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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Unio

species Unio nanus Lamarck, 1819: nominal species
     47 specimens available.

Unio nana Lamarck, 1819
Lamarck, 1819, Hist. Natur. Anim. sans Vert. 6: 76, sp. 27.

Unio nanaLamarck (1819), Westerlund (1890), Kobelt & Haas (1910)
Unio littoralisLea (1834), Hanley (1843)
Margarita (Unio) litoralisLea (1836), Lea (1838)
Margaron (Unio) litoralisLea (1852), Lea (1870)
Unio (Batavi) nanusDupuy (1852)
Unio (Lymnium) batavus var. L nanusMoquin-Tandon (1855)
Unio batavus var. nanusde Joannis (1859)
Unio litoralisReeve (1865)
Unio nanusKobelt (1879), Lopes-Lima et al. (2024)
Unio (Groupe de l’U. mancus) nanusLocard (1882)
Unio (Nanusiana) nanusLocard (1889)
Unio (sec. Hyridella) batavus vr. nanusPaetel (1890)
Unio (Groupe de l’U. nanus) nanusLocard (1893)
Unio (groupe de l’U. nanus) nanusDumas (1895)
Unio (sec. Lymnium) littoralisSimpson (1900), Simpson (1914)
Unio (Rhomboïdiana) nanaGermain (1904)
Unio (Groupe des Ninusiana) nanusCaziot (1907)
Unio mancus var. nanusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Batavusiana (s.s.) nanaZatravkin (1983), Zatravkin (1987)
Unio (Crassiana) crassus nanusFalkner et al. (2001), Falkner et al. (2002)
Crassiana (Bataviana) nanaStarobogatov et al. (2004)
Crassiana nana nanaKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassusGraf (2007), Anisimova et al. (2018), Bogatov et al. (2018)
Unio crassus nanusPrié & Puillandre (2014)
Unio (Crassiana) nanaVinarski & Kantor (2016)
Unio crassus crassusKlishko et al. (2017)
Unio crassus nanaBogatov (2022)

Unio cytherea Küster, 1833
Küster, 1833, Icones Moll. Test. Abthl. 2: pl. 5, fig. 3.

Unio cythereaKüster (1833)
Unio (Mysca) conusMörch (1863)
Unio cytherea cythereaHaas (1913)
Unio crassus cythereaHaas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus cythereaFalkner et al. (2001), Falkner et al. (2002)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio crassusAraujo et al. (2018)
Unio nanusLopes-Lima et al. (2024)

Unio fusculus Rossmässler, 1836
‘Z.’ Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 21.
‘Ziegler’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 30, pl. 15, fig. 211.

Unio fusculusRossmässler (1836), Zelebor (1851)
Unio batavusRossmässler (1835), Westerlund (1890)
Margarita (Unio) batavaLea (1836), Lea (1838)
Mysca batava var. fusculaSchlüter (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Lymnium) batavus var. E ripariusMoquin-Tandon (1855)
Unio batavus var. crassusClessin (1877)
Unio (Groupe de l’U. fusculus) fusculusLocard (1882), Locard (1893)
Unio (Fusculusiana) fusculusLocard (1889), Germain (1904)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus var. piscinalisGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Batavusiana (s.s.) fusculaZatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) fuscula fusculaStadnichenko (1984)
Crassiana (Bataviana) fusculaStarobogatov et al. (2004)
Crassiana fusculaKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassusGraf (2007)
Unio (Crassiana) fusculusVinarski & Kantor (2016)
Unio crassus crassusKlishko et al. (2017)
Unio crassus musivusBogatov (2022)
Unio nanusLopes-Lima et al. (2024)

Unio riparia Pfeiffer, 1821
Bruguière, 1797, Encycl. Méth. 2: pl. 249, fig. 4a-b [as Unio sp.].
Pfeiffer, 1821, Natur. Deutsch. Land Sussw. Moll. 1: 118, pl. 5, fig. 13.
Scholtz, 1843, Land- und Wassermoll. Schlesiens: 129. [nomen nudum]

Unio ripariaPfeiffer (1821), Krynicki (1837)
Unio ripariusMenke (1828), Menke (1830), Servain (1885), Westerlund (1890)
Unio batavusRossmässler (1835)
Margarita (Unio) batavaLea (1836), Lea (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Lymnium) batavus var. E ripariusMoquin-Tandon (1855)
Unio (Groupe de l’U. fusculus) ripariusLocard (1882)
Unio (Amniciana) ripariusServain (1888)
Unio (Amnicusiana) ripariusLocard (1889)
Unio (sec. Hyridella) batavus vr. ripariusPaetel (1890)
Unio (Groupe de l’U. amnicus) ripariusLocard (1893)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Socardiana) ripariusGermain (1904)
Unio mancus var. amnicus form ripariusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio (Crassiana) crassus ripariusFalkner et al. (2001), Falkner et al. (2002)
Unio crassus crassusLopes-Lima et al. (2017)
Unio nanusLopes-Lima et al. (2024)

Unio decurvatus Rossmässler, 1835
Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 22, pl. 9, fig. 131.
Rossmässler, 1837, Icon. Land- und Süssw.-Moll. 1 (5-6): 21, pl. 24, fig. 339.

