|
Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae tribe PLEUROBEMINI Hannibal, 1912: cladomics
9 genera
Cladomics Summary
number of species = 108
number of subtaxa (rank: genus) = 9
analyzed in 144 cladograms in 87 publications
maximum number of terminals in an analysis = 58, mean = 6.62
maximum number of subtaxa (genus) in an analysis = 9, mean = 3.04
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 100
maximum number of included terminals = 58, mean = 9.09
maximum number of subtaxa (genus) = 9, mean = 3.94
proportion of cladograms that support basic monophyly = 0.78
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 91
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 58, mean = 9.11
maximum number of subtaxa (genus) = 9, mean = 4.1
proportion of cladograms that support rigorous monophyly = 0.76
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
36 |
35 |
9 |
18 |
18 |
4 |
0.0 |
*** Top Monophyly Cladogram ***
+-------------------------------- Quadrulini
|
| +-------------- Elliptoideus **
| +--------------+
| | | +----------- Plectomerus
--+ | +--+
| | | +-------- Obovaria_unicolor
| | +--+
| | | +----- Obovaria_olivaria
| | +--+
| | | +-- Lampsilis
+--+ +--+
| +-- Lemiox
|
| +-------- Cyrtonaias
| +-----------------+
| | | +----- Reginaia
| | +--+
| | | +-- Popenaiadini
+--+ +--+
| +-- Amblema
|
| +----------------------- Elliptio **
| |
+--+ +-------------------- Eurynia **
| |
| | +----------------- Parvaspina **
+--+ |
| | +-- Hemistena **
| +--------------+
+--+ +-- Plethobasus **
|
| +-------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum **
+--+ +--------+
| | +-- Fusconaia **
| |
+--+ +-- Pleurobema_stabile **
| +-----+
| | +-- Pleurobema_sintoxia **
+--+
| +----- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
+--+
+-- Pleurobema_pyriforme **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
28 |
9 |
72 |
60 |
4 |
44.0 |
*** Top Sister-Group Cladogram ***
+-------------------- Quadrulini
|
| +-- Plectomerus
| +-----+
--+ | +-- Reginaia
| +--------+
| | | +----- Lampsilini
| | +--+
| | | +-- Popenaiadini
+--+ +--+
| +-- Amblema
|
| +-------------- Elliptoideus **
| |
+--+ +----------- Hemistena **
| |
| | +-- Plethobasus **
+--+ +-----+
| | +-- Elliptio **
| |
+--+ +----- Pleuronaia **
| |
| +----- Fusconaia **
+--+
+----- Pleurobema **
|
| +-- Eurynia **
+--+
+-- Parvaspina **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
58 |
50 |
9 |
46 |
24 |
4 |
136.0 |
+----------------------------------------------- Reginaia
|
+----------------------------------------------- Amblema
|
| +----- Popenaiadini
| |
| | +-- Dromus
| | |
| | +-- Epioblasma
| | |
+-----------------------------------------+ +-- Glebula
| | |
| | +-- Lampsilis_ovata
| | |
| | +-- Lampsilis_virescens
| | |
| | +-- Lemiox
--+ +--+
| +-- Obliquaria
| |
| +-- Obovaria_olivaria
| |
| +-- Obovaria_unicolor
| |
| +-- Plectomerus
| |
| +-- Potamilus_alatus
| |
| +-- Potamilus_capax
|
| +-------------------------------------------- Quadrulini
| |
| | +----------------------------------------- Elliptoideus **
| | |
| | | +-- Elliptio_lanceolata **
+--+ | +-----------------------------------+
| | | +-- Parvaspina **
| | |
| | | +----------------------------------- Plethobasus **
| | | |
| | | +----------------------------------- Eurynia **
+--+ | |
| | +----------------------------------- Hemistena **
| | |
| | | +-- Elliptio_complanata **
| | | |
| | | +-- Elliptio_hopetonensis **
| | | |
| | | +-- Elliptio_producta__1 **
| | | |
| | | +-- Elliptio_shepardiana **
+--+ | |
| | +-- Elliptio_angustata **
| | |
| | +-- Elliptio_arca **
| | |
| | +-- Elliptio_arctata **
| | |
| | +-- Elliptio_jayensis **
| | |
| | +-- Elliptio_crassidens **
| | |
| +--------------------------------+-- Elliptio_dariensis **
| | |
| | +-- Elliptio_fisheriana **
| | |
| | +-- Elliptio_folliculata **
| | |
+--+ +-- Elliptio_fumata **
| |
| +-- Elliptio_icterina **
| |
| +-- Elliptio_mcmichaeli **
| |
| +-- Elliptio_producta__2 **
| |
| +-- Elliptio_pullata **
| |
| +-- Elliptio_purpurella **
| |
| +-- Elliptio_congaraea **
|
| +-------------------------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum **
| | +--------------------------+
| | | +-- Fusconaia **
| | |
| | | +-------------------------- Pleurobema_plenum **
| | | |
| | | | +----------------------- Pleurobema_stabile **
| | | | |
| | | | | +----- Pleurobema_pyriforme **
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum__2 **
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni **
| | | | | | |
+--+ | +--+ | | +----- Pleurobema_decisum **
| | | | | |
| | | | +--+ +-- Pleurobema_georgianum__1 **
| | | | | |
| | | | +--+-- Pleurobema_hanleyianum__1 **
| | | | |
| | | | +-- Pleurobema_hanleyianum__2 **
| | | |
| | +--+ +-------------- Pleurobema_beadleianum__2 **
| | | |
| | | +--+ +----------- Pleurobema_strodeanum__2 **
| | | | | |
| | | | +--+----------- Pleurobema_rubellum **
| | | | |
| | | | | +-------- Pleurobema_beadleianum__1 **
| | | | +--+
+--+ | | | +----- Pleurobema_taitianum **
| | | +--+
| | | | +-- Pleurobema_perovatum **
| +--+ +--+
| | +-- Pleurobema_strodeanum__1 **
| |
| | +----- Pleurobema_oviforme__6 **
| | |
| | +--+----- Pleurobema_clava__3 **
| | | |
| | | | +-- Pleurobema_oviforme__5 **
| | | +--+
| +--------+ +-- Pleurobema_clava__2 **
| |
| | +----- Pleurobema_oviforme__2 **
| | |
| | +----- Pleurobema_oviforme__3 **
| +--+
| +----- Pleurobema_oviforme__4 **
| |
| | +-- Pleurobema_clava__1 **
| +--+
| +-- Pleurobema_oviforme__1 **
|
| +----------------- Pleurobema_riddellii **
| |
| | +-------------- Pleurobema_sintoxia__34 **
| | |
| | | +----------- Pleurobema_sintoxia__31 **
| | | |
+-----------+ | +----------- Pleurobema_sintoxia__32 **
| | |
| | +----------- Pleurobema_sintoxia__33 **
| | |
| | | +-------- Pleurobema__4 **
| | | |
| | | +-------- Pleurobema__5 **
+--+ | |
| | +-------- Pleurobema_sintoxia__2 **
| | |
| | +-------- Pleurobema_sintoxia__3 **
| | |
| | +-------- Pleurobema__8 **
| | |
| | +-------- Pleurobema__9 **
| | |
| | +-------- Pleurobema__10 **
| | |
| | +-------- Pleurobema_sintoxia__5 **
+--+ |
| +-------- Pleurobema_sintoxia__6 **
| |
| +-------- Pleurobema_sintoxia__7 **
| |
| +-------- Pleurobema_sintoxia__8 **
| |
| +-------- Pleurobema_sintoxia__9 **
| |
| +-------- Pleurobema_sintoxia__10 **
| |
| +-------- Pleurobema_sintoxia__11 **
| |
| +-------- Pleurobema_sintoxia__12 **
| |
| +-------- Pleurobema_sintoxia__13 **
| |
| +-------- Pleurobema_sintoxia__14 **
| |
| +-------- Pleurobema_sintoxia__15 **
| |
| +-------- Pleurobema_sintoxia__16 **
+--+
+-------- Pleurobema_sintoxia__17 **
|
+-------- Pleurobema_sintoxia__18 **
|
+-------- Pleurobema_sintoxia__19 **
|
+-------- Pleurobema_sintoxia__20 **
