|
Bivalvia | Unionoida | Unionoidea | Unionidae subfamily AMBLEMINAE Rafinesque, 1820: cladomics
5 tribes
Cladomics Summary
number of species = 416
number of subtaxa (rank: tribe) = 5
analyzed in 289 cladograms in 172 publications
maximum number of terminals in an analysis = 90, mean = 11.07
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.59
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 248
maximum number of included terminals = 90, mean = 12.74
maximum number of subtaxa (tribe) = 5, mean = 2.85
proportion of cladograms that support basic monophyly = 0.88
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 176
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 90, mean = 10.81
maximum number of subtaxa (tribe) = 5, mean = 3.06
proportion of cladograms that support rigorous monophyly = 0.82
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Wu et al., 2024 [Publ 4350], fig. 8a_S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
5 |
75 |
69 |
3 |
0.0 |
*** Top Monophyly Cladogram ***
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
125 |
70 |
4 |
173 |
134 |
3 |
128.0 |
*** Top Sister-Group Cladogram ***
+-------------- Parreysiinae
|
| +----------- Chamberlainia
| |
| +----------- Megalonaias **
| |
| +----------- Uniomerus **
| |
--+ +----------- Quadrulina **
| |
| +----------- Sinohyriopsis
| |
| +----------- Lamprotulini
| |
| +----------- Gonideini
| |
| +----------- Pseudodontini
+--+
+----------- Unioninae
|
| +-- Rectidentini
+--------+
| +-- Contradentini
|
| +-------- Amblema **
| |
| +-------- Toxolasma **
| |
| +-------- Pleurobemini **
| |
| | +-- Obliquaria **
| +-----+
| | +-- Plectomerus **
+--+
| +----- Ellipsaria **
| |
| +----- Potamilus_leptodon **
| |
| +----- Medionidus **
| |
| +----- Potamilus_alatus **
| |
| +----- Ptychobranchus **
+--+
+----- Truncilla **
|
+----- Venustaconcha **
|
+----- Cambarunio **
|
+----- Leaunio **
|
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 90 |
90 |
81 |
5 |
12 |
10 |
2 |
99.0 |
+-- Chamberlainiini
+-----------------------------+
| +-- Unioninae
|
--+ +-- Gonideini
| +--------------------------+
| | +-- Lamprotulini
| |
+--+ +-------------------------- Quadrulini **
| |
| | +----------------------- Pleurobemini **
+--+ |
| | +----- Popenaiadini **
| | +--------------+
| | | | +-- Amblema **
+--+ | +--+
| | +-- Reginaia **
| |
| | +-- Plectomerus **
| | +--------------+
+--+ | +-- Toxolasma **
| |
| | +----- Obliquaria **
| | +--------+
| | | | +-- Cyrtonaias **
| | | +--+
| | | +-- Glebula **
| | |
+--+ | +----------- Sagittunio **
| | |
| | +----------- Leaunio **
| | |
| | +----------- Potamilus **
| | |
| | +----------- Hamiota **
| | |
| | | +-- Ellipsaria **
+--+ +--------+
| | +-- Truncilla **
| |
| | +----- Lampsilis_teres **
| +-----+
| | | +-- Ortmanniana_ligamentina **
| | +--+
| | +-- Lampsilis_siliquoidea **
| |
| | +----- Villosa **
| +-----+
| | | +-- Ligumia **
| | +--+
| | +-- Cambarunio **
| |
+--+ +-------- Ortmanniana_pectorosa **
+--+
| | +----- Lampsilis_ornata **
| +--+
| | +-- Lampsilis_cardium **
| +--+
| +-- Lampsilis_ovata **
|
| +-------- Obovaria_olivaria **
| |
| +-------- Venustaconcha_ellipsiformis__2 **
+--+
| +-------- Epioblasma **
| |
| | +----- Obovaria_unicolor **
| +--+
| | +-- Obovaria_arkansasensis **
| +--+
| +-- Obovaria_subrotunda **
|
| +-- Ptychobranchus **
| +-----+
| | +-- Venustaconcha_ellipsiformis__1 **
| |
+--+ +----- Medionidus_conradicus **
| |
| | +-- Lemiox **
+--+--+
| +-- Medionidus_acutissimus **
|
| +-- Dromus **
+--+
+-- Cyprogenia **
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 90 |
90 |
92 |
5 |
12 |
10 |
2 |
102.0 |
+-------------------- Chamberlainiini
|
+-------------------- Unioninae
--+
| +-- Gonideini
| +--------------+
| | +-- Lamprotulini
+--+
| +-------------- Quadrulini **
| |
+--+ +----------- Pleurobemini **
| |
+--+ +-- Plectomerus **
| +-----+
| | +-- Reginaia **
+--+
| +----- Lampsilini **
+--+
| +-- Popenaiadini **
+--+
+-- Amblema **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 82 |
82 |
59 |
5 |
22 |
21 |
2 |
179.0 |
+-------------- Unioninae
|
| +----------- Gonideinae
--+ |
| | +-------- Reginaia **
| | |
+--+ +-------- Amblema **
| |
| | +-- Quadrulini **
| +-----+
| | +-- Pleurobemini **
+--+
| +----- Popenaiadini **
| |
| | +-- Dromus **
| | |
| | +-- Epioblasma **
| | |
+--+ +-- Glebula **
| |
| +-- Lampsilis_ovata **
| |
| +-- Lampsilis_virescens **
| |
| +-- Lemiox **
+--+
+-- Obliquaria **
|
+-- Obovaria_olivaria **
|
+-- Obovaria_unicolor **
|
+-- Plectomerus **
|
+-- Potamilus_alatus **
|
+-- Potamilus_capax **
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 73 |
73 |
67 |
5 |
0 |
0 |
0 |
72.0 |
+----------------------------------- Quadrulini **
|
--+ +-------------------------------- Pleurobemini **
| |
| | +-- Amblema **
+--+ +--------------------------+
| | +-- Reginaia **
| |
+--+ +-------------------------- Popenaiadini **
| |
| | +-- Glebula **
+--+ +--------------------+
| | +-- Paetulunio **
| |
+--+ +-- Toxolasma **
| +-----------------+
| | +-- Plectomerus **
| |
+--+ +----------------- Obliquaria **
| |
| | +----- Potamilus **
+--+ +--------+
| | | +-- Dromus **
| | +--+
+--+ +-- Lemiox **
|
| +----------- Lampsilis_ovata **
| |
+--+ +-------- Lampsilis_virescens **
| |
+--+ +-- Epioblasma **
| +--+
| | +-- Obovaria_arkansasensis **
+--+
| +-- Obovaria_olivaria **
+--+
+-- Venustaconcha **
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 73 |
73 |
63 |
5 |
0 |
0 |
0 |
75.0 |
+-------------- Quadrulini **
|