Unio decurvatusRossmässler (1835), Rossmässler (1837), J.F. Schmidt (1840), C.B. Adams (1847), Küster (1854), Sowerby (1867), Westerlund (1890)
Margarita (Unio) batavaLea (1836), Lea (1838)
Mysca decurvataSchlüter (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Sandriana) decurvatusLocard (1889)
Unio (s.s.) decurvatusPaetel (1890)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus decurvatusFalkner et al. (2001)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio batavus var. albensis Hazay, 1885
Hazay, 1885, Jb. Dtsch. Malak. Ges. 12: 43.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. albensisHazay (1885), Westerlund (1890)
Unio albensisKobelt (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassus albensisFalkner et al. (2001)
Unio crassus batavusLopes-Lima et al. (2017)
Unio nanusLopes-Lima et al. (2024)

Unio batavus var. ondovensis Hazay, 1885
Hazay, 1885, Jb. Dtsch. Malak. Ges. 12: 43.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. ondovensisHazay (1885), Westerlund (1890)
Unio ondovensisKobelt (1890)
Unio crassus cythereaHaas (1940), Haas (1969), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus ondovensisFalkner et al. (2001)
Unio nanusLopes-Lima et al. (2024)

Mya ovata Coxe, 1789
Studer in Coxe, 1789, unknown publication: 392. [nomen nudum]

Mya ovataCoxe (1789)
Unio nanusLopes-Lima et al. (2024)

Mya pictorum Coxe, 1789
Coxe, 1789, unknown publication: 392. [nomen nudum]
(misidentified reference to Mya pictorum Linnaeus, 1758)

Mya pictorumCoxe (1789)
Unio nanusLopes-Lima et al. (2024)

Unio dilatatus Studer, 1820
Bruguière, 1797, Encycl. Méth. 2: pl. 249, fig. 4a-b [as Unio sp.].
Studer, 1820, System. Verzeich. bekannt geword. Schweizer-Conchylien: 30.

Unio dilatatusStuder (1820)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Lymnium) batavusMoquin-Tandon (1855)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio inflatus Studer, 1820
Draparnaud, 1806, Hist. Moll. France: pl. 11, figs. 2. [in part]
Studer, 1820, System. Verzeich. bekannt geword. Schweizer-Conchylien: 30.
(misidentified reference to Unio pictorum Draparnaud, 1806)

Unio inflatusStuder (1820)
Unio limosusCharpentier (1837)
Margaron (Unio) pictorumLea (1870)
Unio pictorumHe & Zhuang (2013)
Unio (s.s.) pictorumVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio ovatus Studer, 1820
Schröter, 1779, Gesch. Flußconch.: pl. 3, figs. 2.
Studer, 1820, System. Verzeich. bekannt geword. Schweizer-Conchylien: 30.
(homonym of Unio ovatus Say, 1817)

Unio ovatusStuder (1820), Servain (1885), Westerlund (1890)
Unio batavus var. ovatusCharpentier (1837)
Margaron (Unio) tumidusLea (1852), Lea (1870)
Unio (Lymnium) batavus var. C ovatusMoquin-Tandon (1855)
Unio (Lymnium) batavusDrouët (1857)
Unio (Fusculusiana) ovatusLocard (1889)
Unio (Groupe de l’U. fusculus) ovatusLocard (1893)
Unio consentaneus ovatusKobelt & Haas (1910)
Unio crassus var. piscinalisGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio planus Studer, 1820
Studer, 1820, System. Verzeich. bekannt geword. Schweizer-Conchylien: 30.

Unio planusStuder (1820)
Margarita (Unio) batavaLea (1836), Lea (1838)
Unio limosusCharpentier (1837)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio rostratus Studer, 1820
Studer, 1820, System. Verzeich. bekannt geword. Schweizer-Conchylien: 30.
(homonym of Unio rostrata Lamarck, 1819)

Unio rostratusStuder (1820)
Margarita (Unio) ovalisLea (1836)
Unio tumidusCharpentier (1837), Reeve (1865)
Margaron (Unio) tumidusLea (1852), Lea (1870)
Unio (Lymnium) tumidusMoquin-Tandon (1855)
Unio nanusLopes-Lima et al. (2024)

Unio sinuatus Studer, 1820
Schröter, 1779, Gesch. Flußconch.: 182, pl. 2, fig. 2 [Mya testa crassa]. [non-binomial]
Studer, 1820, System. Verzeich. bekannt geword. Schweizer-Conchylien: 30.
(homonym of Unio sinuata Lamarck, 1819)

Unio sinuatusStuder (1820)
Margarita (Unio) batavaLea (1836), Lea (1838)
Unio batavus var. sinuatusCharpentier (1837)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Lymnium) batavus var. M sinuatusMoquin-Tandon (1855)
Margaron (Margaritana) margaritiferaLea (1870)
Unio (Batavusiana) financeiLocard (1889)
Unio squamosusDrouët (1889)
Unio batavusGermain (1931)
Unio crassus cythereaHaas (1940)
Unio nanusLopes-Lima et al. (2024)

Unio castaneus Fitzinger, 1833
Fitzinger, 1833, Syst, Verz. Österr. Weichth.: 119.
(homonym of Unio castaneus Lea, 1831)

Unio castaneusFitzinger (1833)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio dubius Fitzinger, 1833
Fitzinger, 1833, Syst, Verz. Österr. Weichth.: 119. [nomen nudum]

Unio dubiusFitzinger (1833), Zelebor (1851)
Unio (Lymnium) aterDrouët (1857)
Margaron (Unio) pictorumLea (1870)
Unio (sec. Lymnium) pictorumSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio nigricans Fitzinger, 1833
Fitzinger, 1833, Syst, Verz. Österr. Weichth.: 119. [nomen nudum]

Unio nigricansFitzinger (1833), Zelebor (1851)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio sinuolata Küster, 1833
Küster, 1833, Icones Moll. Test. Abthl. 2: pl. 5, fig. 5.

Unio sinuolataKüster (1833)
Unio cytherea cythereaHaas (1913)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio labacensis Rossmässler, 1835
‘Ziegler’ Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 21.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio labacensisRossmässler (1835), J.F. Schmidt (1840)
Unio batavusRossmässler (1835), Küster (1855), Westerlund (1890)
Margarita (Unio) batavaLea (1836), Lea (1838)
Mysca batava var. labacensisSchlüter (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio batavus var. crassusClessin (1877)
Unio amnicusDrouët (1889)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio vitreus Rossmässler, 1835
‘Stenz’ Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 21.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio vitreusRossmässler (1835)
Margaron (Unio) batavusLea (1870)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio amnicus Rossmässler, 1836
Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 20.
‘Ziegl.’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 31, pl. 15, fig. 212.