|
+-------- Pleurobema_sintoxia__21 **
|
+-------- Pleurobema_sintoxia__22 **
|
+-------- Pleurobema_sintoxia__23 **
|
+-------- Pleurobema_sintoxia__24 **
|
+-------- Pleurobema_sintoxia__25 **
|
+-------- Pleurobema_sintoxia__26 **
|
+-------- Pleurobema_sintoxia__27 **
|
+-------- Pleurobema_sintoxia__28 **
|
+-------- Pleurobema_sintoxia__29 **
|
+-------- Pleurobema_sintoxia__30 **
|
| +-- Pleurobema__6 **
+-----+
| +-- Pleurobema_sintoxia__1 **
|
| +-- Pleurobema__7 **
+-----+
| +-- Pleurobema_sintoxia__4 **
|
| +----- Pleurobema__3 **
+--+
| +-- Pleurobema__1 **
+--+
+-- Pleurobema__2 **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
35 |
9 |
32 |
32 |
4 |
50.0 |
+-------------------------------- Quadrulini
|
| +-- Plectomerus
| +-----+
| | +-- Reginaia
--+ +--------------------+
| | | +----- Lampsilini
| | +--+
| | | +-- Popenaiadini
| | +--+
| | +-- Amblema
+--+
| +-------------------------- Elliptoideus **
| |
| | +----------------------- Pleurobema_stabile **
| | |
| | | +----- Pleurobema_riddellii **
+--+ | +-----------+
| | | | +-- Pleurobema_sintoxia **
| | | +--+
| | +--+ +-- Pleurobema_cordatum **
| | | |
+--+ | | +-------------- Fusconaia **
| | | |
| | +--+ +----------- Pleuronaia **
| | | |
| | +--+ +-- Eurynia **
| | | +-----+
| | | | +-- Parvaspina **
| | +--+
+--+ | +----- Elliptio **
| +--+
| | +-- Hemistena **
| +--+
| +-- Plethobasus **
|
| +-- Pleurobema_clava **
| +-----------+
| | +-- Pleurobema_oviforme **
| |
+-----+ +----------- Pleurobema_pyriforme **
| |
| | +-- Pleurobema_athearni **
| | +--+
+--+ | +-- Pleurobema_georgianum **
| +--+
| | | +-- Pleurobema_decisum **
| | +--+
| | +-- Pleurobema_hanleyianum **
+--+
| +----- Pleurobema_beadleianum **
| |
| +----- Pleurobema_strodeanum **
+--+
+----- Pleurobema_rubellum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
36 |
9 |
32 |
32 |
4 |
48.0 |
+----------------------------------- Quadrulini
|
| +----------- Elliptoideus **
| |
| | +-------- Elliptio **
| +--------------------+ |
| | | | +-- Eurynia **
--+ | | +-----+
| | | | +-- Parvaspina **
| | +--+
| | | +-- Hemistena **
| | +-----+
| | | +-- Plethobasus **
| | |
+--+ | +----- Pleuronaia **
| +--+
| | +-- Fusconaia **
| +--+
| +-- Pleurobema **
|
| +-- Amblema
| +--------------------------+
| | +-- Reginaia
| |
+--+ +-------------------------- Popenaiadini
| |
| | +-- Glebula
+--+ +--------------------+
| | +-- Paetulunio
| |
+--+ +-- Toxolasma
| +-----------------+
| | +-- Plectomerus
| |
+--+ +----------------- Obliquaria
| |
| | +----- Potamilus
+--+ +--------+
| | | +-- Dromus
| | +--+
+--+ +-- Lemiox
|
| +----------- Lampsilis_ovata
| |
+--+ +-------- Lampsilis_virescens
| |
+--+ +-- Epioblasma
| +--+
| | +-- Obovaria_arkansasensis
+--+
| +-- Obovaria_olivaria
+--+
+-- Venustaconcha
Burlakova et al., 2012 [Publ 3486], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
30 |
8 |
0 |
0 |
0 |
45.0 |
+----------------------------------- Parvaspina **
|
| +-------------------------------- Pleurobema_plenum **
--+ |
| | +-- Plethobasus **
| | +--+
+--+ | +-- Hemistena **
| +-----------------------+
| | | +-- Eurynia **
| | +--+
| | +-- Elliptio **
+--+
| +-------------------------- Pleurobema_cordatum__2 **
| |
| | +----------------------- Fusconaia **
+--+ |
| | +-------- Pleurobema_stabile **
| | |
| | +-----------+ +----- Pleurobema_riddellii **
+--+ | | |
| | +--+ +-- Pleurobema_cordatum__1 **
| | | |
| | | +-- Pleurobema_sintoxia__1 **
| | +--+
+--+ +-- Pleurobema_sintoxia__2 **
| |
| +-- Pleurobema_sintoxia__3 **
|
| +----------------- Pleuronaia **
| |
| | +-- Pleurobema_georgianum **
+--+ +-----------+
| | +-- Pleurobema_pyriforme **
| |
+--+ +-- Pleurobema_decisum **
| +--------+
| | +-- Pleurobema_hanleyianum **
| |
+--+ +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +----- Pleurobema_rubellum **
| |
| +----- Pleurobema_beadleianum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Burlakova et al., 2012 [Publ 3486], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
25 |
8 |
1 |
1 |
1 |
44.0 |
+-------------------------- Amblemini
|
| +----------------------- Hemistena **
--+ |
| +----------------------- Pleurobema_stabile **
| |
+--+ +-------------------- Eurynia **
| |
| | +-------- Pleurobema_riddellii **
| | |
| | +--------+ +----- Pleurobema_sintoxia__2 **
+--+ | | |
| | +--+----- Pleurobema_sintoxia__3 **
| | |
| | | +-- Pleurobema_cordatum **
| | +--+
| | +-- Pleurobema_sintoxia__1 **
+--+
| +-------------- Parvaspina **
| |
| +-------------- Pleuronaia **
| |
| +-------------- Fusconaia **
| |
| | +-- Plethobasus **
+--+-----------+
| +-- Elliptio **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
| |
| | +-------- Pleurobema_decisum **
+--+ |
| +-------- Pleurobema_pyriforme **
| |
| +-------- Pleurobema_hanleyianum **
+--+
+-------- Pleurobema_georgianum **
|
| +----- Pleurobema_rubellum **
| |
| +----- Pleurobema_beadleianum **
+--+
+----- Pleurobema_strodeanum **
|
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Burlakova et al., 2012 [Publ 3486], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
36 |
8 |
6 |
6 |
3 |
50.0 |
+----------------------------------------------- Amblema
|
--+ +-- Lampsilis
| +-----------------------------------------+
| | +-- Plectomerus
+--+
| +----------------------------------------- Popenaiadini
| |
+--+ +-------------------------------------- Cyrtonaias
| |
+--+ +----------------------------------- Parvaspina **
| |
| | +-- Plethobasus **
+--+ +-----------------------------+
| | +-- Elliptio **
| |
+--+ +----------------------------- Fusconaia **
| |
| | +-- Eurynia **
+--+ +-----------------------+
| | +-- Hemistena **
| |
| | +----------- Pleuronaia **
+--+ +-----------+
| | | +-------- Pleurobema_riddellii **
| | +--+
| | | +----- Pleurobema_sintoxia__2 **
| | +--+
+--+ | +-- Pleurobema_sintoxia__1 **
| +--+
| +-- Pleurobema_cordatum **
|
| +-------------------- Pleurobema_stabile **
| |
+--+ +----------------- Pleurobema_pyriforme **
| |
| | +----- Pleurobema_georgianum **
+--+ +--------+
| | | +-- Pleurobema_hanleyianum **
| | +--+
| | +-- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_beadleianum **
| |
+--+ +-- Pleurobema_rubellum **
| +--+
| | +-- Pleurobema_strodeanum **
+--+
| +-- Pleurobema_perovatum **
+--+
+-- Pleurobema_taitianum **
Perkins et al., 2017 [Publ 3777], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
33 |
7 |
2 |
2 |
1 |
57.0 |
+-------------------- Amblema
|
--+-------------------- Plectomerus
|
| +----------------- Pleurobema **
+--+
| +-------------- Pleuronaia **
+--+
| +----------- Fusconaia **
+--+
| +-------- Parvaspina **
+--+
| +----- Elliptio **
+--+
| +-- Eurynia **
+--+
+-- Hemistena **
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