--+ +----------- Pleurobemini **
| |
+--+ +-- Plectomerus **
| +-----+
| | +-- Reginaia **
+--+
| +----- Lampsilini **
+--+
| +-- Popenaiadini **
+--+
+-- Amblema **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 55 |
55 |
47 |
4 |
2 |
1 |
1 |
55.0 |
+----- Pleurobemini **
|
--+ +-- Amblemini **
| |
+--+-- Popenaiadini **
|
+-- Lampsilini **
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 54 |
54 |
49 |
4 |
2 |
1 |
1 |
55.0 |
+----------- Unioninae
--+
| +-------- Lampsilini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Popenaiadini **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
42 |
5 |
0 |
0 |
0 |
59.0 |
+-------------------------------- Quadrulini **
|
| +-------------- Elliptoideus **
| +--------------+
| | | +----------- Plectomerus **
--+ | +--+
| | | +-------- Obovaria_unicolor **
| | +--+
| | | +----- Obovaria_olivaria **
| | +--+
| | | +-- Lampsilis **
+--+ +--+
| +-- Lemiox **
|
| +-------- Cyrtonaias **
| +-----------------+
| | | +----- Reginaia **
| | +--+
| | | +-- Popenaiadini **
+--+ +--+
| +-- Amblema **
|
| +----------------------- Elliptio **
| |
+--+ +-------------------- Eurynia **
| |
| | +----------------- Parvaspina **
+--+ |
| | +-- Hemistena **
| +--------------+
+--+ +-- Plethobasus **
|
| +-------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum **
+--+ +--------+
| | +-- Fusconaia **
| |
+--+ +-- Pleurobema_stabile **
| +-----+
| | +-- Pleurobema_sintoxia **
+--+
| +----- Pleurobema_decisum **
+--+
| +-- Pleurobema_clava **
+--+
+-- Pleurobema_pyriforme **
Campbell & Lydeard, 2012 [Publ 3308], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
41 |
5 |
0 |
0 |
0 |
52.0 |
+-------------------- Quadrulini **
|
--+ +----------------- Pleurobemini **
| |
| | +-- Popenaiadini **
+--+ +-----------+
| | +-- Amblema **
| |
+--+ +-- Cyrtonaias **
| +--------+
| | +-- Reginaia **
+--+
| +-------- Plectomerus **
+--+
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
Campbell & Lydeard, 2012 [Publ 3308], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
36 |
5 |
0 |
0 |
0 |
50.0 |
+-------------------- Quadrulini **
|
--+ +----------------- Pleurobemini **
| |
+--+ +-- Reginaia **
| +-----------+
| | +-- Amblema **
+--+
| +----------- Popenaiadini **
| |
+--+ +-------- Cyrtonaias **
| |
+--+-------- Plectomerus **
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
Campbell & Lydeard, 2012 [Publ 3308], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
40 |
5 |
0 |
0 |
0 |
52.0 |
+----------------- Quadrulini **
|
--+ +-------------- Pleurobemini **
| |
| | +-- Popenaiadini **
+--+ +--------+
| | +-- Amblema **
| |
+--+ +-------- Plectomerus **
| |
| | +-- Cyrtonaias **
+--+-----+
| +-- Reginaia **
|
| +----- Lemiox **
+--+
| +-- Lampsilis **
+--+
+-- Obovaria **
Inoue et al., 2020 [Publ 3941], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
41 |
4 |
1 |
1 |
1 |
43.0 |
+----------- Unioninae
--+
| +-------- Quadrulini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Smith et al., 2019 [Publ 3951], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
40 |
4 |
0 |
0 |
0 |
43.0 |
+-------------------------------------- Quadrulini **
|
| +-- Amblema **
--+ +--------------------------------+
| | +-- Reginaia **
| |
+--+ +-------------------------------- Plectomerus **
| |
| | +----------------------------- Toxolasma **
+--+ |
| | +----- Popenaiadini **
| | +--------------------+
+--+ | | +-- Glebula **
| | +--+
| | +-- Cyrtonaias **
| |
+--+ +----------------------- Obliquaria **
| |
| | +-- Medionidus **
| | +--+
| | | +-- Lemiox **
+--+ +--------------+
| | | +-- Cyprogenia **
| | +--+
| | +-- Ptychobranchus **
| |
+--+ +-------- Potamilus **
| +--------+
| | | +----- Atlanticoncha **
| | +--+
| | | +-- Ellipsaria **
| | +--+
+--+ +-- Truncilla **
|
| +-- Hamiota **
| +-----------+
| | +-- Villosa **
| |
+--+ +----------- Ligumia **
| |
| | +----- Venustaconcha **
+--+ +--+
| | | +-- Obovaria **
| | +--+
+--+ +-- Epioblasma **
|
| +----- Ortmanniana_pectorosa **
+--+
| +-- Ortmanniana_ligamentina **
+--+
+-- Lampsilis **
Burlakova et al., 2012 [Publ 3486], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 41 |
41 |
42 |
4 |
0 |
0 |
0 |
57.0 |
+----------- Amblema **
|
--+ +-- Lampsilis **
| +-----+
| | +-- Plectomerus **
+--+
| +----- Popenaiadini **
+--+
| +-- Cyrtonaias **
+--+
+-- Pleurobemini **
Inoue et al., 2020 [Publ 3941], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
40 |
4 |
1 |
1 |
1 |
44.0 |
+-------------- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Reginaia **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblema **
+--+
+-- Lampsilini **
Pfeiffer et al., 2016 [Publ 3660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 37 |
37 |
33 |
4 |
1 |
1 |
1 |
47.0 |
+-------------------------------- Unioninae
|
--+ +----------------------------- Amblemini **
| |
+--+ +-------------------------- Lampsilini **
| |
+--+ +----------------------- Elliptoideus **
| |
+--+ +-------------------- Quadrulini **
| |
+--+ +----------------- Elliptio **
| |
| | +-------------- Parvaspina **
+--+ |
| +-------------- Plethobasus **
| |
| | +-- Hemistena **
+--+-----------+
| +-- Eurynia **
|
| +----------- Fusconaia **
| |
| | +----- Pleurobema_riddellii **
+--+ +--+
| | | +-- Pleurobema_sintoxia **
| | +--+
+--+ +-- Pleurobema_cordatum **
|
| +----- Pleuronaia **
| |
+--+ +-- Pleurobema_pyriforme **
| |
+--+-- Pleurobema_strodeanum **
|
+-- Pleurobema_clava **
Perkins et al., 2017 [Publ 3777], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
2 |
0 |
0 |
0 |
48.0 |