Unio amnicusRossmässler (1836), J.F. Schmidt (1840), C.B. Adams (1847), Küster (1854), Servain (1885), Drouët (1889), Westerlund (1890)
Unio batavusRossmässler (1835)
Margarita (Unio) batavaLea (1836), Lea (1838)
Mysca amnicaSchlüter (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Batavi) nanusDupuy (1852)
Unio (Lymnium) batavus var. L nanusMoquin-Tandon (1855)
Unio batavus var. crassusClessin (1877)
Unio (Groupe de l’U. fusculus) amnicusLocard (1882)
Unio (Amniciana) amnicusServain (1888)
Unio (Amnicusiana) amnicusLocard (1889)
Unio (sec. Hyridella) batavus vr. amnicusPaetel (1890)
Unio (Groupe de l’U. amnicus) amnicusLocard (1893)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (Groupe des Amnicusiana) amnicusCaziot (1907)
Unio consentaneus minorKobelt & Haas (1910)
Unio mancus var. amnicusGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio atrovirens Rossmässler, 1836
‘Schmidt’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 28, pl. 15, figs. 206-207.

Unio atrovirensRossmässler (1836), J.F. Schmidt (1840)
Mysca decurvata subvar. atrovirensSchlüter (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio decurvatusKüster (1854)
Unio batavus var. aterClessin (1877)
Unio (s.s.) decurvatus vr. atrovirensPaetel (1890)
Unio decurvatus var. atrovirensWesterlund (1890)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio carinthiacus Rossmässler, 1836
‘Z.’ Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 21.
‘Ziegl.’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 30, pl. 15, fig. 209.
Type(s): PARATYPE BMNH_1965216, Carinthia.

Unio carinthiacusRossmässler (1836), Reeve (1865)
Unio batavusRossmässler (1835), Küster (1855), Westerlund (1890), Germain (1931)
Margarita (Unio) batavaLea (1836), Lea (1838)
Mysca batava var. carynthiacaSchlüter (1838)
Unio carynthiacusJ.F. Schmidt (1840), C.B. Adams (1847), Schröder (1885)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio batavus var. crassusClessin (1877)
Unio (Bataviana) carynthiacusServain (1888)
Unio (Batavusiana) carynthiacusLocard (1889), Germain (1904)
Unio (Groupe de l’U. batavus) carinthiacusLocard (1893)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus Rossmässler, 1836
‘Ziegl.’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 29, pl. 15, fig. 208.
‘Ziegl.’ Rossmässler, 1838, Icon. Land- und Süssw.-Moll. 2 (1-2): 42, pl. 40, fig. 544.
Rossmässler, 1842, Icon. Land- und Süssw.-Moll. 2 (5): 14, pl. 55, fig. 742.

Unio consentaneusRossmässler (1836), Rossmässler (1838), J.F. Schmidt (1840), Rossmässler (1842), Zelebor (1851), Kobelt (1910), Germain (1931)
Margarita (Unio) batavaLea (1836), Lea (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Potamida) crassusMörch (1853)
Unio aterKüster (1854), Westerlund (1890)
Unio (Lymnium) aterMoquin-Tandon (1855), Drouët (1857)
Unio batavus var. aterClessin (1877)
Unio ater vr. consentaneusPaetel (1890)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio consentaneus var.Kobelt & Haas (1910)
Unio crassus consentaneusOrtmann (1912)
Unio cytherea cythereaHaas (1913)
Unio (s.s.) crassus consentancusZhadin (1938)
Unio crassus cythereaHaas (1940), Haas (1969)
Batavusiana (s.s.) nanaZatravkin (1987)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio piscinalis Rossmässler, 1836
‘Z.’ Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 21.
‘Ziegl.’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 30, pl. 15, fig. 210.

Unio piscinalisRossmässler (1836)
Unio batavusRossmässler (1835)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio batavus var. crassusClessin (1877)
Unio (Groupe de l’U. fusculus) piscinalisLocard (1882), Locard (1893)
Unio (Fusculusiana) piscinalisLocard (1889)
Unio batavus var. piscinalisWesterlund (1890)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus var. piscinalisGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio reniformis Rossmässler, 1836
‘Schmidt’ Rossmässler, 1836, Icon. Land- und Süssw.-Moll. 1 (3): 31, pl. 15, fig. 213.

Unio reniformisRossmässler (1836), J.F. Schmidt (1840), Küster (1854), Sowerby (1868), Westerlund (1890)
Margarita (Unio) batavaLea (1836), Lea (1838)
Mysca reniformisSchlüter (1838)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Potamida) reniformisMörch (1853)
Unio (Lymnium) batavus var. N reniformisMoquin-Tandon (1855)
Unio batavus var. aterClessin (1877)
Unio (Groupe de l’U. fusculus) reniformisLocard (1882), Locard (1893)
Unio (Fusculusiana) reniformisLocard (1889)
Unio ater vr. reniformisPaetel (1890)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus var. piscinalisGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio retusa Held, 1836
Held, 1836, Isis (Oken) 1836: 281.
(homonym of Unio retusa Lamarck, 1819)

Unio retusaHeld (1836)
Margaron (Unio) pictorumLea (1852), Lea (1870)
Unio pictorumDrouët (1889)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus var. elongatus Held, 1837
Held, 1837, Isis (Oken) 1837 (4): 306.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio elongata Lamarck, 1819)

Unio batavus var. elongatusHeld (1837)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus var. gibbosus Held, 1837
Held, 1837, Isis (Oken) 1837 (4): 306.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio gibbosus Barnes, 1823)