27 |
9 |
72 |
60 |
4 |
39.0 |
+-------------------------------------- Quadrulini
|
| +-------------------------------- Elliptoideus **
| |
| | +----------------------------- Plethobasus **
| +--+ |
| | | +----------------------------- Parvaspina **
| | | |
| | | +----------------------------- Elliptio **
| | +--+
| | | +-------------------------- Eurynia **
| | | |
| | | +-------------------------- Hemistena **
| | | |
| | | | +----------------------- Fusconaia **
| | +--+ |
--+ | | | +----- Pleurobema_sintoxia__2 **
| | | +-----------------+
| | | | | +-- Pleurobema_sintoxia__1 **
| | | | +--+
| | +--+ +-- Pleurobema_cordatum **
| | |
| | | +-------------------- Pleuronaia **
| | | |
| | | | +-------- Pleurobema_rubellum **
| | | | +--------+
| | +--+ | | +----- Pleurobema_strodeanum **
| | | | +--+
| | | | | +-- Pleurobema_perovatum **
| | | | +--+
| | | | +-- Pleurobema_taitianum **
| | +--+
+--+ | +-- Pleurobema_oviforme__1 **
| | |
| | +-----------+-- Pleurobema_oviforme__2 **
| | | |
| | | +-- Pleurobema_clava **
| +--+
| | +----------- Pleurobema_pyriforme **
| | |
| +--+ +-- Pleurobema_athearni **
| | +-----+
| | | +-- Pleurobema_georgianum__2 **
| +--+
| | +----- Pleurobema_decisum **
| | |
| +--+ +-- Pleurobema_hanleyianum__1 **
| | |
| +--+-- Pleurobema_hanleyianum__2 **
| |
| +-- Pleurobema_georgianum__1 **
|
| +----- Popenaiadini
| +--------------+
| | | +-- Amblema
| | +--+
| | +-- Reginaia
| |
| | +-- Plectomerus
| | +--------------+
+--------------+ | +-- Toxolasma
| |
| | +----- Obliquaria
| | +--------+
| | | | +-- Cyrtonaias
| | | +--+
| | | +-- Glebula
| | |
+--+ | +----------- Sagittunio
| | |
| | +----------- Leaunio
| | |
| | +----------- Potamilus
| | |
| | +----------- Hamiota
| | |
| | | +-- Ellipsaria
+--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Burdick & White, 2007 [Publ 2679], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
29 |
30 |
7 |
2 |
2 |
2 |
45.0 |
+-------------------------------- Lampsilini
|
--+ +----------------------------- Parvaspina **
| |
| | +-------------------------- Amblemini
+--+ |
| | +----- Eurynia **
| | +-----------------+
+--+ | | +-- Plethobasus **
| | +--+
| | +-- Elliptio **
| |
| | +-------------------- Pleurobema_plenum **
+--+ |
| +-------------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum__1 **
| | +-----+
| | | +-- Fusconaia **
+--+ +--------+
| | | +----- Pleurobema_sintoxia__1 **
| | | |
| | +--+ +-- Pleurobema_cordatum__2 **
| | | |
| | +--+-- Pleurobema_sintoxia__2 **
| | |
+--+ +-- Pleurobema_sintoxia__3 **
|
| +-------------- Pleurobema_stabile **
| |
| | +-- Pleurobema_rubellum **
| | +-----+
| | | +-- Pleurobema_strodeanum **
+--+ +--+
| | | +----- Pleurobema_beadleianum **
| | +--+
| | | +-- Pleurobema_perovatum **
| | +--+
+--+ +-- Pleurobema_taitianum **
|
| +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +-- Pleurobema_decisum **
| +--+
| | +-- Pleurobema_hanleyianum **
+--+
| +-- Pleurobema_pyriforme **
+--+
+-- Pleurobema_georgianum **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
24 |
9 |
17 |
17 |
4 |
43.0 |
+-------------------------- Quadrulini
|
| +-- Popenaiadini
| +-----------+
| | +-- Amblema
| |
--+ +--------+ +-- Cyrtonaias
| | | +--------+
| | | | +-- Reginaia
| | +--+
| | | +-------- Plectomerus
| | +--+
| | | +----- Lemiox
+--+ +--+
| | +-- Lampsilis
| +--+
| +-- Obovaria
|
| +-------------------- Elliptoideus **
| |
| | +----------------- Pleurobema_stabile **
+--+ |
| | +----- Pleurobema_pyriforme **
| | +--------+
+--+ | | +-- Pleurobema_clava **
| | +--+
| | +-- Pleurobema_decisum **
| |
+--+ +-- Pleurobema_cordatum **
| +--------+
| | +-- Pleurobema_sintoxia **
| |
| | +----- Pleuronaia **
+--+ +--+
| | | +-- Parvaspina **
| | +--+
| | +-- Fusconaia **
+--+
| +-- Eurynia **
| +--+
| | +-- Hemistena **
+--+
| +-- Plethobasus **
+--+
+-- Elliptio **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
24 |
9 |
17 |
17 |
4 |
43.0 |
+-------------------------------- Quadrulini
|
| +-- Popenaiadini
| +--------+
| | +-- Amblema
| |
--+ +-----------------+ +-------- Plectomerus
| | | |
| | | | +-- Cyrtonaias
| | +--+-----+
| | | +-- Reginaia
| | |
| | | +----- Lemiox
+--+ +--+
| | +-- Lampsilis
| +--+
| +-- Obovaria
|
| +-------------------------- Elliptoideus **
| |
+--+ +----------------------- Pleurobema_stabile **
| |
+--+ +-------------------- Hemistena **
| |
+--+ +----------------- Eurynia **
| |
| | +-- Pleurobema_cordatum **
+--+ +-----------+
| | +-- Pleurobema_sintoxia **
| |
| | +----- Pleurobema_clava **
+--+ +-----+
| | | +-- Pleurobema_decisum **
| | +--+
| | +-- Pleurobema_pyriforme **
+--+
| +-- Parvaspina **
| +-----+
| | +-- Pleuronaia **
+--+
| +----- Fusconaia **
+--+
| +-- Plethobasus **
+--+
+-- Elliptio **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
20 |
9 |
17 |
17 |
4 |
39.0 |
+-------------------- Quadrulini
|
| +-- Reginaia
| +-----------+
| | +-- Amblema
| +--+
--+ | | +----------- Popenaiadini
| | | |
| | +--+ +-------- Cyrtonaias
| | | |
| | +--+-------- Plectomerus
| | |
| | | +----- Lemiox
+--+ +--+
| | +-- Lampsilis
| +--+
| +-- Obovaria
|
| +-------- Elliptoideus **
| |
| | +----- Hemistena **
| | |
+--------+ +----- Plethobasus **
| |
| +----- Elliptio **
| |
+--+ +-- Eurynia **
+--+
| +-- Parvaspina **
|
| +-- Pleuronaia **
| |
+--+-- Pleurobema **
|
+-- Fusconaia **
Campbell et al., 2008 [Publ 2675], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
22 |
8 |
2 |
2 |
2 |
52.0 |
+-------------------- Lampsilini
|
+-------------------- Amblemini
--+
| +----------------- Parvaspina **
| |
+--+ +-------------- Hemistena **
| |
+--+ +-- Plethobasus **
| +--------+
| | +-- Elliptio **
+--+
| +-------- Fusconaia **
+--+
| +----- Pleurobema **
+--+
| +-- Eurynia **
+--+
+-- Pleuronaia **
Campbell et al., 2008 [Publ 2675], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
17 |
8 |
2 |
2 |
2 |
48.0 |
+----------------------- Lampsilini
|
+----------------------- Amblemini
|
| +-- Pleurobema_cordatum **
| |
| +--------+-- Pleurobema_sintoxia__1 **
| | |
| | +-- Pleurobema_sintoxia__2 **
--+ +--------+
| | | +-------- Fusconaia **
| | | |
| | +--+ +----- Pleuronaia **
| | | |
| | | | +-- Eurynia **
| | +--+ |
| | | +-- Hemistena **
| | | |
+--+ +--+-- Plethobasus **
| |
| +-- Parvaspina **
| |
| +-- Elliptio **
|
| +----------------- Pleurobema_stabile **
| |
| | +-------------- Pleurobema_pyriforme **
| | |
+--+ | +-- Pleurobema_georgianum__2 **
| +-----------+
| | +-- Pleurobema_athearni **
| |
| | +----- Pleurobema_decisum **
+--+--------+
| | +-- Pleurobema_hanleyianum **
| +--+
| +-- Pleurobema_georgianum__1 **
|
| +-- Pleurobema_clava **
| +--------+
| | +-- Pleurobema_oviforme **
+--+
| +-------- Pleurobema_rubellum **
