio insolidus Lea, 1872

Lemiox rimosus (Rafinesque, 1831)
+ Unio caelatus Conrad, 1834

Leptodea amphichaena (Frierson, 1898)

Leptodea fragilis (Rafinesque, 1820)
+ Unio gracilis Barnes, 1823

Leptodea laevissima (Lea, 1829)

Leptodea leptodon (Rafinesque, 1820)
+ Anodon purpurascens Swainson, 1823
+ Symphynota tenuissima Lea, 1829
+ Unio velum Say, 1829

Lexingtonia dolabelloides (Lea, 1840)
+ Unio maculatus Conrad, 1834
+ Unio mooresianus Lea, 1857
+ Unio thorntonii Lea, 1857
+ Unio subglobatus Lea, 1871
+ Unio circumactus Lea, 1871
+ Pleurobema conradi Vanatta, 1915

Lexingtonia subplana (Conrad, 1834)

Ligumia nasuta (Say, 1817)
+ Obliquaria (Ellipsaria) attenuata Rafinesque, 1820
+ Unio rostrata Valenciennes, 1827
+ Unio nashvillianus Lea, 1834
+ Unio vaughanianus Sowerby, 1868

Ligumia recta (Lamarck, 1819)
+ Unio (Eurynia) latissima Rafinesque, 1820
+ Unio sageri Conrad, 1836

Margaritifera falcata (Gould, 1850)

Margaritifera hembeli (Conrad, 1838)

Margaritifera margaritifera (Linnaeus, 1758)

Medionidus acutissimus (Lea, 1831)
+ Unio rubellinus Lea, 1857

Medionidus conradicus (Lea, 1834)

Medionidus mcglameriae van der Schalie, 1939

Medionidus parvulus (Lea, 1860)

Medionidus penicillatus (Lea, 1857)
+ Unio walkeri B.H. Wright, 1897
+ Unio kingii B.H. Wright, 1900
+ Medionidus simpsonianus Walker, 1905

Megalonaias gigantea (Barnes, 1823)
+ Unio undulatus Barnes, 1823
+ Unio heros Say, 1829
+ Unio multiplicatus Lea, 1831
+ Unio boykinianus Lea, 1840

Obliquaria atroviolacea Rafinesque, 1820

Obliquaria cliffordiana Rafinesque, 1820

Obliquaria interrupta Rafinesque, 1820

Obliquaria nodulata Rafinesque, 1820
+ Unio pustulatus Lea, 1831

Obliquaria obliquata Rafinesque, 1820

Obliquaria pusilla Rafinesque, 1820

Obliquaria reflexa Rafinesque, 1820
+ Unio cornutus Barnes, 1823

Obliquaria retusa Rafinesque, 1820

Obliquaria rubra Rafinesque, 1820

Obliquaria sintoxia Rafinesque, 1820

Obliquaria sinuata Rafinesque, 1820

Obliquaria triangularis Rafinesque, 1820

Obovaria jacksoniana (Frierson, 1912)

Obovaria obovalis Rafinesque, 1820

Obovaria olivaria (Rafinesque, 1820)
+ Unio ellipsis Lea, 1828

Obovaria pachostea Rafinesque, 1820

Obovaria retusa (Lamarck, 1819)
+ Obovaria torsa Rafinesque, 1820

Obovaria rotulata (Wright, B.H., 1899)

Obovaria subrotunda (Rafinesque, 1820)
+ Obovaria striata Rafinesque, 1820
+ Unio circulus Lea, 1829
+ Unio depygis Conrad, 1866

Obovaria unicolor (Lea, 1845)

Plectomerus dombeyanus (Valenciennes, 1833)
+ Unio interruptus Say, 1831
+ Unio trapezoides Lea, 1831
+ Bariosta ponderosus Rafinesque, 1831

Plethobasus cooperianus (Lea, 1834)

Plethobasus cyphyus (Rafinesque, 1820)
+ Unio aesopus Green, 1827
+ Unio cicatricosus Say, 1829
+ Unio varicosus Lea, 1831

Pleurobema aldrichianum Goodrich, 1931

Pleurobema altum (Conrad, 1854)

Pleurobema avellanum Simpson, 1900

Pleurobema chattanogaense [sic] (Lea, 1858)