Unio batavus var. gibbosusHeld (1837)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus var. squamosus Charpentier, 1837
Charpentier, 1837, Cat. Moll. Suisse: 25, pl. 2, fig. 22.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. squamosusCharpentier (1837)
Unio (Lymnium) batavus var. H squamosusMoquin-Tandon (1855)
Unio squamosusDrouët (1881), Drouët (1889), Westerlund (1890)
Unio (sec. Hyridella) batavus vr. squamosusPaetel (1890)
Unio consentaneus squamosusKobelt (1910), Kobelt & Haas (1910)
Unio consentaneusGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus legitimus Rossmässler, 1838
Rossmässler, 1838, Icon. Land- und Süssw.-Moll. 2 (1-2): 25, pl. 35, fig. 491a. [infra-subsp.]
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus legitimusRossmässler (1838)
Unio consentaneus form legitimusRossmässler (1838)
Unio crassus cythereaHaas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus minor Rossmässler, 1838
Rossmässler, 1838, Icon. Land- und Süssw.-Moll. 2 (1-2): 25, pl. 35, fig. 491c. [infra-subsp.]
(homonym of Unio margaritifer var. minor Rossmässler, 1835; intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus minorRossmässler (1838), Kobelt & Haas (1910)
Unio consentaneus form minorRossmässler (1838)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus pulcher Rossmässler, 1838
Rossmässler, 1838, Icon. Land- und Süssw.-Moll. 2 (1-2): 25, pl. 35, fig. 491b. [infra-subsp.]
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836; homonym of Unio pulcher Lea, 1838)

Unio consentaneus pulcherRossmässler (1838)
Unio consentaneus form pulcherRossmässler (1838)
Unio nanusLopes-Lima et al. (2024)

Unio pruinosus J.F. Schmidt, 1840
J.F. Schmidt, 1840, Bull. Soc. imp. Nat. Moscou 13: 445, pl. 9, fig. 3.

Unio pruinosusJ.F. Schmidt (1840), Drouët (1881), Drouët (1889), Westerlund (1890)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio (Groupe de l’U. fusculus) fusculusLocard (1882)
Unio (Fusculusiana) pruinosusLocard (1889)
Unio (Groupe de l’U. fusculus) pruinosusLocard (1893)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus var. piscinalisGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio heldii Küster, 1854
Küster, 1854, Conchyl. Cab. 9(2): 111, pl. 30, fig. 5-7.

Unio heldiiKüster (1854), Drouët (1881), Drouët (1881), Westerlund (1890), Kobelt & Haas (1910)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio nessorhynchus Küster, 1854
Küster, 1854, Conchyl. Cab. 9(2): 116, pl. 31, fig. 6.

Unio nessorhynchusKüster (1854)
Margaron (Unio) batavusLea (1852), Lea (1870)
Unio aterKüster (1854)
Unio ater var. nessorhynchusWesterlund (1890)
Unio cytherea nessorhynchusHaas (1913)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio bosnensis Möllendorff, 1874
Möllendorff, 1874, Beitr. Fauna Bosniens: 62, text figs. 21-22.
(renamed Unio bosnicus Locard, 1889)

Unio bosnensisMöllendorff (1874), Kobelt (1879)
Unio (Fusculusiana) bosnicusLocard (1889)
Unio (sec. Hyridella) bosnensisPaetel (1890)
Unio bosnicusWesterlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio phaseolus Kobelt, 1879
‘Held’ Kobelt, 1879, Icon. Land- und Süssw.-Moll. 6: 44, pl. 163, fig. 1649.
(homonym of Unio phaseolus Hildreth, 1828)

Unio phaseolusKobelt (1879)
Unio heldiiKüster (1854), Drouët (1881), Westerlund (1890), Kobelt & Haas (1910)
Unio batavus var. crassusClessin (1877)
Unio (sec. Hyridella) batavus vr. phaseolusPaetel (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio croaticus Drouët, 1881
Drouët, 1881, J. de Conch. 29: 245.

Unio croaticusDrouët (1881), Kobelt (1886), Westerlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio neocomensis Drouët, 1881
Drouët, 1881, J. de Conch. 29: 247.

Unio neocomensisDrouët (1881), Westerlund (1890)
Unio neocomiensisKobelt (1886)
Unio (Sandriana) necomensisLocard (1889)
Unio (Groupe de l’U. macrorhynchus) necomensisLocard (1893)
Unio mancus var. macrorhynchusGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio danemorae Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 284, 354 [as ‘danemoræ’].
‘Mörch’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 147 [41] [as ‘danemoræ’].
‘Mörch’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 177 [as ‘danemoræ’].
Type(s): IDIOTYPE MNHN_22186, lac d’Ondres (Basses-Pyrénées).

Unio danemoraeLocard (1882)
Unio (Groupe de l’U. ater) danemoræLocard (1882), Locard (1893)
Unio (Ateriana) danemoræLocard (1889)
Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio dubisianus Locard, 1882
‘Coutagne’ Locard, 1882, Prodr. Malac. Franç. 1: 291 [as ‘dubisanus’], 360.
‘Coutagne’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 129 [23].
‘Coutagne’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 158.
(correct name for Unio dubisanus Locard, 1882)
Type(s): SYNTYPE MNHN_22198, La Clange (Jura).

Unio dubisianusLocard (1882), Drouët (1889), Westerlund (1890)
Unio (Groupe de l’U. mancus) dubisanusLocard (1882)
Unio (Mancusiana) dubisianusLocard (1889)
Unio (Groupe de l’U. mancus) dubisianusLocard (1893)
Unio (Groupe des Mancusiana) dubisianusCaziot (1907)
Unio mancusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus mancusPrie (2012)
Unio nanusLopes-Lima et al. (2024)

Unio dubisiopsis Locard, 1882
Locard, 1882, Prodr. Malac. Franç. 1: 291 [as ‘dubisopsis’], 360.
Locard, 1889, Contr. Faune Malacol. Franc. 13: 128 [22] [as ‘dubisianopsis’].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 155 [as ‘dubisianopsis’].
(correct name for Unio dubisianopsis Locard, 1889)

Unio dubisiopsisLocard (1882)
Unio (Groupe de l’U. mancus) dubisiopsisLocard (1882)
Unio (Melasiana) dubisianopsisLocard (1889)
Unio dubisianus var. dubisianopsisWesterlund (1890)
Unio (Groupe de l’U. melas) dubisianopsisLocard (1893)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio macrorhynchus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 292, 361.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 132 [26].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 158, fig. 172.
Type(s): LECTOTYPE MNHN_22326, lac du Bourget.