| |
+--+-------- Pleurobema_strodeanum **
|
| +----- Pleurobema_beadleianum **
+--+
| +-- Pleurobema_taitianum **
+--+
+-- Pleurobema_perovatum **
Campbell et al., 2008 [Publ 2675], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
23 |
5 |
8 |
3 |
3 |
3 |
45.0 |
+----------- Quadrulini
|
+----------- Amblemini
|
| +-- Elliptio_arctata **
| +-----+
| | +-- Elliptio_crassidens **
| |
| | +----- Pleurobema_decisum **
--+ | |
| | +----- Pleurobema_rubellum **
| | |
| | +----- Pleurobema_georgianum__2 **
| | |
| | +----- Pleurobema_stabile **
| | |
| | +----- Pleurobema_pyriforme **
| | |
+--+ +----- Pleurobema_oviforme__2 **
| |
| +----- Pleurobema_hanleyianum__2 **
| |
| +----- Pleuronaia **
| |
| +----- Pleurobema_georgianum__3 **
| |
| +----- Pleurobema_cordatum **
| |
| +----- Pleurobema_beadleianum **
| |
| +----- Hemistena **
+--+
+----- Fusconaia_flava **
|
+----- Fusconaia_cerina **
|
+----- Eurynia **
|
+----- Elliptio_arca **
|
+----- Lampsilini
|
| +-- Plethobasus **
+--+
| +-- Parvaspina **
|
| +-- Pleurobema_sintoxia **
| |
+--+-- Pleurobema_oviforme__1 **
| |
| +-- Pleurobema_clava **
|
| +-- Pleurobema_athearni **
| |
+--+-- Pleurobema_georgianum__1 **
|
+-- Pleurobema_hanleyianum__1 **
Perkins et al., 2017 [Publ 3777], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
9 |
7 |
2 |
2 |
2 |
45.0 |
+-------------------- Amblemini
|
+-------------------- Quadrulini
|
--+ +----------------- Parvaspina **
| |
| | +-------------- Elliptio_arca **
| | |
+--+ +-------------- Hemistena **
| |
| | +-- Elliptio_arctata **
| +-----------+
| | +-- Elliptio_crassidens **
+--+
| +----------- Eurynia **
| |
| | +-------- Pleurobema_oviforme **
| | |
| | +-------- Pleurobema_clava **
| | |
+--+ +-------- Pleurobema_pyriforme **
| |
| +-------- Pleurobema_georgianum **
| |
| +-------- Pleurobema_beadleianum **
| |
+--+-------- Pleurobema_hanleyianum **
|
+-------- Pleurobema_decisum **
|
| +-- Pleurobema_sintoxia **
+-----+
| +-- Pleurobema_rubellum **
|
| +----- Elliptio_spinosa **
| |
+--+ +-- Fusconaia_escambia **
| |
| +-- Fusconaia_flava **
+--+
+-- Fusconaia_cerina **
|
+-- Pleuronaia **
Campbell et al., 2008 [Publ 2675], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
23 |
8 |
8 |
3 |
3 |
3 |
42.0 |
+----------------------- Quadrulini
|
+----------------------- Amblemini
|
| +-------------------- Lampsilini
| |
| +-------------------- Eurynia **
| |
| +-------------------- Fusconaia_cerina **
--+ |
| +-------------------- Fusconaia_flava **
| |
| +-------------------- Hemistena **
| |
| +-------------------- Plethobasus **
| |
| +-------------------- Parvaspina **
| |
+--+-------------------- Pleurobema_cordatum **
|
+-------------------- Pleuronaia **
|
+-------------------- Pleurobema_oviforme__2 **
|
+-------------------- Pleurobema_stabile **
|
+-------------------- Elliptio **
|
| +----------------- Pleurobema_decisum **
| |
| +----------------- Pleurobema_sintoxia **
| |
| +----------------- Pleurobema_clava **
+--+
+----------------- Pleurobema_oviforme__1 **
|
| +-------------- Pleurobema_beadleianum **
| |
+--+ +----------- Pleurobema_pyriforme **
| |
+--+ +-------- Pleurobema_rubellum **
| |
| +-------- Pleurobema_hanleyianum__2 **
+--+
+-------- Pleurobema_georgianum__2 **
|
| +----- Pleurobema_hanleyianum__1 **
+--+
| +-- Pleurobema_georgianum__1 **
+--+
+-- Pleurobema_athearni **
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
27 |
23 |
9 |
17 |
16 |
3 |
45.0 |
+----------------------------- Amblemini
|
--+ +-------------------------- Lampsilini
| |
+--+ +----------------------- Elliptoideus **
| |
+--+ +-------------------- Quadrulini
| |
+--+ +----------------- Elliptio **
| |
| | +-------------- Parvaspina **
+--+ |
| +-------------- Plethobasus **
| |
| | +-- Hemistena **
+--+-----------+
| +-- Eurynia **
|
| +----------- Fusconaia **
| |
| | +----- Pleurobema_riddellii **
+--+ +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
+--+ +-- Pleurobema_cordatum **
|
| +----- Pleuronaia **
| |
+--+ +-- Pleurobema_pyriforme **
| |
+--+-- Pleurobema_strodeanum **
|
+-- Pleurobema_clava **
Smith et al., 2020 [Publ 4136], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
12 |
9 |
1 |
1 |
1 |
41.0 |
+----------------- Quadrulini
|
--+ +-------------- Elliptoideus **
| |
+--+ +----------- Elliptio **
| |
| +----------- Plethobasus **
+--+
| +-------- Parvaspina **
| |
| | +-- Hemistena **
+--+-----+
| +-- Eurynia **
|
| +----- Fusconaia **
+--+
| +-- Pleuronaia **
+--+
+-- Pleurobema **
Pieri et al., 2018 [Publ 3867], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
20 |
3 |
3 |
2 |
1 |
38.0 |
+----- Quadrulini
|
--+ +-- Eurynia **
| |
+--+-- Pleurobema **
|
+-- Fusconaia **
Smith et al., 2020 [Publ 4136], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
2 |
0 |
0 |
0 |
42.0 |
+-- Pleurobema **
--+
+-- Fusconaia **
Morrison et al., 2021 [Publ 4362], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
2 |
0 |
0 |
0 |
47.0 |
+-------------------- Fusconaia **
|
--+ +-- Pleurobema_clava **
| +--------------+
| | +-- Pleurobema_oviforme **
+--+
| +-------------- Pleurobema_decisum **
| |
+--+ +----------- Pleurobema_hanleyianum **
| |
+--+ +-- Pleurobema_taitianum **
| +-----+
| | +-- Pleurobema_perovatum **
+--+
| +----- Pleurobema_rubellum **
+--+
| +-- Pleurobema_strodeanum **
+--+
+-- Pleurobema_beadleianum **
Morrison et al., 2021 [Publ 4362], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
2 |
0 |
0 |
0 |
47.0 |
+-- Fusconaia **
--+
+-- Pleurobema **
Inoue et al., 2020 [Publ 3941], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
6 |
34 |
33 |
3 |
41.0 |
+----------------- Quadrulini
|
--+ +-- Amblemini
| +-----------+
| | +-- Lampsilini
+--+
| +----------- Parvaspina **
| |
+--+ +-- Eurynia **
| +-----+
| | +-- Hemistena **
+--+
| +----- Pleurobema **
+--+
| +-- Pleuronaia **
+--+
+-- Fusconaia **
Inoue et al., 2020 [Publ 3941], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
6 |
33 |
32 |
3 |
41.0 |
+-------------------- Quadrulini
|
--+ +----------------- Reginaia
| |
+--+ +-- Amblema
| +-----------+
| | +-- Lampsilini
+--+
| +----------- Parvaspina **
| |
+--+ +-- Eurynia **
| +-----+
| | +-- Hemistena **
+--+
| +----- Pleurobema **
+--+
| +-- Pleuronaia **
+--+
+-- Fusconaia **
Kandl et al., 2001 [Publ 3384], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
6 |
3 |
2 |
2 |
2 |
40.0 |
+-------------------- Amblemini
|
+-------------------- Quadrulini
--+
| +----------------- Elliptio **
| |
+--+ +-------------- Fusconaia **
| |
+--+ +----------- Pleurobema_sintoxia **
| |
| | +-- Pleurobema_clava **
+--+ +-----+
| | +-- Pleurobema_strodeanum **
| |
+--+ +----- Pleurobema_pyriforme__6 **
| |
| | +-- Pleurobema_pyriforme__1 **
+--+ |