+-- Amblema **
|
--+-- Plectomerus **
|
+-- Pleurobemini **
Burlakova et al., 2012 [Publ 3486], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
26 |
2 |
0 |
0 |
0 |
51.0 |
+-- Amblemini **
--+
+-- Pleurobemini **
Burlakova et al., 2012 [Publ 3486], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
30 |
1 |
0 |
0 |
0 |
53.0 |
-- (Pleurobemini
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 34 |
34 |
33 |
5 |
1 |
1 |
1 |
42.0 |
+-------------------------------------- Gonideinae
|
--+ +----------------------------------- Quadrulini **
| |
+--+ +-------------------------------- Pleurobemini **
| |
+--+ +----------------------------- Amblemini **
| |
| | +-- Popenaias **
+--+ +-----------------------+
| | +-- Psoronaias **
| |
+--+ +-- Cyrtonaias_tampicoensis **
| +--------------------+
| | +-- Glebula **
| |
+--+ +-------------------- Potamilus **
| |
| | +-- Truncilla **
+--+ +--------------+
| | +-- Ptychobranchus **
| |
+--+ +-------------- Pachynaias **
| |
| | +-- Cyrtonaias_explicata **
+--+ +--------+
| | +-- Sphenonaias **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Ortmanniana **
+--+
| +----- Cambarunio **
+--+
| +-- Epioblasma **
+--+
+-- Ligumia **
Burlakova et al., 2019 [Publ 3899], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 33 |
33 |
21 |
5 |
0 |
0 |
0 |
38.0 |
+-------------------------- Quadrulini **
|
| +----------------------- Plectomerus **
--+ |
| | +-------------------- Popenaiadini **
| | |
| | +-------------------- Pleurobemini **
+--+ |
| | +-- Cyrtonaias **
| | +--+
| | | +-- Glebula **
| +--------------+
| | | +-- Reginaia **
+--+ +--+
| +-- Obliquaria **
|
| +----------------- Toxolasma **
| |
| | +-------------- Potamilus **
| | |
| | | +-- Ellipsaria **
| | +-----------+
+--+ | +-- Truncilla **
| |
| | +----- Medionidus **
| +--------+
| | | +-- Lemiox **
| | +--+
+--+ +-- Dromus **
|
| +----------- Cambarunio **
| |
| +----------- Leaunio **
| |
| +----------- Villosa **
| |
| | +-- Sagittunio **
| +--------+
+--+ +-- Lampsilis_ovata **
|
| +----- Venustaconcha **
+-----+
| | +-- Epioblasma **
| +--+
| +-- Obovaria **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Lampsilis_hydiana **
+--+
+-- Lampsilis_straminea **
Burdick & White, 2007 [Publ 2679], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 30 |
30 |
31 |
3 |
0 |
0 |
0 |
46.0 |
+-------------------------------- Lampsilini **
|
--+ +----------------------------- Parvaspina **
| |
| | +-------------------------- Amblemini **
+--+ |
| | +----- Eurynia **
| | +-----------------+
+--+ | | +-- Plethobasus **
| | +--+
| | +-- Elliptio **
| |
| | +-------------------- Pleurobema_plenum **
+--+ |
| +-------------------- Pleuronaia **
| |
| | +-- Pleurobema_cordatum__1 **
| | +-----+
| | | +-- Fusconaia **
+--+ +--------+
| | | +----- Pleurobema_sintoxia__1 **
| | | |
| | +--+ +-- Pleurobema_cordatum__2 **
| | | |
| | +--+-- Pleurobema_sintoxia__2 **
| | |
+--+ +-- Pleurobema_sintoxia__3 **
|
| +-------------- Pleurobema_stabile **
| |
| | +-- Pleurobema_rubellum **
| | +-----+
| | | +-- Pleurobema_strodeanum **
+--+ +--+
| | | +----- Pleurobema_beadleianum **
| | +--+
| | | +-- Pleurobema_perovatum **
| | +--+
+--+ +-- Pleurobema_taitianum **
|
| +-- Pleurobema_clava **
| +-----+
| | +-- Pleurobema_oviforme **
| |
+--+ +-- Pleurobema_decisum **
| +--+
| | +-- Pleurobema_hanleyianum **
+--+
| +-- Pleurobema_pyriforme **
+--+
+-- Pleurobema_georgianum **
Serb et al., 2003 [Publ 2659], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
4 |
0 |
0 |
0 |
40.0 |
+-------------------------- Lampsilis **
|
--+ +----------------------- Potamilus **
| |
| | +----- Medionidus **
+--+ +--------------+
| | | +-- Dromus **
| | +--+
+--+ +-- Cyprogenia **
|
| +----------------- Obliquaria **
| |
+--+ +-------------- Amblema **
| |
| | +----- Reginaia **
+--+ +-----+
| | | +-- Elliptoideus **
| | +--+
+--+ +-- Plectomerus **
|
| +-------- Quadrulini **
+--+
| +----- Hemistena **
+--+
| +-- Elliptio **
+--+
+-- Fusconaia **
Boyer et al., 2011 [Publ 2718], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
25 |
4 |
9 |
9 |
1 |
36.0 |
+----------------------------------- Unioninae
|
--+ +-------------------------------- Quadrulina **
| |
+--+ +----------------------------- Megalonaias **
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +----------------------- Amblema **
| |
| | +-------------------- Pleurobemini **
+--+ |
| | +-- Reginaia **
| +-----------------+
+--+ +-- Obliquaria **
|
| +----------------- Potamilus **
| |
| | +-- Ellipsaria **
+--+ +-----------+
| | +-- Truncilla **
| |
+--+ +----------- Lampsilis_cardium **
| |
| | +----- Ligumia **
| +-----+
+--+ | +-- Obovaria **
| +--+
| +-- Epioblasma **
|
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis_higginsii **
Serb et al., 2003 [Publ 2659], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
20 |
4 |
0 |
0 |
0 |
38.0 |
+----------- Lampsilis **
|
| +-------- Obliquaria **
| |
| +-------- Amblema **
| |
--+ +-------- Reginaia **
| |
| +-------- Potamilus **
| |
| +-------- Quadrulini **
| |
+--+ +-- Elliptoideus **
+-----+
| +-- Plectomerus **
|
| +----- Medionidus **
+--+
| | +-- Dromus **
| +--+
| +-- Cyprogenia **
|
| +----- Hemistena **
+--+
| +-- Elliptio **
+--+
+-- Fusconaia **
Boyer et al., 2011 [Publ 2718], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
22 |
4 |
6 |
6 |
1 |
35.0 |
+----------- Unioninae
|
--+ +-------- Reginaia **
| |
+--+ +-- Pleurobemini **
| +--+
| | +-- Quadrulini **
+--+
| +-- Amblema **
+--+
+-- Lampsilini **