Unio macrorhynchusLocard (1882), Westerlund (1890)
Unio (Groupe de l’U. sandrii) macrorhynchusLocard (1882)
Unio (Sandriana) macrorhynchusLocard (1889)
Unio (Groupe de l’U. macrorhynchus) macrorhynchusLocard (1893)
Unio mancus var. macrorhynchusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio melas Locard, 1882
Coutagne in Locard, 1882, Prodr. Malac. Franç. 1: 285, 355.
‘Coutagne’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 128 [22].
‘Coutagne’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 155, fig. 169.
Type(s): SYNTYPE MNHN_22363, Saulcy-sur-Meurthe (Vosges) [written on shell].

Unio melasLocard (1882)
Unio (Groupe de l’U. crassus) melasLocard (1882)
Unio (Melasiana) melasLocard (1889), Germain (1904)
Unio (Groupe de l’U. melas) melasLocard (1893)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio potamius Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 289, 359.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 140 [34].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 169.
Type(s): IDIOTYPE MNHN_22470, La Brizotte (Côte-d’Or).

Unio potamiusLocard (1882), Bourguignat (1883)
Unio (Groupe de l’U. Batavus) potamiusLocard (1882)
Unio (Ellipsopsisiana) potamiusLocard (1889)
Unio batavus var. potamiusWesterlund (1890)
Unio (Groupe de l’U. batavus) potamiusLocard (1893)
Unio (Batavusiana) potamiusGermain (1904)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio savensis Drouët, 1882
Drouët, 1882, Unionidae de Serbie: 15.

Unio savensisDrouët (1882), Kobelt (1884), Westerlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio socardianus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 286, 355.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 134 [28].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 161.

Unio socardianusLocard (1882)
Unio (Crassiana) socardianusServain (1882)
Unio (Groupe de l’U. crassus) socardianusLocard (1882)
Unio (Socardiana) socardianusLocard (1889)
Unio (Groupe de l’U. feliciani) socardianusLocard (1893)
Unio crassusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio striatulus Drouët, 1882
Drouët, 1882, Unionidae de Serbie: 19.
(homonym of Unio striatulus Lea, 1857; renamed Unio striolatus Drouët, 1884)

Unio striatulusDrouët (1882), Kobelt (1886)
Unio striolatusDrouët (1889), Westerlund (1890)
Unio crassus cythereaHaas (1940)
Unio nanusLopes-Lima et al. (2024)

Unio truncatulus Drouët, 1882
Drouët, 1882, Unionidae de Serbie: 12.
(homonym of Unio rhomboideus var. truncatulus de Joannis, 1859)

Unio truncatulusDrouët (1882), Kobelt (1886), Westerlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio dokici Drouët, 1884
Drouët, 1884, Suppl. Unionidae de Serbie: 5, pl. 1, fig. 1.

Unio dokiciDrouët (1884), Kobelt (1886), Westerlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio rivalis Drouët, 1884
Drouët, 1884, Suppl. Unionidae de Serbie: 13.

Unio rivalisDrouët (1884), Kobelt (1886), Westerlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio serbicus Drouët, 1884
Drouët, 1884, Suppl. Unionidae de Serbie: 9, pl. 1, fig. 1.

Unio serbicusDrouët (1884), Kobelt (1886), Westerlund (1890)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio striolatus Drouët, 1884
Drouët, 1884, Suppl. Unionidae de Serbie: 9.
(new name for Unio striatulus Drouët, 1882)

Unio striolatusDrouët (1884), Drouët (1889), Westerlund (1890)
Unio crassus cythereaHaas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio duregicus Servain, 1885
Servain, 1885, Bull. Soc. Malac. France 2: 331.

Unio duregicusServain (1885), Westerlund (1890)
Unio (Fusculusiana) duregicusLocard (1889)
Unio consentaneus ostiorumKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio ostiorum Servain, 1885
‘Brot, mss.’ Servain, 1885, Bull. Soc. Malac. France 2: 327.

Unio ostiorumServain (1885)
Unio (Sandriana) necomensisLocard (1889)
Unio neocomensisWesterlund (1890)
Unio consentaneus ostiorumKobelt & Haas (1910)
Unio mancus var. macrorhynchusGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio sandriopsis Servain, 1885
‘Bourguignat, mss.’ Servain, 1885, Bull. Soc. Malac. France 2: 325.

Unio sandriopsisServain (1885), Westerlund (1890)
Unio (Sandriana) sandriopsisLocard (1889)
Unio (Groupe de l’U. macrorhynchus) sandriopsisLocard (1893)
Unio mancus var. macrorhynchusGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio subrobustus Servain, 1885
‘Bourguignat, mss.’ Servain, 1885, Bull. Soc. Malac. France 2: 332.

Unio subrobustusServain (1885), Westerlund (1890)
Unio (Socardiana) subrobustusLocard (1889)
Unio (Groupe de l’U. feliciani) subrobustusLocard (1893)
Unio crassusGermain (1931)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio tiguricus Servain, 1885
Servain, 1885, Bull. Soc. Malac. France 2: 329.

Unio tiguricusServain (1885), Westerlund (1890)
Unio (Sandriana) tiguricusLocard (1889)
Unio consentaneus ostiorumKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio turicus Servain, 1885
Servain, 1885, Bull. Soc. Malac. France 2: 327.