| +-- Pleurobema_pyriforme__2 **
| |
+--+-- Pleurobema_pyriforme__3 **
|
+-- Pleurobema_pyriforme__4 **
|
+-- Pleurobema_pyriforme__5 **
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
4 |
47 |
3 |
3 |
45.0 |
+-------- Pleurobema **
--+
| +----- Fusconaia **
+--+
| +-- Elliptio **
+--+
+-- Eurynia **
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
5 |
29 |
28 |
4 |
42.0 |
+----------------------------------- Quadrulini
|
| +-------------- Elliptoideus **
| +-----------------+
| | | +----------- Elliptio **
--+ | +--+
| | | +-------- Pleuronaia **
| | +--+
| | | +----- Pleurobema_riddellii **
| | +--+
+--+ | +-- Fusconaia **
| +--+
| +-- Pleurobema_strodeanum **
|
| +----------------------------- Amblemini
| |
| | +-- Popenaias
+--+ +-----------------------+
| | +-- Psoronaias
| |
+--+ +-- Cyrtonaias_tampicoensis
| +--------------------+
| | +-- Glebula
| |
+--+ +-------------------- Potamilus
| |
| | +-- Truncilla
+--+ +--------------+
| | +-- Ptychobranchus
| |
+--+ +-------------- Pachynaias
| |
| | +-- Cyrtonaias_explicata
+--+ +--------+
| | +-- Sphenonaias
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Ortmanniana
+--+
| +----- Cambarunio
+--+
| +-- Epioblasma
+--+
+-- Ligumia
Serb et al., 2003 [Publ 2659], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
27 |
18 |
4 |
24 |
24 |
3 |
57.0 |
+----------- Lampsilis
|
| +-------- Obliquaria
| |
| +-------- Amblema
| |
--+ +-------- Reginaia
| |
| +-------- Potamilus
| |
| +-------- Quadrulini
| |
+--+ +-- Elliptoideus **
+-----+
| +-- Plectomerus
|
| +----- Medionidus
+--+
| | +-- Dromus
| +--+
| +-- Cyprogenia
|
| +----- Hemistena **
+--+
| +-- Elliptio **
+--+
+-- Fusconaia **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
191 |
20 |
3 |
42.0 |
+----------------------------------- Quadrulini
|
| +-------- Elliptoideus **
| |
--+ +-----------------------+ +-- Elliptio **
| | | +--+
| | | | +-- Eurynia **
| | +--+
| | | +-- Fusconaia **
+--+ +--+
| +-- Pleurobema **
|
| +----------------------------- Amblema
| |
+--+ +-------------------------- Toxolasma
| |
+--+ +-- Obliquaria
| +--------------------+
| | +-- Plectomerus
+--+
| +-------------------- Potamilus_leptodon
| |
+--+ +----------------- Potamilus_alatus
| |
| | +-- Medionidus
+--+ +-----------+
| | +-- Ptychobranchus
| |
+--+ +-- Ellipsaria
| +--------+
| | +-- Truncilla
+--+
| +-------- Leaunio
| |
+--+ +-- Venustaconcha
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
5 |
52 |
50 |
3 |
44.0 |
+-- Amblemini
|
+--------+-- Popenaiadini
| |
| +-- Lampsilini
--+
| +-------- Pleuronaia **
| |
+--+-------- Pleurobema **
|
| +----- Fusconaia **
+--+
| +-- Hemistena **
+--+
+-- Eurynia **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
193 |
20 |
3 |
40.0 |
+-------------- Amblema
|
+-------------- Toxolasma
|
| +-- Obliquaria
+-----------+
| +-- Plectomerus
|
| +----------- Elliptoideus **
+--+
| | +-------- Elliptio **
| +--+
| | +----- Eurynia **
--+ +--+
| | +-- Fusconaia **
| +--+
| +-- Pleurobema **
|
| +----- Ellipsaria
| |
| +----- Potamilus_leptodon
| |
| +----- Medionidus
| |
| +----- Potamilus_alatus
| |
| +----- Ptychobranchus
+--------+
+----- Truncilla
|
+----- Venustaconcha
|
+----- Cambarunio
|
+----- Leaunio
|
| +-- Ortmanniana
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
193 |
20 |
3 |
42.0 |
+-------------------------- Quadrulini
|
| +-------- Elliptoideus **
| |
--+ +--------------+ +-- Elliptio **
| | | +--+
| | | | +-- Eurynia **
| | +--+
| | | +-- Fusconaia **
+--+ +--+
| +-- Pleurobema **
|
| +-------------------- Amblema
| |
| | +----- Toxolasma
+--+ +-----------+
| | | +-- Obliquaria
| | +--+
| | +-- Plectomerus
+--+
| +-------------- Potamilus_leptodon
| |
| | +----------- Potamilus_alatus
+--+ |
| | +-------- Venustaconcha
| | |
+--+ +-------- Leaunio
| |
| | +-- Ellipsaria
| +-----+
+--+ +-- Truncilla
|
| +-- Medionidus
+-----+
| +-- Ptychobranchus
|
| +----- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Serb et al., 2003 [Publ 2659], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
21 |
19 |
4 |
24 |
24 |
3 |
51.0 |
+-------------------------- Lampsilis
|
--+ +----------------------- Potamilus
| |
| | +----- Medionidus
+--+ +--------------+
| | | +-- Dromus
| | +--+
+--+ +-- Cyprogenia
|
| +----------------- Obliquaria
| |
+--+ +-------------- Amblema
| |
| | +----- Reginaia
+--+ +-----+
| | | +-- Elliptoideus **
| | +--+
+--+ +-- Plectomerus
|
| +-------- Quadrulini
+--+
| +----- Hemistena **
+--+
| +-- Elliptio **
+--+
+-- Fusconaia **
Kandl et al., 2001 [Publ 3384], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
6 |
4 |
2 |
1 |
1 |
1 |
42.5 |
+-- Elliptio **
|
--+-- Quadrulini
|
+-- Pleurobema **
Lydeard et al., 2000 [Publ 2654], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
4 |
2 |
10 |
8 |
3 |
39.0 |
+----------- Lampsilini
--+
| +-------- Amblemini
+--+
| +----- Quadrulini
+--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
4 |
52 |
50 |
3 |
42.0 |
+-------- Lampsilini
|
| +-- Amblemini
--+ +--+
| | +-- Popenaiadini
| |
+--+ +-- Pleurobema **
| |
| +-- Eurynia **
+--+
+-- Pleuronaia **
|
+-- Fusconaia **
Lydeard et al., 2000 [Publ 2654], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
4 |
2 |
10 |
8 |
3 |
39.0 |
+----------- Amblemini
--+
| +-------- Lampsilini
+--+
| +----- Quadrulini
+--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
4 |
2 |
10 |
8 |
3 |
39.0 |
+----- Quadrulini
--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Lydeard et al., 2000 [Publ 2654], fig. 2c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
10 |
8 |
3 |
39.0 |
+----- Amblemini
--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Boyer et al., 2011 [Publ 2718], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
31 |
25 |
3 |
42.0 |
+-------------- Reginaia
|
--+ +-- Amblema
| +--------+
| | +-- Lampsilini
+--+
| +-------- Quadrulini
| |
+--+ +-- Fusconaia **
| +--+
| | +-- Pleurobema **
+--+
| +-- Eurynia **
+--+
+-- Elliptio **
Boyer et al., 2011 [Publ 2718], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
34 |
25 |
3 |
44.0 |
+-------------------------------- Quadrulina
|
--+ +----------------------------- Megalonaias
| |
+--+ +-------------------------- Toxolasma
| |
| | +----------------------- Amblema
+--+ |
| | +-- Reginaia
| | +-----------------+
| | | +-- Obliquaria
+--+ |
| | +-------- Pleurobema **
| +-----------+
| | | +----- Elliptio **
| | +--+
+--+ | +-- Fusconaia **
| +--+
| +-- Eurynia **
|
| +----------------- Potamilus
| |
| | +-- Ellipsaria
+--+ +-----------+
| | +-- Truncilla
| |
+--+ +----------- Lampsilis_cardium
| |
| | +----- Ligumia
| +-----+
+--+ | +-- Obovaria
| +--+
| +-- Epioblasma
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis_higginsii