Smith et al., 2020 [Publ 4129], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
5 |
0 |
0 |
0 |
32.0 |
+----------- Quadrulini **
--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Smith et al., 2020 [Publ 4129], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
5 |
0 |
0 |
0 |
32.0 |
+----------- Quadrulini **
--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Campbell et al., 2008 [Publ 2675], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
18 |
3 |
0 |
0 |
0 |
41.0 |
+-- Lampsilini **
|
--+-- Amblemini **
|
+-- Pleurobemini **
Campbell et al., 2008 [Publ 2675], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
23 |
3 |
0 |
0 |
0 |
40.0 |
+-- Lampsilini **
|
--+-- Amblemini **
|
+-- Pleurobemini **
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
24 |
5 |
1 |
1 |
1 |
32.0 |
+-------------- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Lampsilini **
+--+
+-- Popenaiadini **
Inoue et al., 2020 [Publ 4128], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 26 |
26 |
17 |
4 |
0 |
0 |
0 |
36.0 |
+-------------------------- Quadrulini **
|
--+ +----------------------- Reginaia **
| |
+--+ +-------------------- Amblema **
| |
| | +-- Popenaias **
+--+ +--------------+
| | +-- Psoronaias **
| |
+--+ +-- Glebula **
| +-----------+
| | +-- Cyrtonaias_tampicoensis **
| |
+--+ +----------- Potamilus **
| |
| | +-- Ptychobranchus **
+--+ +-----+
| | +-- Truncilla **
| |
| | +----- Ortmanniana **
+--+ |
| +----- Epioblasma **
| |
| +----- Lampsilis **
| |
+--+----- Ligumia **
|
+----- Cambarunio **
|
+----- Pachynaias **
|
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
20 |
4 |
173 |
134 |
3 |
23.0 |
+-------------------------------------- Parreysiinae
|
--+ +----------------------------------- Unioninae
| |
| | +-- Rectidentini
+--+ +-----------------------------+
| | +-- Contradentini
| |
| | +-------- Chamberlainiini
+--+ +--------------------+
| | | +----- Lamprotulini
| | +--+
| | | +-- Gonideini
+--+ +--+
| +-- Pseudodontini
|
| +-------------------------- Quadrulini **
| |
+--+ +----------------------- Pleurobemini **
| |
+--+ +-------------------- Amblema **
| |
| | +----- Toxolasma **
+--+ +-----------+
| | | +-- Obliquaria **
| | +--+
| | +-- Plectomerus **
+--+
| +-------------- Potamilus_leptodon **
| |
| | +----------- Potamilus_alatus **
+--+ |
| | +-------- Venustaconcha **
| | |
+--+ +-------- Leaunio **
| |
| | +-- Ellipsaria **
| +-----+
+--+ +-- Truncilla **
|
| +-- Medionidus **
+-----+
| +-- Ptychobranchus **
|
| +----- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
23 |
4 |
171 |
132 |
3 |
22.0 |
+----------------------------------------------------- Parreysiinae
|
--+ +-------------------------------------------------- Pseudodontini
| |
+--+ +-- Rectidentini
| +--------------------------------------------+
| | +-- Contradentini
+--+
| +-------------------------------------------- Unioninae
| |
+--+ +----------------------------------------- Chamberlainiini
| |
+--+ +-- Lamprotulini
| +-----------------------------------+
| | +-- Gonideini
+--+
| +----------------------------------- Quadrulini **
| |
+--+ +-------------------------------- Pleurobemini **
| |
+--+ +----------------------------- Amblema **
| |
+--+ +-------------------------- Toxolasma **
| |
+--+ +-- Obliquaria **
| +--------------------+
| | +-- Plectomerus **
+--+
| +-------------------- Potamilus_leptodon **
| |
+--+ +----------------- Potamilus_alatus **
| |
| | +-- Medionidus **
+--+ +-----------+
| | +-- Ptychobranchus **
| |
+--+ +-- Ellipsaria **
| +--------+
| | +-- Truncilla **
+--+
| +-------- Leaunio **
| |
+--+ +-- Venustaconcha **
| +--+
| | +-- Cambarunio **
+--+
| +-- Ortmanniana **
+--+
+-- Lampsilis **
Campbell et al., 2008 [Publ 2675], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
9 |
4 |
0 |
0 |
0 |
41.0 |
+----------------------- Quadrulini **
|
+----------------------- Amblemini **
|
| +-------------------- Lampsilini **
| |
| +-------------------- Eurynia **
| |
| +-------------------- Fusconaia_cerina **
--+ |
| +-------------------- Fusconaia_flava **
| |
| +-------------------- Hemistena **
| |
| +-------------------- Plethobasus **
| |
| +-------------------- Parvaspina **
| |
+--+-------------------- Pleurobema_cordatum **
|
+-------------------- Pleuronaia **
|
+-------------------- Pleurobema_oviforme__2 **
|
+-------------------- Pleurobema_stabile **
|
+-------------------- Elliptio **
|
| +----------------- Pleurobema_decisum **
| |
| +----------------- Pleurobema_sintoxia **
| |
| +----------------- Pleurobema_clava **
+--+
+----------------- Pleurobema_oviforme__1 **
|
| +-------------- Pleurobema_beadleianum **
| |
+--+ +----------- Pleurobema_pyriforme **
| |
+--+ +-------- Pleurobema_rubellum **
| |
| +-------- Pleurobema_hanleyianum__2 **
+--+
+-------- Pleurobema_georgianum__2 **
|
| +----- Pleurobema_hanleyianum__1 **
+--+
| +-- Pleurobema_georgianum__1 **
+--+
+-- Pleurobema_athearni **
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
23 |
4 |
4 |
2 |
1 |
30.0 |
+-------------- Unioninae
|
--+ +----------- Megalonaias **
| |
+--+ +-- Uniomerus **
| +-----+
| | +-- Quadrulina **
+--+
| +----- Pleurobemini **
+--+
| +-- Lampsilini **
+--+
+-- Amblemini **
Inoue et al., 2020 [Publ 4128], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
27 |
4 |
0 |
0 |
0 |
35.0 |
+----------------------------------------- Quadrulini **
|
--+ +-------------------------------------- Reginaia **
| |
+--+ +----------------------------------- Amblema **
| |
+--+ +-- Popenaias **
| +-----------------------------+