Pleurobema clava (Lamarck, 1819)
+ Unio (Aximedia) elliptica Rafinesque, 1820
+ Pleurobema cuneata Rafinesque, 1820
+ Pleurobema mytiloides Rafinesque, 1820
+ Obliquaria (Scalenaria) scalenia Rafinesque, 1820
+ Unio patulus Lea, 1829

Pleurobema collina (Conrad, 1837)

Pleurobema cordatum (Rafinesque, 1820)
+ Obliquaria (Sintoxia) lateralis Rafinesque, 1820
+ Unio catillus Conrad, 1836
+ Unio solidus Lea, 1838
+ Unio plenus Lea, 1840

Pleurobema curtum (Lea, 1859)

Pleurobema decisum (Lea, 1831)

Pleurobema flavidulum (Lea, 1861)

Pleurobema furvum (Conrad, 1834)

Pleurobema georgianum (Lea, 1841)
+ Unio favosus Lea, 1857

Pleurobema hagleri (Frierson, 1900)

Pleurobema hanleyanum [sic] (Lea, 1852)

Pleurobema irrasum (Lea, 1861)

Pleurobema johannis (Lea, 1859)

Pleurobema marshalli Frierson, 1927

Pleurobema meredithii (Lea, 1858)

Pleurobema murrayense (Lea, 1868)

Pleurobema nucleopse (Conrad, 1849)

Pleurobema nux (Lea, 1852)

Pleurobema oviforme (Conrad, 1834)
+ Unio holstonensis Lea, 1840
+ Unio argenteus Lea, 1841
+ Unio mundus Lea, 1857
+ Unio lawii Lea, 1871
+ Unio bellulus Lea, 1872

Pleurobema perovatum (Conrad, 1834)

Pleurobema pyriforme (Lea, 1857)
+ Unio striatus Lea, 1840
+ Unio bulbosus Lea, 1857
+ Unio modicus Lea, 1857
+ Unio amabilis Lea, 1865
+ Unio reclusus B.H. Wright, 1898
+ Unio harperi B.H. Wright, 1899
+ Pleurobema simpsoni Vanatta, 1915

Pleurobema rubellum (Conrad, 1834)

Pleurobema showalteri (Lea, 1860)

Pleurobema simulans (Lea, 1871)

Pleurobema stabile (Lea, 1861)

Pleurobema strodeanus (Wright, B.H., 1898)
+ Pleurobema patsaligensis Simpson, 1900

Pleurobema tombigbeanum Frierson, 1908

Pleurobema troschelianum (Lea, 1852)

Pleurobema verum (Lea, 1861)

Popenaias buckleyi (Lea, 1841)
+ Unio cunninghami B.H. Wright, 1883
+ Unio dallii B.H. Wright, 1888
+ Unio dorei B.H. Wright, 1888
+ Unio hinkleyi B.H. Wright, 1888
+ Unio orcuttii S.H. Wright, 1888
+ Unio ferrissii Marsh, 1891
+ Unio subluridus Simpson, 1892
+ Unio pinei B.H. Wright, 1897
+ Unio caloosaensis Dall, 1898
+ Unio tenuisculus Frierson, 1911
+ Unio (Elliptio) sanctorumjohanium B.H. Wright, 1933
+ Unio (Elliptio) webbianus hartii B.H. Wright, 1934
+ Unio (Elliptio) webbianus B.H. Wright, 1934

Popenaias popei (Lea, 1857)

Proptera alata (Say, 1816)
+ Metaptera megaptera Rafinesque, 1820

Proptera capax (Green, 1832)

Proptera purpurata (Lamarck, 1819)

Ptychobranchus fasciolare (Rafinesque, 1820)
+ Unio phaseolus Hildreth, 1828

Ptychobranchus greeni (Conrad, 1834)
+ Unio foremanianus Lea, 1842
+ Unio flavescens Lea, 1845
+ Unio simplex Lea, 1845
+ Unio velatus Conrad, 1854
+ Unio woodwardius Lea, 1857
+ Unio woodwardianus Lea, 1859
+ Unio trinacrus Lea, 1861

Ptychobranchus jonesi (van der Schalie, 1934)

Ptychobranchus occidentalis (Conrad, 1836)