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
5 |
3 |
4 |
29 |
17 |
3 |
43.0 |
+-- Megalonaias
+-----------------------+
| +-- Quadrulina
|
| +----------- Elliptoideus **
| +-----------+
--+ | | +-------- Pleurobema **
| | +--+
| | | +----- Uniomerus
| | +--+
| | | +-- Fusconaia **
+--+ +--+
| +-- Eurynia **
|
| +-------------------- Amblema
| |
| | +----- Toxolasma
| | +-----------+
+--+ | | +-- Plectomerus
| | +--+
| | +-- Obliquaria
| |
+--+ +----- Ptychobranchus
| +--------+
| | | +-- Potamilus_fragilis
| | +--+
| | +-- Medionidus
+--+
| +-- Villosa
| +--------+
| | +-- Ellipsaria
+--+
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Potamilus_purpuratus
+--+
+-- Potamilus_alatus
Johnson et al., 2023 [Publ 4323], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
44.0 |
+-- Fusconaia **
--+
+-- Pleurobema **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
12 |
4 |
43.0 |
+----------- Quadrulini
|
| +----- Elliptoideus **
--+ +--+
| | | +-- Elliptio **
| | +--+
+--+ +-- Pleurobema **
|
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
17 |
0 |
3 |
19 |
9 |
3 |
49.0 |
+-- Gonideini
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea
|
+-- Lampsilis_teres
|
+-- Megalonaias
|
+-- Obliquaria
|
--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio **
|
+-- Fusconaia **
|
+-- Plectomerus
|
+-- Pleurobema **
|
+-- Unionini
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
28 |
8 |
3 |
41.0 |
+-- Glebula
+--------------+
| +-- Cyrtonaias
|
--+ +-------------- Toxolasma
| |
| | +----- Potamilus
| +--------+
+--+ | +-- Ligumia
| +--+
| +-- Ortmanniana
|
| +----------- Amblemini
+--+
| +-------- Quadrulini
+--+
| +----- Eurynia **
+--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
28 |
8 |
3 |
41.0 |
+-------------- Amblemini
--+
| +----------- Lampsilini
+--+
| +-------- Quadrulini
+--+
| +----- Eurynia **
+--+
| +-- Fusconaia **
+--+
+-- Pleurobema **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
0 |
3 |
16 |
8 |
3 |
44.0 |
+-- Elliptio **
|
+-- Fusconaia **
|
--+-- Amblemini
|
+-- Pleurobema **
|
+-- Unionini
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
44 |
11 |
3 |
41.0 |
+-------------- Quadrulini
--+
| +----------- Amblemini
+--+
| +-------- Lampsilini
+--+
| +----- Eurynia **
+--+
| +-- Fusconaia **
+--+
+-- Pleurobema **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
27 |
14 |
3 |
41.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Eurynia **
+--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
43 |
10 |
3 |
41.0 |
+-- Obliquaria
+--------+
| +-- Quadrulini
|
| +-------- Amblemini
--+ |
| | +----- Eurynia **
| +--+
| | | +-- Pleurobema **
+--+ +--+
| +-- Fusconaia **
|
| +----- Truncilla
| |
| +----- Ptychobranchus
+--+
| +-- Ortmanniana
| |
| +-- Cambarunio
+--+
+-- Ligumia
|
+-- Epioblasma
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
77 |
6 |
3 |
42.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Elliptio **
+--+
| +-- Pleurobema **
+--+
+-- Eurynia **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
73 |
6 |
3 |
42.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Elliptio **
+--+
| +-- Pleurobema **
+--+
+-- Eurynia **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
28 |
16 |
3 |
28 |
8 |
3 |
59.0 |
+-------------------- Amblemini
|
+-------------------- Coelaturini
|
+-------------------- Eurynia **
|
+-------------------- Fusconaia **
|
+-------------------- Gonideini
|
+-------------------- Pleurobema **
|
+-------------------- Quadrulini
--+
| +----------- Ortmanniana
+--------+
| | +-------- Glebula
| +--+
| | +----- Potamilus
| +--+
| | +-- Ligumia
| +--+
| +-- Toxolasma
|
| +----------------- Cyrtonaias
| |
+--+ +-------------- Unionini
| |
| | +----------- Anodontini
+--+ |
| | +----- Mutela
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaea
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
27 |
14 |
3 |
41.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Eurynia **
+--+
| +-- Pleurobema **
+--+
+-- Fusconaia **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
12 |
4 |
43.0 |
+-------------- Quadrulini
|
--+ +-- Popenaiadini
| +--------+
| | +-- Lampsilini
+--+
| +-------- Amblemini
+--+
| +----- Elliptoideus **
+--+
| +-- Elliptio **
+--+
+-- Pleurobema **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
16 |
12 |
3 |
28 |
8 |
3 |
52.0 |
+----------------------- Fusconaia **
|
+----------------------- Amblemini
--+
| +-------------------- Quadrulini
| |
+--+ +----------------- Pleurobema **
| |
| | +-------------- Eurynia **
+--+ |
| | +----- Cyrtonaias
| +--------+
+--+ | +-- Unionini
| +--+
| +-- Anodontini
|
| +----------- Ortmanniana
+--+
| +-------- Glebula
+--+
| +----- Potamilus
+--+
| +-- Ligumia
+--+
+-- Toxolasma
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
136 |
6 |
3 |
44.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Pleurobema **
+--+
| +-- Eurynia **
+--+
+-- Elliptio **
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
44 |
11 |
3 |
41.0 |
+----------- Amblemini
|
+--------+ +-- Obliquaria
| | +-----+
| | | +-- Quadrulini
| +--+
| | +----- Eurynia **
--+ +--+
| | +-- Pleurobema **
| +--+
| +-- Fusconaia **
|
| +----------------- Ptychobranchus
+--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
3 |
31 |
8 |
3 |
42.0 |
+-------- Toxolasma
|
+-------- Amblemini
|
| +-- Glebula
+-----+
| +-- Cyrtonaias
|
--+ +----- Potamilus
+--+
| | +-- Ortmanniana
| +--+
| +-- Ligumia
|
| +----- Quadrulini
| |
+--+ +-- Eurynia **
| |
+--+-- Pleurobema **
|
+-- Fusconaia **
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
1 |
3 |
24 |
15 |
3 |
42.0 |
+-- Megalonaias
+-----------------------+
| +-- Quadrulina
|
| +----- Elliptoideus **
--+ |
| +-----------------+ +-- Eurynia **
| | | |
| | +--+-- Pleurobema **
| | |
+--+ +-- Uniomerus
|
| +-------------------- Amblema
| |
+--+ +-- Plectomerus
| +--------------+
| | +-- Obliquaria
+--+
| +-------------- Potamilus_fragilis
| |
+--+ +----------- Ptychobranchus
| |
+--+ +-------- Medionidus
| |
+--+ +----- Potamilus_purpuratus
| |
+--+----- Lampsilis
|
| +-- Ellipsaria
+--+
+-- Villosa
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
136 |
6 |
3 |
44.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Pleurobema **
+--+
| +-- Elliptio **
+--+
+-- Eurynia **
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
3 |
43 |
10 |
2 |
42.0 |
+-- Obliquaria
+--+
| +-- Quadrulini
+--------------+
| | +-- Eurynia **
| | |
| +--+-- Pleurobema **
--+ |
| +-- Fusconaia **
|
| +----------------- Ptychobranchus
+--+
| +-------------- Truncilla
+--+
| +----------- Ortmanniana
+--+
| +-------- Lampsilis
+--+
| +----- Cambarunio
+--+
| +-- Ligumia
+--+
+-- Epioblasma
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
3 |
25 |
15 |