| | +-- Psoronaias **
+--+
| +----------------------------- Cyrtonaias_tampicoensis **
| |
+--+ +-------------------------- Ptychobranchus **
| |
+--+ +----------------------- Ligumia **
| |
+--+ +-------------------- Potamilus **
| |
+--+ +----------------- Pachynaias **
| |
| | +-- Epioblasma **
+--+ +-----------+
| | +-- Truncilla **
| |
+--+ +-- Ortmanniana **
| +--------+
| | +-- Lampsilis **
+--+
| +-------- Cambarunio **
+--+
| +----- Sphenonaias_microdon__2 **
+--+
| +-- Cyrtonaias_explicata **
+--+
+-- Sphenonaias_microdon__1 **
Campbell et al., 2008 [Publ 2675], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 25 |
25 |
6 |
4 |
0 |
0 |
0 |
44.0 |
+----------- Quadrulini **
|
+----------- Amblemini **
|
| +-- Elliptio_arctata **
| +-----+
| | +-- Elliptio_crassidens **
| |
| | +----- Pleurobema_decisum **
--+ | |
| | +----- Pleurobema_rubellum **
| | |
| | +----- Pleurobema_georgianum__2 **
| | |
| | +----- Pleurobema_stabile **
| | |
| | +----- Pleurobema_pyriforme **
| | |
+--+ +----- Pleurobema_oviforme__2 **
| |
| +----- Pleurobema_hanleyianum__2 **
| |
| +----- Pleuronaia **
| |
| +----- Pleurobema_georgianum__3 **
| |
| +----- Pleurobema_cordatum **
| |
| +----- Pleurobema_beadleianum **
| |
| +----- Hemistena **
+--+
+----- Fusconaia_flava **
|
+----- Fusconaia_cerina **
|
+----- Eurynia **
|
+----- Elliptio_arca **
|
+----- Lampsilini **
|
| +-- Plethobasus **
+--+
| +-- Parvaspina **
|
| +-- Pleurobema_sintoxia **
| |
+--+-- Pleurobema_oviforme__1 **
| |
| +-- Pleurobema_clava **
|
| +-- Pleurobema_athearni **
| |
+--+-- Pleurobema_georgianum__1 **
|
+-- Pleurobema_hanleyianum__1 **
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
21 |
4 |
4 |
2 |
1 |
33.0 |
+----------- Unioninae
--+
| +-------- Amblemini **
+--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
Perkins et al., 2017 [Publ 3777], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 24 |
24 |
10 |
3 |
0 |
0 |
0 |
40.0 |
+-- Amblemini **
|
--+-- Quadrulini **
|
+-- Pleurobemini **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 22 |
22 |
20 |
4 |
4 |
2 |
1 |
29.0 |
+----------- Unioninae
--+
| +-------- Quadrulini **
+--+
| +----- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
20 |
4 |
12 |
10 |
1 |
31.0 |
+----------------------------- Unioninae
|
| +-- Megalonaias **
| +-----------------------+
--+ | +-- Quadrulina **
| |
| | +----------- Elliptoideus **
| | +-----------+
+--+ | | +-------- Pleurobema **
| | +--+
| | | +----- Uniomerus **
| | +--+
| | | +-- Fusconaia **
+--+ +--+
| +-- Eurynia **
|
| +-------------------- Amblema **
| |
| | +----- Toxolasma **
| | +-----------+
+--+ | | +-- Plectomerus **
| | +--+
| | +-- Obliquaria **
| |
+--+ +----- Ptychobranchus **
| +--------+
| | | +-- Potamilus_fragilis **
| | +--+
| | +-- Medionidus **
+--+
| +-- Villosa **
| +--------+
| | +-- Ellipsaria **
+--+
| +-------- Lampsilis_teres **
+--+
| +----- Lampsilis_siliquoidea **
+--+
| +-- Potamilus_purpuratus **
+--+
+-- Potamilus_alatus **
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
5 |
1 |
1 |
1 |
26.0 |
+----------------- Gonideinae
--+
| +-------------- Quadrulini **
+--+
| +----------- Pleurobemini **
+--+
| +-------- Amblema **
+--+
| +----- Plectomerus **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
5 |
1 |
1 |
1 |
26.0 |
+----------------- Gonideinae
--+
| +-------------- Quadrulini **
+--+
| +----------- Pleurobemini **
+--+
| +-------- Amblema **
+--+
| +----- Plectomerus **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Smith et al., 2020 [Publ 4136], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
13 |
2 |
0 |
0 |
0 |
33.0 |
+-- Quadrulini **
--+
+-- Pleurobemini **
Pieri et al., 2018 [Publ 3867], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
22 |
2 |
1 |
1 |
1 |
38.0 |
+----- Unioninae
--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
2 |
0 |
0 |
0 |
27.0 |
+-- Amblemini **
--+
+-- Lampsilini **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
32 |
31 |
4 |
19 |
14 |
2 |
38.0 |
+-------------- Quadrulina **
|
| +----- Lamprotulini
--+ +-----+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Unioninae
|
| +-------- Megalonaias **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
16 |
4 |
10 |
6 |
1 |
30.0 |
+----------------- Unioninae
|
| +-- Megalonaias **
--+ +-----------+
| | +-- Quadrulina **
| |
+--+ +----- Amblema **
| +-----+
| | | +-- Plectomerus **
| | +--+
+--+ +-- Lampsilini **
|
| +-------- Elliptoideus **
+--+
| +----- Uniomerus **
+--+
| +-- Eurynia **
+--+
+-- Pleurobema **
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
13 |
2 |
0 |
0 |
0 |
27.0 |
+-------------------- Obliquaria **
|
+-------------------- Amblemini **
|
| +----------------- Lampsilis_teres **
| |
| +----------------- Obovaria **
--+ |
| | +-- Leaunio **
| +--------------+
| | +-- Cambarunio **
| |
| | +-------- Ortmanniana **
| | |
+--+--------+ +-- Lampsilis_hydiana **
| | +--+
| | | +-- Lampsilis_virescens **
| +--+
| | +-- Lampsilis_siliquoidea **
| +--+
| +-- Lampsilis_radiata **
|
| +-------------- Lampsilis_bracteata **
| |
+--+ +----------- Lampsilis_ornata **
| |
+--+ +-------- Lampsilis_ovata **
| |
+--+ +----- Lampsilis_cardium__3 **
| |
+--+ +-- Lampsilis_satura **
| |
+--+-- Lampsilis_cardium__1 **
|
+-- Lampsilis_cardium__2 **
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
14 |
4 |
9 |
6 |
1 |
30.0 |
+----------------------------- Unioninae
|
| +-- Megalonaias **
--+ +-----------------------+
| | +-- Quadrulina **
| |
| | +----- Elliptoideus **
+--+ |