Ptychobranchus subtentum (Say, 1825)

Quadrula archeri Frierson, 1905

Quadrula aurea (Lea, 1859)

Quadrula cylindrica (Say, 1817)

Quadrula intermedia (Conrad, 1836)
+ Unio tuberosus Lea, 1840

Quadrula metanevra (Rafinesque, 1820)
+ Unio nodosus Barnes, 1823
+ Unio wardii Lea, 1861

Quadrula pustulosa (Lea, 1831)
+ Obliquaria (Quadrula) bullata Rafinesque, 1820
+ Unio prasinus Conrad, 1834
+ Unio nodiferus Conrad, 1841
+ Unio pernodosus Lea, 1845
+ Unio asperatus Lea, 1861
+ Unio vallatus Lea, 1868
+ Unio refulgens Lea, 1868
+ Unio sphaericus Lea, 1868
+ Unio cahabensis Lea, 1871

Quadrula quadrula (Rafinesque, 1820)
+ Unio rugosus Barnes, 1823
+ Unio lacrymosus Lea, 1828
+ Unio apiculatus Say, 1829
+ Unio asperrimus Lea, 1831
+ Unio asper Lea, 1831
+ Unio fragosus Conrad, 1835
+ Unio rumphianus Lea, 1852
+ Unio nobilis Conrad, 1854
+ Unio blandianus Lea, 1857
+ Unio forsheyi Lea, 1859
+ Unio speciosus Lea, 1862
+ Quadrula quadrula contraryensis Utterback, 1915
+ Quadrula (Quadrula) quadrula bullocki F.C. Baker, 1928

Quadrula stapes (Lea, 1831)

Quincuncina burkei Walker in Ortmann & Walker, 1922

Quincuncina guadalupensis Wurtz, 1950

Quincuncina infucata (Conrad, 1834)
+ Unio securiformis Conrad, 1849
+ Unio kleinianus Lea, 1852

Simpsoniconcha ambigua (Say, 1825)

Strophitus subvexa (Conrad, 1834)
+ Margaritana connasaugaensis Lea, 1858
+ Margaritana gesnerii Lea, 1858
+ Margaritana spillmanii Lea, 1858
+ Margaritana tombecbeensis Lea, 1858
+ Margaritana alabamensis Lea, 1861
+ Margaritana tombigbeensis Lea, 1862
+ Margaritana columbensis Lea, 1867

Strophitus undulatus (Say, 1817)
+ Anodonta pensylvanica Lamarck, 1819
+ Anodon rugosus Swainson, 1822
+ Anodon areolatus Swainson, 1829
+ Alasmodonta edentula Say, 1829
+ Anodonta virgata Conrad, 1836
+ Anodonta pavonia Lea, 1838
+ Anodonta wardiana Lea, 1838
+ Anodon unadilla De Kay, 1843
+ Anodonta tetragona Lea, 1845
+ Anodonta arkansensis Lea, 1852
+ Anodonta shaefferiana Lea, 1852
+ Anodon papyracea J.G. Anthony, 1865
+ Alasmodon rhombica J.G. Anthony, 1865
+ Anodon annulatus Sowerby, 1867
+ Anodon quadriplicatus Sowerby, 1870
+ Anodonta salmonea Clessin, 1873
+ Strophitus undulatus ovatus Frierson, 1927
+ Strophitus rugosus lacustris F.C. Baker, 1928
+ Strophitus rugosus pepinensis F.C. Baker, 1928
+ Strophitus rugosus winnebagoensis F.C. Baker, 1928

Tritogonia verrucosa (Rafinesque, 1820)

Truncilla donaciformis (Lea, 1828)
+ Unio zigzag Lea, 1829

Truncilla macrodon (Lea, 1859)