3 |
42.0 |
+-- Megalonaias
+-----------+
| +-- Quadrulina
|
--+ +----- Amblema
| +-----+
| | | +-- Plectomerus
| | +--+
+--+ +-- Lampsilini
|
| +-------- Elliptoideus **
+--+
| +----- Uniomerus
+--+
| +-- Eurynia **
+--+
+-- Pleurobema **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
9 |
0 |
3 |
122 |
6 |
3 |
48.0 |
+-- Amblemini
|
+-- Ortmanniana
|
+-- Lampsilis
|
+-- Cambarunio
|
--+-- Elliptio **
|
+-- Eurynia **
|
+-- Pleurobema **
|
+-- Quadrula_quadrula
|
+-- Quadrula_verrucosa
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
73 |
6 |
3 |
44.0 |
+-- Amblemini
+--------+
| +-- Lampsilini
--+
| +-------- Quadrulini
+--+
| +----- Pleurobema **
+--+
| +-- Eurynia **
+--+
+-- Elliptio **
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
28 |
8 |
3 |
41.0 |
+-------------- Amblemini
--+
| +----------- Lampsilini
+--+
| +-------- Quadrulini
+--+
| +----- Eurynia **
+--+
| +-- Fusconaia **
+--+
+-- Pleurobema **
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
90 |
15 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
72 |
13 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Pleurobema **
| +-----+
| | +-- Elliptio **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Smith et al., 2020 [Publ 4129], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
26 |
26 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Pleurobema **
| +-----+
| | +-- Elliptoideus **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
11 |
0 |
2 |
23 |
11 |
3 |
51.0 |
+----- Eurynia **
+-----------+
| | +-- Cambarunio
| +--+
| +-- Ptychobranchus
--+
| +-- Amblemini
| +-----------+
| | +-- Pleurobema **
+--+
| +----------- Truncilla
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Epioblasma
+--+
| +----- Ligumia
+--+
| +-- Obliquaria
+--+
+-- Ortmanniana
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
51 |
13 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Smith et al., 2020 [Publ 4129], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
26 |
26 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Pleurobema **
| +-----+
| | +-- Elliptoideus **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
24 |
20 |
3 |
-- |
+-------- Quadrulini
--+
| +----- Amblemini
+--+
| +-- Pleurobema **
+--+
+-- Lampsilini
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
26 |
22 |
3 |
-- |
+----------- Amblemini
--+
| +-------- Lampsilini
+--+
| +----- Quadrulini
+--+
| +-- Pleurobema **
+--+
+-- Elliptio **
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
18 |
7 |
3 |
-- |
+-------- Quadrulini
--+
| +----- Elliptio **
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
7 |
0 |
0 |
-- |
+-- Fusconaia **
--+
+-- Pleurobema **
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
19 |
18 |
4 |
-- |
+-------------- Quadrulini
|
--+ +-- Pleurobema **
| +--------+
| | +-- Fusconaia **
+--+
| +-------- Amblema
+--+
| +----- Plectomerus
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
72 |
13 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Pleurobema **
| +-----+
| | +-- Elliptio **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
94 |
15 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Wu et al., 2024 [Publ 4388], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
52 |
8 |
3 |
-- |
+-------- Quadrulini
|
--+ +-- Elliptio **
| +--+
| | +-- Pleurobema **
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
13 |
4 |
-- |
+-------------- Toxolasma
+--+
| | +----------- Popenaiadini
| +--+
| | +-------- Amblemini
| +--+
| | +----- Quadrulini
--+ +--+
| | +-- Pleurobema **
| +--+
| +-- Elliptio **
|
| +----------- Potamilus
| |
+-----+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
94 |
15 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
17 |
12 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
13 |
0 |
2 |
81 |
5 |
3 |
50.0 |
+-- Lamprotula
|
+-- Potomida
|
+-- Pronodularia
|
+-- Gonidea
|
+-- Leguminaia
|
+-- Microcondylaea
|
--+-- Amblemini
|
+-- Ortmanniana
|
+-- Lampsilis
|
+-- Cambarunio
|
+-- Elliptio **
|
+-- Pleurobema **
|
+-- Quadrulini
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
19 |
18 |
4 |
-- |
+-------------- Quadrulini
|
--+ +-- Pleurobema **
| +--------+
| | +-- Fusconaia **
+--+
| +-------- Amblema
+--+
| +----- Plectomerus
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
72 |
13 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Pleurobema **
| +-----+
| | +-- Elliptio **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
50 |
10 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
32 |
12 |
3 |
43.0 |
+----------------------------- Megalonaias
|
| +----- Elliptoideus **
--+ +--------------------+
| | | +-- Uniomerus
| | +--+
+--+ +-- Pleurobema **
|
| +----------------------- Amblema
| |
+--+ +-- Obliquaria
| +-----------------+
| | +-- Plectomerus
+--+
| +----------------- Medionidus
+--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis
+--+
| +-- Villosa
+--+
+-- Ellipsaria
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
90 |
15 |
4 |
-- |
+----------- Quadrulini
|
--+ +-- Elliptio **
| +-----+
| | +-- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
13 |
3 |
-- |
+-------- Quadrula_quadrula
|
+-------- Quadrula_verrucosa
--+
| +-- Eurynia **
| +--+
| | +-- Pleurobema **
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
17 |
0 |
2 |
20 |
9 |
3 |
49.0 |
+-- Gonideini
|
+-- Pyganodon_grandis
|
+-- Utterbackia
|
+-- Pyganodon_cataracta
|
+-- Lampsilis_straminea
|
+-- Lampsilis_teres
|
+-- Megalonaias
|
+-- Obliquaria
|
--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio **
|
+-- Fusconaia **
|
+-- Plectomerus
|
+-- Aculamprotula
|
+-- Unio
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
27 |
23 |
3 |
-- |
+----------- Megalonaias
|
--+ +-- Uniomerus
| +-----+
| | +-- Quadrulina
+--+
| +----- Pleurobema **
+--+
| +-- Lampsilini
+--+
+-- Amblemini
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
7 |
0 |
2 |
34 |
5 |
3 |
47.0 |
+-- Amblemini
|
+-- Ortmanniana
|
+-- Lampsilis
|
--+-- Cambarunio
|
+-- Elliptio **
|
+-- Pleurobema **
|
+-- Quadrulini
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
32 |
12 |
3 |
43.0 |
+----------- Megalonaias
|
| +----- Elliptoideus **
| +--+
--+ | | +-- Uniomerus
| | +--+
| | +-- Pleurobema **
| |
| | +----- Potamilus_fragilis
+--+ |
| +----- Medionidus
| |
| +----- Amblema
| |
| | +-- Obliquaria
+--+--+
| +-- Plectomerus
|
| +-- Villosa
| |
| +-- Ellipsaria
| |
+--+-- Lampsilis
|
+-- Potamilus_purpuratus
|
+-- Ptychobranchus
Burlakova et al., 2019 [Publ 3899], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
31 |
31 |
4 |
-- |
+-------------------------- Quadrulini
|
| +----------------------- Plectomerus
| |
--+ | +-------------------- Popenaiadini
| | |