| +-----------------+ +-- Eurynia **
| | | |
| | +--+-- Pleurobema **
| | |
+--+ +-- Uniomerus **
|
| +-------------------- Amblema **
| |
+--+ +-- Plectomerus **
| +--------------+
| | +-- Obliquaria **
+--+
| +-------------- Potamilus_fragilis **
| |
+--+ +----------- Ptychobranchus **
| |
+--+ +-------- Medionidus **
| |
+--+ +----- Potamilus_purpuratus **
| |
+--+----- Lampsilis **
|
| +-- Ellipsaria **
+--+
+-- Villosa **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
5 |
79 |
68 |
3 |
0.0 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
5 |
79 |
68 |
3 |
0.0 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
14 |
5 |
75 |
69 |
3 |
1.0 |
+----- Gonideinae
+--------+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
11 |
4 |
13 |
8 |
1 |
28.0 |
+----------- Unionini
|
--+ +-------- Anodontini
| |
+--+ +-- Quadrulini **
| +--+
| | +-- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Grobler et al., 2011 [Publ 3359], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
13 |
2 |
0 |
0 |
0 |
26.0 |
+-- Amblemini **
--+
+-- Lampsilini **
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
18 |
16 |
4 |
21 |
16 |
1 |
29.0 |
+----------- Unioninae
|
--+ +-------- Lampsilini **
| |
+--+ +-- Hyriinae
| +--+
| | +-- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
4 |
13 |
8 |
1 |
28.0 |
+----------- Unionini
|
--+ +-------- Anodontini
| |
+--+ +-- Quadrulini **
| +--+
| | +-- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Szumowski et al., 2012 [Publ 3569], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
12 |
2 |
0 |
0 |
0 |
23.0 |
+-- Amblemini **
--+
+-- Lampsilini **
Johnson et al., 2018 [Publ 3844], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
3 |
0 |
0 |
0 |
28.0 |
+----- Amblemini **
--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
Inoue et al., 2020 [Publ 3941], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 16 |
16 |
14 |
1 |
0 |
0 |
0 |
25.0 |
-- (Lampsilini
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
42 |
35 |
3 |
17.0 |
+----- Gonideinae
+--------+
| | +-- Unioninae
| +--+
--+ +-- Parreysiinae
|
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Porto-Hannes et al., 2021 [Publ 4227], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
4 |
0 |
0 |
0 |
23.0 |
+-------- Quadrula_quadrula **
|
--+-------- Quadrula_verrucosa **
|
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
59 |
48 |
2 |
2.0 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
59 |
50 |
2 |
2.0 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
42 |
35 |
3 |
19.0 |
+----------- Gonideinae
|
| +-- Unioninae
+--------+
--+ +-- Parreysiinae
|
| +-------- Quadrulini **
| |
+--+ +-- Pleurobemini **
| +--+
| | +-- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
59 |
48 |
2 |
2.0 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
0 |
0 |
0 |
10.0 |
+----------- Toxolasma **
+--+
| | +-------- Popenaiadini **
| +--+
| | +----- Amblemini **
| +--+
--+ | +-- Quadrulini **
| +--+
| +-- Pleurobemini **
|
| +----------- Potamilus **
| |
+--+ +-- Lampsilis_cardium **
| +-----+
| | +-- Lampsilis_ornata **
+--+
| +----- Venustaconcha **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
5 |
38 |
36 |
2 |
2.0 |
+-- Unioninae
+-----------+
| +-- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
4 |
2 |
2 |
2 |
26.0 |
+----------------- Gonideinae
|
+----------------- Unioninae
|
--+ +-- Pleurobemini **
| +-----------+
| | +-- Quadrulini **
| |
+--+ +-- Paetulunio **
| +--------+
| | +-- Amblemini **
+--+
| +-------- Lampsilis **
| |
+--+ +----- Leaunio **
| |
+--+----- Obovaria **
|
| +-- Villosa **
+--+
+-- Cambarunio **
Szumowski et al., 2012 [Publ 3569], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
0 |
0 |
0 |
30.0 |
-- (Quadrulini
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
4 |
20 |
18 |
2 |
27.0 |
+-------------------------------------------- Unioninae
|
--+ +----------------------------------------- Sinosolenaia
| |
+--+ +-------------------------------------- Lamprotulini
| |
+--+ +----------------------------------- Chamberlainiini
| |
+--+ +-------------------------------- Ptychorhynchus
| |
+--+ +----------------------------- Megalonaias **
| |
| | +----- Elliptoideus **
+--+ +--------------------+
| | | +-- Uniomerus **
| | +--+
+--+ +-- Pleurobema **
|
| +----------------------- Amblema **
| |
+--+ +-- Obliquaria **
| +-----------------+
| | +-- Plectomerus **
+--+
| +----------------- Medionidus **
+--+
| +-------------- Potamilus_fragilis **
+--+
| +----------- Ptychobranchus **
+--+
| +-------- Potamilus_purpuratus **
+--+
| +----- Lampsilis **
+--+
| +-- Villosa **
+--+
+-- Ellipsaria **
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
6 |
4 |
20 |
18 |
2 |
27.0 |
+----------------- Unioninae
|
| +-------------- Ptychorhynchus
| |
--+ +-------------- Chamberlainiini
| |
| +-------------- Lamprotulini
| |
+--+-------------- Sinosolenaia
|
| +----------- Megalonaias **
| |
| | +----- Elliptoideus **
| | +--+
+--+ | | +-- Uniomerus **
| | +--+
| | +-- Pleurobema **
| |
| | +----- Potamilus_fragilis **
+--+ |
| +----- Medionidus **
| |
| +----- Amblema **
| |
| | +-- Obliquaria **
+--+--+
| +-- Plectomerus **
|
| +-- Villosa **
| |
| +-- Ellipsaria **
| |
+--+-- Lampsilis **
|
+-- Potamilus_purpuratus **
|
+-- Ptychobranchus **
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
10 |
4 |
33 |
11 |
3 |
26.0 |
+----------------- Pseudodontini
|
--+ +-------------- Parreysiinae
| |
| +-------------- Contradentini
+--+
+-------------- Unioninae
|
| +----------- Gonideini
+--+