Unio turicusServain (1885), Westerlund (1890)
Unio (Sandriana) turicusLocard (1889)
Unio consentaneus ostiorumKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio kochii Kobelt, 1886
Sandberger & Koch, 1851, Jb. Nassau. Ver. Naturk. 7: 245 [as ‘U. moquinianus’].
Kobelt, 1886, Jb. Nassau. Ver. Naturk. 39: 97, pl. 6, fig. 1.
Kobelt, 1886, Icon. Land- und Süssw.-Moll. 2: 46, pl. 56, figs. 295-296.

Unio kochiiKobelt (1886)
Unio kochiKobelt (1886), Westerlund (1890)
Unio crassus crassusHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio batavus var. taunica Kobelt, 1886
Kobelt, 1886, Jb. Nassau. Ver. Naturk. 39: 96, pl. 7, fig. 5.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. taunicaKobelt (1886)
Unio batavus var. aterClessin (1877)
Unio (sec. Hyridella) batavus vr. taunicusPaetel (1890)
Unio batavus var. taunicusWesterlund (1890)
Unio batavus taunicusHaas (1910)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio badiellus Drouët, 1888
Drouët, 1888, J. de Conch. 36: 107.
Drouët, 1889, Mém. Acad. Sci. Dijon 1: 76, pl. 1, fig. 4.

Unio badiellusDrouët (1888), Drouët (1889), Westerlund (1890)
Unio (Locardiana) badiellusLocard (1889)
Unio (Groupe de l’U. batavus) badiellusLocard (1893)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio bosnicus Locard, 1889
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 132 [26].
(new name for Unio bosnensis Möllendorff, 1874)

Unio bosnicusLocard (1889), Westerlund (1890)
Unio (Fusculusiana) bosnicusLocard (1889)
Unio crassus cythereaHaas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio euthymeanus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 166 [60], 245 [139].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 203.
Type(s): LECTOTYPE MNHN_22202, La Saône à Collonges (Rhône).

Unio euthymeanusLocard (1889), Westerlund (1890)
Unio (Vinceleusiana) euthymeanusLocard (1889)
Unio (Groupe de l’U. tumens) euthymeanusLocard (1893)
Unio batavusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio ignari Locard, 1889
‘Nilss.’ Drouët, 1857, Étud. Naïades. France, Unio: 72, pl. 4, fig. 1 [as ‘U. ater’].
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 146 [40], 213 [107].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 176.
Type(s): SYNTYPES MNHN_22270, Le Mandrezey, à Saulcy-sur-Meurthe (Vosges).

Unio ignariLocard (1889), Westerlund (1890)
Unio (Ateriana) ignariLocard (1889)
Unio (Groupe de l’U. ater) ignariLocard (1893)
Unio consentaneusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio jurianus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 129 [23], 187 [81].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 157, fig. 171 [as ‘juranus’].
Type(s): LECTOTYPE MNHN_22292, La Valouze (Jura).

Unio jurianusLocard (1889), Westerlund (1890)
Unio (Mancusiana) jurianusLocard (1889)
Unio (Groupe de l’U. mancus) juranusLocard (1893)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio manculus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 130 [24], 190 [84].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 156.
Type(s): LECTOTYPE MNHN_22329, Gigny (Jura).

Unio manculusLocard (1889), Westerlund (1890)
Unio (Mancusiana) manculusLocard (1889)
Unio (Groupe de l’U. mancus) manculusLocard (1893)
Unio mancusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus mancusPrie (2012)
Unio nanusLopes-Lima et al. (2024)

Unio nubilus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 138 [32], 198 [92].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 165, fig. 179.
(homonym of Unio nubilus Lea, 1868)
Type(s): LECTOTYPE MNHN_22414, Verpillière (Isère).

Unio nubilusLocard (1889), Westerlund (1890), Rochebrune (1903)
Unio (Nubilusiana) nubilusLocard (1889)
Unio (Groupe de l’U. locardianus) nubilusLocard (1893)
Unio mancus var. elongatulusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio orbus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 138 [32], 195 [89].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 165.
Type(s): LECTOTYPE MNHN_22427, Pentoise (Jura).

Unio orbusLocard (1889), Westerlund (1890)
Unio (Locardiana) orbusLocard (1889)
Unio (Groupe de l’U. locardianus) orbusLocard (1893)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio passavanti Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 159 [53], 238 [132].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 178.
Type(s): POSSIBLE SYNTYPE MNHN_22442, Passavant (Haute-Seine).

Unio passavantiLocard (1889), Westerlund (1890)
Unio (Mariana) passavantiLocard (1889)
Unio (Groupe de l’U. ater) passavantiLocard (1893)
Unio consentaneus var. moquiniGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus moquinianusPrie (2012)
Unio nanusLopes-Lima et al. (2024)

Unio redactus Drouët, 1889
Drouët, 1889, Mém. Acad. Sci. Dijon 1: 79, pl. 1, fig. 2.

Unio redactusDrouët (1889), Kobelt (1893)
Unio (Amnicusiana) amnicusLocard (1889)
Unio amnicusWesterlund (1890)
Unio nanaKobelt & Haas (1910)
Unio mancus var. amnicusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio scotinus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 148 [42], 218 [112].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 178, fig. 192.
Type(s): LECTOTYPE MNHN_22518, Selles (Haute-Saône).

Unio scotinusLocard (1889), Westerlund (1890)
Unio (Ateriana) scotinusLocard (1889)
Unio (Groupe de l’U. ater) scotinusLocard (1893)
Unio consentaneus var. moquiniGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus moquinianusPrie (2012)
Unio nanusLopes-Lima et al. (2024)

Unio stygnus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 148 [42], 219 [113].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 179.
Type(s): LECTOTYPE MNHN_22556, Passavant (Haute-Saône).