| | | +-- Elliptio **
| | +-----------------+
+--+ | +-- Pleurobema **
| |
| | +-- Cyrtonaias
| | +--+
| | | +-- Glebula
| +--------------+
+--+ | +-- Reginaia
| +--+
| +-- Obliquaria
|
| +----------------- Toxolasma
| |
| | +-------------- Potamilus
| | |
| | | +-- Ellipsaria
| | +-----------+
+--+ | +-- Truncilla
| |
| | +----- Medionidus
| +--------+
| | | +-- Lemiox
| | +--+
+--+ +-- Dromus
|
| +----------- Cambarunio
| |
| +----------- Leaunio
| |
| +----------- Villosa
| |
| | +-- Sagittunio
| +--------+
+--+ +-- Lampsilis_ovata
|
| +----- Venustaconcha
+-----+
| | +-- Epioblasma
| +--+
| +-- Obovaria
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
Manendo et al., 2008 [Publ 2669], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
11 |
3 |
2 |
-- |
+-------- Lampsilini
--+
| +----- Amblemini
+--+
| +-- Pleuronaia **
+--+
+-- Fusconaia **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
17 |
3 |
-- |
+-------- Quadrulini
--+
| +----- Fusconaia **
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
3 |
-- |
+-------- Quadrula
|
--+-------- Pustulosa
|
| +----- Pleurobema **
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Johnson et al., 2018 [Publ 3844], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
15 |
2 |
-- |
+----- Amblemini
--+
| +-- Elliptio **
+--+
+-- Quadrulini
Soroka, 2008 [Publ 3695], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Fusconaia_flava
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
5 |
2 |
-- |
-- Fusconaia_flava
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
3 |
3 |
-- |
-- Eurynia_dilatata
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
5 |
1 |
-- |
+-- Fusconaia **
--+
+-- Lampsilini
Wu et al., 2021 [Publ 4366], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
90 |
5 |
2 |
-- |
+----- Lampsilini
--+
| +-- Quadrulini
+--+
+-- Elliptio **
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
5 |
1 |
-- |
-- Fusconaia_flava
Keogh & Simons, 2019 [Publ 3935], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
9 |
2 |
-- |
+----- Lampsilini
--+
| +-- Amblemini
+--+
+-- Fusconaia **
Metzger et al., 2018 [Publ 3952], fig. 1C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Elliptio **
--+
+-- Lampsilini
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
3 |
2 |
-- |
+----- Lampsilini
--+
| +-- Fusconaia **
+--+
+-- Quadrulini
Soroka, 2008 [Publ 3695], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Fusconaia_flava
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
3 |
2 |
-- |
+----- Lampsilini
--+
| +-- Eurynia **
+--+
+-- Amblemini
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
1 |
1 |
-- |
-- Elliptio_complanata
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
1 |
1 |
-- |
-- Elliptio_complanata
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
1 |
1 |
-- |
-- Elliptio
Wu et al., 2023 [Publ 4339], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
18 |
5 |
2 |
-- |
+----- Lampsilini
--+
| +-- Elliptio **
+--+
+-- Quadrulini
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
9 |
2 |
-- |
+----- Quadrulini
--+
| +-- Elliptio **
+--+
+-- Lampsilini
Krebs, 2004 [Publ 2692], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
2 |
2 |
-- |
+-- Lampsilini
--+
+-- Fusconaia **
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
4 |
2 |
-- |
+-- Quadrulini
|
--+-- Parvaspina **
|
+-- Lampsilini
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
37 |
16 |
3 |
-- |
+-- Fusconaia **
--+
+-- Quadrulini
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
14 |
3 |
-- |
+-- Eurynia **
+-----------+
| +-- Quadrulini
|
--+ +-- Paetulunio
| +--------+
| | +-- Amblemini
+--+
| +-------- Lampsilis
| |
+--+ +----- Leaunio
| |
+--+----- Obovaria
|
| +-- Villosa
+--+
+-- Cambarunio
Giribet & Distel, 2003 [Publ 3670], fig. 3f5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
1 |
1 |
-- |
-- Elliptio_complanata
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
1 |
1 |
-- |
-- Elliptio_complanata
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
90 |
5 |
2 |
-- |
+----- Quadrulini
--+
| +-- Elliptio **
+--+
+-- Lampsilini
Wu et al., 2021 [Publ 4366], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
11 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Popenaiadini
+--+
+-- Lampsilini
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
4 |
3 |
-- |
+-------- Pleurobema **
--+
| +----- Quadrulini
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
5 |
1 |
-- |
+-- Fusconaia **
--+
+-- Lampsilini
Campbell, 2000 [Publ 2724], fig. 1-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Elliptio_complanata
Krebs, 2004 [Publ 2692], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
1 |
1 |
-- |
+-- Lampsilini
--+
+-- Fusconaia **
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
5 |
1 |
-- |
+-- Fusconaia **
--+
+-- Lampsilini
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
26 |
25 |
4 |
-- |
+----------- Quadrulini
--+
| +-------- Pleurobema **
+--+
| +----- Amblemini
+--+
| +-- Lampsilini
+--+
+-- Popenaiadini
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
5 |
1 |
-- |
+-- Fusconaia **
--+
+-- Lampsilini
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
7 |
1 |
-- |
+-- Fusconaia **
--+
+-- Lampsilini
Hoeh et al., 1996 [Publ 2694], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Fusconaia_flava
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
5 |
1 |
-- |
+----------- Cyrtonaias
|
--+----------- Fusconaia **
|
| +-------- Potamilus
+--+
| +----- Lampsilis
+--+
| +-- Ligumia
+--+
+-- Ortmanniana
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
11 |
3 |
-- |
+----- Lampsilini
--+
| +-- Amblemini
+--+
+-- Eurynia **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
119 |
3 |
3 |
-- |
+-- Amblemini
|
+-- Elliptio **
--+
+-- Lampsilini
|
+-- Quadrulini
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
9 |
1 |
-- |
+-------------- Fusconaia **
|
+-------------- Obliquaria
--+
| +----------- Potamilus_capax
| |
| +----------- Lampsilis
+--+
| +-------- Potamilus_fragilis
| |
| | +----- Potamilus_inflatus
+--+ |
| | +-- Potamilus_ohiensis
| +--+
+--+ +-- Potamilus_amphichaenus
|
| +-- Potamilus_purpuratus__1
| |
+--+-- Potamilus_purpuratus__2
|
+-- Potamilus_alatus
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
35 |
4 |
3 |
-- |
+----- Quadrulini
|
--+----- Pleurobema **
|
| +-- Amblemini
+--+
+-- Lampsilini
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
4 |
3 |
-- |
+-------- Quadrulini
--+
| +----- Eurynia **
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
53 |
10 |
2 |
-- |
+----- Quadrulini
--+
| +-- Elliptio **
+--+
+-- Lampsilini
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
5 |
1 |
-- |
+-- Fusconaia **
--+
+-- Lampsilini
| 