| +-------- Quadrulini **
+--+
| +----- Amblemini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
5 |
5 |
3 |
2 |
3.0 |
+-- Unioninae
+-----------+
| +-- Gonideinae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
4 |
33 |
11 |
3 |
28.0 |
+----------------------------- Unioninae
|
--+ +-------------------------- Contradentini
| |
| | +-- Gonideini
+--+ +--------------------+
| | +-- Pseudodontini
| |
| | +-------- Amblemini **
+--+ +-----------+
| | | +----- Pleurobemini **
| | +--+
| | | +-- Obliquaria **
+--+ +--+
| +-- Quadrulini **
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
10 |
3 |
33 |
11 |
3 |
27.0 |
+-------------------------------- Pseudodontini
|
--+ +----------------------------- Contradentini
| |
+--+ +-------------------------- Unioninae
| |
+--+ +----------------------- Gonideini
| |
| | +----- Pleurobemini **
+--+ +--------------+
| | | +-- Obliquaria **
| | +--+
+--+ +-- Quadrulini **
|
| +----------------- Ptychobranchus **
+--+
| +-------------- Truncilla **
+--+
| +----------- Ortmanniana **
+--+
| +-------- Lampsilis **
+--+
| +----- Cambarunio **
+--+
| +-- Ligumia **
+--+
+-- Epioblasma **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
14 |
12 |
4 |
12 |
12 |
3 |
26.0 |
+-------------------------------- Parreysiinae
|
--+ +----------------------------- Unioninae
| |
+--+ +-------------------------- Lamprotulini
| |
+--+ +----------------------- Quadrulini **
| |
+--+ +-------------------- Gonideini
| |
| | +----- Eurynia **
+--+ +-----------+
| | | +-- Cambarunio **
| | +--+
| | +-- Ptychobranchus **
+--+
| +-- Amblemini **
| +-----------+
| | +-- Pleurobema **
+--+
| +----------- Truncilla **
| |
+--+ +-- Lampsilis **
| +-----+
| | +-- Epioblasma **
+--+
| +----- Ligumia **
+--+
| +-- Obliquaria **
+--+
+-- Ortmanniana **
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
7 |
4 |
33 |
11 |
3 |
25.0 |
+----------------------- Pseudodontini
|
--+ +-------------------- Contradentini
| |
+--+ +----------------- Unioninae
| |
+--+ +-------------- Gonideini
| |
| | +-- Obliquaria **
+--+ +--------+
| | +-- Quadrulini **
| |
+--+ +-------- Amblemini **
| |
| +-------- Pleurobemini **
+--+
| +----- Truncilla **
| |
| +----- Ptychobranchus **
+--+
| +-- Ortmanniana **
| |
| +-- Cambarunio **
+--+
+-- Ligumia **
|
+-- Epioblasma **
Wu et al., 2021 [Publ 4366], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
5 |
57 |
46 |
2 |
5.0 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Lydeard et al., 2000 [Publ 2654], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
11 |
4 |
2 |
1 |
1 |
30.0 |
+-------- Lampsilini **
--+
| +----- Amblemini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
17 |
0 |
4 |
10 |
6 |
2 |
28.0 |
+----- Anodontina
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias **
| |
| +-- Obliquaria **
| |
+--+-- Theliderma **
|
+-- Quadrula **
|
+-- Uniomerus **
|
+-- Amblema **
|
+-- Elliptio **
|
+-- Fusconaia **
|
+-- Plectomerus **
|
+-- Pleurobema **
|
+-- Unionini
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
13 |
11 |
4 |
2 |
1 |
1 |
22.0 |
+-- Quadrulini **
+-----+
| +-- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Amblemini **
+--+
+-- Pleurobemini **
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
5 |
40 |
38 |
2 |
5.0 |
+-- Gonideinae
+-----------+
| +-- Unioninae
--+
| +----------- Quadrulini **
+--+
| +-------- Pleurobemini **
+--+
| +----- Amblemini **
+--+
| +-- Popenaiadini **
+--+
+-- Lampsilini **
Lydeard et al., 2000 [Publ 2654], fig. 2c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
14 |
9 |
4 |
2 |
1 |
1 |
30.0 |
+----- Unioninae
|
+----- Margaritiferinae
|
--+----- Quadrulini **
|
+----- Lampsilini **
|
| +-- Amblemini **
+--+
+-- Pleurobemini **
Lydeard et al., 2000 [Publ 2654], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
14 |
11 |
4 |
2 |
1 |
1 |
31.0 |
+----- Unioninae
|
+----- Margaritiferinae
|
--+----- Lampsilini **
|
+----- Amblemini **
|
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Smith et al., 2020 [Publ 4136], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
1 |
0 |
0 |
0 |
28.0 |
-- (Pleurobemini
Lydeard et al., 2000 [Publ 2654], fig. 2d
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
11 |
4 |
2 |
1 |
1 |
29.0 |
+-------- Amblemini **
--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
5 |
4 |
23 |
7 |
3 |
24.0 |
+-------- Toxolasma **
|
+-------- Amblemini **
|
| +-- Glebula **
+-----+
--+ +-- Cyrtonaias **
|
| +-- Quadrulini **
+-----+
| +-- Pleurobemini **
|
| +----- Potamilus **
+--+
| +-- Ortmanniana **
+--+
+-- Ligumia **
Roe, 2013 [Publ 3571], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
1 |
0 |
0 |
0 |
24.0 |
-- (Lampsilini
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
3 |
43 |
40 |
2 |
6.0 |
+----------- Unioninae
--+
| +-------- Gonideinae
+--+
| +----- Quadrulini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
17 |
0 |
4 |
11 |
7 |
2 |
28.0 |
+----- Anodontina
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea **
| |
| +-- Lampsilis_teres **
| |
| +-- Megalonaias **
| |
| +-- Obliquaria **
| |
+--+-- Theliderma **
|
+-- Quadrula **
|
+-- Uniomerus **
|
+-- Amblema **
|
+-- Elliptio **
|
+-- Fusconaia **
|
+-- Plectomerus **
|
+-- Aculamprotula
|
+-- Unio
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
4 |
20 |
7 |
3 |
25.0 |
+-- Gonideinae
+-----------+
| +-- Unioninae
|
--+ +-- Glebula **
| +--------+
| | +-- Cyrtonaias **
| |
+--+ +-------- Toxolasma **
| |
| | +----- Potamilus **
| +--+
+--+ | +-- Ligumia **
| +--+
| +-- Ortmanniana **
|