Truncilla truncata Rafinesque, 1820
+ Unio elegans Lea, 1831

Uniomerus tetralasmus (Say, 1830)
+ Unio declivis Say, 1831
+ Unio obesus Lea, 1831
+ Unio camptodon Say, 1832
+ Unio blandingianus Lea, 1834
+ Unio geometricus Lea, 1834
+ Unio excultus Conrad, 1838
+ Unio sayii Tappan, 1839
+ Unio parallelus Conrad, 1841
+ Unio buddianus Lea, 1843
+ Unio paludicolus Gould, 1845
+ Unio symmetricus Lea, 1845
+ Unio hebes Lea, 1852
+ Unio ineptus Lea, 1852
+ Unio paludicolor Conrad, 1853
+ Unio rivularis Conrad, 1853
+ Unio porrectus Conrad, 1854
+ Unio subcroceus Conrad, 1854
+ Unio manubius Gould, 1855
+ Unio plantii Lea, 1857
+ Unio columbensis Lea, 1857
+ Unio jamesianus Lea, 1857
+ Unio cicur Lea, 1861
+ Unio squalidus Lea, 1863
+ Unio electrinus Reeve, 1865
+ Unio bisselianus Lea, 1867
+ Unio jewettii Lea, 1867
+ Unio pawensis Lea, 1868
+ Unio rivicolus Conrad, 1868

Villosa amygdala (Lea, 1843)
+ Unio trossulus Lea, 1843
+ Unio trosculus Lea, 1843
+ Unio papyraceus Gould, 1845
+ Unio lepidus Gould, 1856
+ Unio vesicularis Lea, 1872
+ Unio singleyanus Marsh, 1891

Villosa brevicula (Call, 1887)

Villosa choctawensis Athearn, 1964

Villosa constricta (Conrad, 1838)
+ Unio genthii Lea, 1857

Villosa delumbis (Conrad, 1834)
+ Unio tenerus Ravenel, 1834
+ Unio vaughanianus Lea, 1838
+ Unio ogeecheensis Conrad, 1849
+ Unio concavus Lea, 1852
+ Unio proximus Lea, 1852
+ Unio contiguus Lea, 1861
+ Unio genuinus Lea, 1868

Villosa fabalis (Lea, 1831)

Villosa iris (Lea, 1829)

Villosa lienosa (Conrad, 1834)
+ Unio saxeus Conrad, 1838
+ Unio haleianus Lea, 1842
+ Unio caliginosus Lea, 1845
+ Unio gouldii Lea, 1845
+ Unio nigerrimus Lea, 1852
+ Unio apicinus Lea, 1857
+ Unio concestator Lea, 1857
+ Unio fallax Lea, 1857
+ Unio intercedens Lea, 1857
+ Unio linguaeformis Lea, 1860
+ Unio porphyreus Lea, 1861
+ Unio fontanus Conrad, 1866
+ Unio unicostatus B.H. Wright, 1899

Villosa nebulosa (Conrad, 1834)
+ Unio notatus Lea, 1838
+ Unio obscurus Lea, 1838
+ Unio simus Lea, 1838
+ Unio amoenus Lea, 1840
+ Margaron (Unio) cumberlandianus Lea, 1852
+ Unio radians Lea, 1857
+ Unio jonesii Lea, 1859
+ Unio planicostatus Lea, 1860
+ Unio scitulus Lea, 1860
+ Unio plancus Lea, 1860
+ Unio difficilis Lea, 1868
+ Unio sparus Lea, 1868

Villosa ortmanni (Walker, 1925)

Villosa taeniata (Conrad, 1834)
+ Unio pictus Lea, 1834
+ Unio punctatus Lea, 1865

Villosa trabalis (Conrad, 1834)
+ Unio perpurpureus Lea, 1861

Villosa vanuxemensis umbrans (Lea, 1857)
+ Unio tenebricus Lea, 1857
+ Unio fabaceus Lea, 1861
+ Unio proprius Lea, 1865

Villosa vanuxemensis vanuxemensis (Lea, 1838)

Villosa vibex (Conrad, 1834)
+ Unio modioliformis Lea, 1834
+ Unio exiguus Lea, 1840
+ Unio stagnalis Conrad, 1849
+ Unio nigrinus Lea, 1852
+ Unio prevostianus Lea, 1852
+ Unio gracilior Lea, 1857
+ Unio obfuscus Lea, 1857
+ Unio rutilans Lea, 1857
+ Unio subellipsis Lea, 1857
+ Unio sudus Lea, 1857
+ Unio dispar Lea, 1860
+ Unio averellii B.H. Wright, 1888

Villosa villosa (Wright, B.H., 1898)
+ Lampsilis (Villosa) wrightiana Frierson, 1927

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