Unio stygnusLocard (1889), Westerlund (1890)
Unio (Ateriana) stygnusLocard (1889)
Unio (Groupe de l’U. ater) stygnusLocard (1893)
Unio consentaneus var. moquiniGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus moquinianusPrie (2012)
Unio nanusLopes-Lima et al. (2024)

Unio valliericus Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 139 [33], 199 [93].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 165.
Type(s): SYNTYPE MNHN_22592, l’Albane (Côte-d’Or).

Unio valliericusLocard (1889), Westerlund (1890)
Unio (Nubilusiana) valliericusLocard (1889)
Unio (Groupe de l’U. locardianus) valliericusLocard (1893)
Unio mancus var. elongatulusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio zoasthenus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 128 [22], 186 [80].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 155.
Type(s): LECTOTYPE MNHN_22602, Bois-Vieux (Jura).

Unio zoasthenusLocard (1889), Westerlund (1890), Rochebrune (1903)
Unio (Melasiana) zoasthenusLocard (1889)
Unio (Groupe de l’U. melas) zoasthenusLocard (1893)
Unio mancusGermain (1931)
Unio crassus batavusHaas (1940), Haas (1969)
Unio mancus mancusPrie (2012)
Unio nanusLopes-Lima et al. (2024)

Unio hassiae Haas, 1908
Haas, 1908, Nachr. Bl. Dtsch. Malak. Ges. 40: 175.

Unio hassiaeHaas (1908)
Unio batavus hassiaeKobelt & Haas (1910)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio pseudocrassus Haas, 1909
Haas, 1909, Beitr. Kenntn. Mittleleurop. Najadeen, Beilage zu... 41: 29.

Unio pseudocrassusHaas (1909)
Unio batavus pseudocrassusKobelt & Haas (1910)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio brandisi Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 82, pl. 419, figs. 2434-2435.

Unio brandisiKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio nanusLopes-Lima et al. (2024)

Unio travnicensis Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 83, pl. 420, figs. 2436-2438.

Unio travnicensisKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus badensis Haas, 1910
Haas, 1910, Mitteleur. Najad., Beiträge Nachr. Bl. Dtsch. Malak. Ges. (4): 60.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus badensisHaas (1910)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus sabulosus Haas, 1910
Haas, 1910, Mitteleur. Najad., Beiträge Nachr. Bl. Dtsch. Malak. Ges. (4): 59.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus sabulosusHaas (1910)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus forma elongata Clessin, 1910
Clessin, 1910, Nachr. Bl. Dtsch. Malak. Ges. 42: 66.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio elongata Lamarck, 1819)

Unio batavus forma elongataClessin (1910)
Unio consentaneus elongatusKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus abnobae Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 23, pl. 458, figs. 2473-2477.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus abnobaeKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus biandeti Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 22, pl. 457, fig. 2470.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus biandetiKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus bodamicus Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 21, pl. 456, figs. 2465-2466.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus bodamicusKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus dilatatus Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 18, pl. 453, figs. 2447-2448; 23, pl. 457, fig. 2472.

Unio consentaneus dilatatusKobelt & Haas (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus elongatus Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 20, pl. 455, figs. 241.
(homonym of Unio elongata Lamarck, 1819; intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus elongatusKobelt & Haas (1910)
Unio crassus cythereaHaas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus hohenwarti Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 78, pl. 418, fig. 2427.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus hohenwartiKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus langi Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 70, pl. 415, fig. 2414.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus langiKobelt (1910)
Unio consentaneus var. langiKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus marisaensis Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 57, pl. 479, figs. 2559-2562.
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: pl. 479 [caption, as “marisensis”].

Unio consentaneus marisaensisKobelt & Haas (1910)
Unio crassus cythereaHaas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus subterminalis Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 80, pl. 418, fig. 2429.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus subterminalisKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio consentaneus thayacus Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 69, pl. 414, figs. 2410-2411; pl. 415, fig. 2413.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836)

Unio consentaneus thayacusKobelt (1910)
Unio consentaneus var. thayacusKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio (consentaneus var. ?) gallensteini Kobelt, 1910
Kobelt, 1910, Icon. Land- und Süssw.-Moll. 15: 79, pl. 418, fig. 2428.
(intro'd as intra-sp. of Unio consentaneus Rossmässler, 1836; homonym of Unio lijacensis var. gallensteini Kobelt, 1890)

Unio (consentaneus var. ?) gallensteiniKobelt (1910)
Unio crassus cythereaHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus hexameri Haas, 1911
Haas, 1911, Verh. Naturhist. Ver. Preuß. Rheinl. 68: 526, pl. 4, fig. 2.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus hexameriHaas (1911)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus palatinus Haas, 1911
Haas, 1911, Verh. Naturhist. Ver. Preuß. Rheinl. 68: 525, pl. 4, fig. 3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus palatinusHaas (1911)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus probavaricus Haas, 1911
Haas, 1911, Verh. Naturhist. Ver. Preuß. Rheinl. 68: 526, pl. 4, fig. 9.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus probavaricusHaas (1911)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus forma pseudoconsentaneus Geyer, 1911
Geyer, 1911, Jh. Ver. Vaterl. Naturkde. Württemb. 67: 365, pl. 5, fig. 3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus forma pseudoconsentaneusGeyer (1911)
Unio crassus batavusHaas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus distinguendus C. Boettger, 1912
C. Boettger, 1912, Arch. Naturg. 78: 305, pl. 1, fig. 7.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus distinguendusC. Boettger (1912)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus navensis C. Boettger, 1912
C. Boettger, 1912, Arch. Naturg. 78: 305, pl. 5, figs. 1-3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus navensisC. Boettger (1912)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio batavus kobeltianus Haas, 1913
Haas, 1913, Nachr. Bl. Dtsch. Malak. Ges. 45: 105.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus kobeltianusHaas (1913)
Unio crassus batavusHaas (1940), Haas (1969)
Unio nanusLopes-Lima et al. (2024)

Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
(misidentified reference to Unio crassus Retzius, 1788)

Unio crassusSoroka (2005)
Unio nanusLopes-Lima et al. (2024)

Taxonomic History (since 1900)

Unio (sec. Lymnium) littoralisSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) batavusSimpson (1900), Simpson (1914)
Unio (sec. Lymnium) pictorumSimpson (1900), Simpson (1914)
Unio nubilusRochebrune (1903)
Unio zoasthenusRochebrune (1903)
Unio (Rhomboïdiana) nanaGermain (1904)
Unio (Socardiana) ripariusGermain (1904)
Unio (Batavusiana) carynthiacusGermain (1904)
Unio (Fusculusiana) fusculusGermain (1904)
Unio (Melasiana) melasGermain (1904)
Unio (Batavusiana) potamiusGermain (1904)
Unio (Groupe des Ninusiana) nanusCaziot (1907)
Unio (Groupe des Amnicusiana) amnicusCaziot (1907)
Unio (Groupe des Mancusiana) dubisianusCaziot (1907)
Unio hassiaeHaas (1908)
Unio pseudocrassusHaas (1909)
Unio nanaKobelt & Haas (1910)
Unio consentaneus ovatusKobelt & Haas (1910)
Unio consentaneus minorKobelt & Haas (1910)
Unio consentaneusKobelt (1910), Germain (1931)
Unio consentaneus var.Kobelt & Haas (1910)
Unio consentaneus squamosusKobelt (1910), Kobelt & Haas (1910)
Unio heldiiKobelt & Haas (1910)
Unio consentaneus ostiorumKobelt & Haas (1910)
Unio batavus taunicusHaas (1910)
Unio batavus hassiaeKobelt & Haas (1910)
Unio batavus pseudocrassusKobelt & Haas (1910)
Unio brandisiKobelt (1910)
Unio travnicensisKobelt (1910)
Unio batavus badensisHaas (1910)
Unio batavus sabulosusHaas (1910)
Unio batavus forma elongataClessin (1910)
Unio consentaneus elongatusKobelt & Haas (1910)
Unio consentaneus abnobaeKobelt & Haas (1910)
Unio consentaneus biandetiKobelt & Haas (1910)
Unio consentaneus bodamicusKobelt & Haas (1910)
Unio consentaneus dilatatusKobelt & Haas (1910)
Unio consentaneus hohenwartiKobelt (1910)
Unio consentaneus langiKobelt (1910)
Unio consentaneus var. langiKobelt (1910)
Unio consentaneus marisaensisKobelt & Haas (1910)
Unio consentaneus subterminalisKobelt (1910)
Unio consentaneus thayacusKobelt (1910)
Unio consentaneus var. thayacusKobelt (1910)
Unio (consentaneus var. ?) gallensteiniKobelt (1910)
Unio batavus hexameriHaas (1911)
Unio batavus palatinusHaas (1911)
Unio batavus probavaricusHaas (1911)
Unio batavus forma pseudoconsentaneusGeyer (1911)
Unio crassus consentaneusOrtmann (1912)
Unio batavus distinguendusC. Boettger (1912)
Unio batavus navensisC. Boettger (1912)
Unio cytherea cythereaHaas (1913)
Unio cytherea nessorhynchusHaas (1913)
Unio batavus kobeltianusHaas (1913)
Unio mancus var. nanusGermain (1931)
Unio crassusGermain (1931), Soroka (2005), Graf (2007), Anisimova et al. (2018), Araujo et al. (2018), Bogatov et al. (2018)
Unio crassus var. piscinalisGermain (1931)
Unio batavusGermain (1931)
Unio mancus var. amnicus form ripariusGermain (1931)
Unio mancus var. amnicusGermain (1931)
Unio mancus var. macrorhynchusGermain (1931)
Unio mancusGermain (1931)
Unio mancus var. elongatulusGermain (1931)
Unio consentaneus var. moquiniGermain (1931)
Unio (s.s.) crassus consentancusZhadin (1938)
Unio crassus batavusHaas (1940), Haas (1969), Lopes-Lima et al. (2017)
Unio crassus crassusHaas (1940), Haas (1969), Klishko et al. (2017), Lopes-Lima et al. (2017)
Unio crassus cythereaHaas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Batavusiana (s.s.) nanaZatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) fusculaZatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) fuscula fusculaStadnichenko (1984)
Unio (Crassiana) crassus nanusFalkner et al. (2001), Falkner et al. (2002)
Unio (Crassiana) crassus ripariusFalkner et al. (2001), Falkner et al. (2002)
Unio (Crassiana) crassus cythereaFalkner et al. (2001), Falkner et al. (2002)
Unio (Crassiana) crassus decurvatusFalkner et al. (2001)
Unio (Crassiana) crassus albensisFalkner et al. (2001)
Unio (Crassiana) crassus ondovensisFalkner et al. (2001)
Crassiana (Bataviana) nanaStarobogatov et al. (2004)
Crassiana (Bataviana) fusculaStarobogatov et al. (2004)
Crassiana nana nanaKantor & Sysoev (2005), Kantor et al. (2010)
Crassiana fusculaKantor & Sysoev (2005), Kantor et al. (2010)
Unio mancus mancusPrie (2012)
Unio mancus moquinianusPrie (2012)
Unio pictorumHe & Zhuang (2013)
Unio crassus nanusPrié & Puillandre (2014)
Unio (Crassiana) nanaVinarski & Kantor (2016)
Unio (s.s.) pictorumVinarski & Kantor (2016)
Unio (Crassiana) crassusVinarski & Kantor (2016)
Unio (Crassiana) fusculusVinarski & Kantor (2016)
Unio crassus nanaBogatov (2022)
Unio crassus musivusBogatov (2022)
Unio nanusLopes-Lima et al. (2024)

Page updated: 08:54:06 Wed 06 Nov 2024

 
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