| +----- Amblemini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Hoeh et al., 2001 [Publ 2652], fig. 14f2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
28 |
16 |
4 |
20 |
7 |
3 |
39.0 |
+-------------------- Amblemini **
|
+-------------------- Parreysiinae
|
+-------------------- Eurynia **
|
+-------------------- Fusconaia **
|
+-------------------- Gonideinae
|
+-------------------- Pleurobema **
|
+-------------------- Quadrulini **
--+
| +----------- Ortmanniana **
+--------+
| | +-------- Glebula **
| +--+
| | +----- Potamilus **
| +--+
| | +-- Ligumia **
| +--+
| +-- Toxolasma **
|
| +----------------- Cyrtonaias **
| |
+--+ +-------------- Unionini
| |
| | +----------- Anodontini
+--+ |
| | +----- Iridininae
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaeinae
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
Hoeh et al., 2001 [Publ 2652], fig. 14f4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
20 |
7 |
3 |
24.0 |
+-------------- Gonideinae
--+
| +----------- Unioninae
+--+
| +-------- Amblemini **
+--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
15 |
2 |
4 |
8 |
5 |
2 |
29.0 |
+-------- Gonideinae
|
| +-- Elliptio **
| |
| +-- Fusconaia **
| |
--+ +--+-- Amblemini **
| | |
| | +-- Pleurobema **
| | |
| | +-- Unionini
| |
| | +-- Pyganodon_cataracta
+--+ |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia
| |
| +-- Lampsilis_teres **
| |
| +-- Lampsilis_straminea **
+--+
+-- Megalonaias **
|
+-- Obliquaria **
|
+-- Theliderma **
|
+-- Quadrula **
|
+-- Uniomerus **
Porto-Hannes et al., 2021 [Publ 4227], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
6 |
4 |
0 |
0 |
0 |
21.0 |
+-------- Quadrula **
|
--+-------- Pustulosa **
|
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
16 |
12 |
4 |
20 |
7 |
3 |
29.0 |
+----------------------------- Gonideinae
|
--+ +-------------------------- Parreysiinae
| |
+--+ +----------------------- Fusconaia **
| |
| +----------------------- Amblemini **
+--+
| +-------------------- Quadrulini **
| |
+--+ +----------------- Pleurobema **
| |
+--+ +-------------- Eurynia **
| |
| | +-- Cyrtonaias **
+--+-----------+
| +-- Unioninae
|
| +----------- Ortmanniana **
+--+
| +-------- Glebula **
+--+
| +----- Potamilus **
+--+
| +-- Ligumia **
+--+
+-- Toxolasma **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
20 |
7 |
3 |
24.0 |
+----------------- Parreysiinae
--+
| +-------------- Unioninae
+--+
| +----------- Gonideinae
+--+
| +-------- Amblemini **
+--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Stodola et al., 2021 [Publ 4266], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
1 |
0 |
0 |
0 |
25.0 |
-- (Lampsilini
Roe & Lydeard, 1998 [Publ 2693], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
5 |
2 |
0 |
0 |
0 |
22.0 |
+-------------- Pleurobemini **
|
+-------------- Obliquaria **
--+
| +----------- Potamilus_capax **
| |
| +----------- Lampsilis **
+--+
| +-------- Potamilus_fragilis **
| |
| | +----- Potamilus_inflatus **
+--+ |
| | +-- Potamilus_ohiensis **
| +--+
+--+ +-- Potamilus_amphichaenus **
|
| +-- Potamilus_purpuratus__1 **
| |
+--+-- Potamilus_purpuratus__2 **
|
+-- Potamilus_alatus **
Keogh & Simons, 2019 [Publ 3935], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
0 |
0 |
0 |
21.0 |
+----- Lampsilini **
--+
| +-- Amblemini **
+--+
+-- Pleurobemini **
Roe et al., 2001 [Publ 2689], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
25.0 |
-- (Lampsilini
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
47 |
37 |
2 |
7.0 |
+----------- Unioninae
--+
| +-------- Gonideinae
+--+
| +----- Quadrulini **
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
71 |
59 |
3 |
19.0 |
+-------- Gonideinae
|
--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
2 |
1 |
1 |
22.0 |
+----------------- Glebula **
|
--+ +-- Amblema **
| +-----------+
| | +-- Quadrulini **
+--+
| +----------- Plectomerus **
+--+
| +-------- Leaunio **
+--+
| +----- Lampsilis_straminea **
+--+
| +-- Lampsilis_teres **
+--+
+-- Potamilus **
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
11 |
7 |
3 |
14.0 |
+----- Gonideinae
+-----+
| | +-- Parreysiinae
| +--+
--+ +-- Unioninae
|
| +-------- Quadrulini **
+--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
51 |
43 |
4 |
44 |
42 |
2 |
61.0 |
+-------- Quadrulini **
|
+-------- Pleurobemini **
|
| +-- Amblemini **
--+-----+
| +-- Lampsilini **
|
| +-- Lamprotulini
| +--+
| | +-- Gonideini
+--+
| +-- Chamberlainiini
+--+
+-- Unioninae
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
130 |
105 |
3 |
19.0 |
+-------------------- Parreysiinae
|
--+ +-- Rectidentini
| +--------------+
| | +-- Contradentini
+--+
| +-------------- Pseudodontini
| |
+--+ +----------- Unioninae
| |
| | +----- Gonideini
+--+ +--+
| | | +-- Chamberlainiini
| | +--+
| | +-- Lamprotulini
+--+
| +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
67 |
57 |
2 |
19.0 |
+-------------------- Pseudodontini
|
--+ +-- Contradentini
| +--------------+
| | +-- Rectidentini
+--+
| +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Unioninae
| |
+--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
3 |
2 |
1 |
18.0 |
+----------- Unioninae
--+
| +-------- Quadrulini **
+--+
| +----- Plectomerus **
+--+
| +-- Amblema **
+--+
+-- Lampsilini **
Kandl et al., 2001 [Publ 3384], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
0 |
0 |
0 |
21.0 |
+-- Amblemini **
|
--+-- Quadrulini **
|
+-- Pleurobemini **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
130 |
105 |
3 |
19.0 |
+-------------------- Parreysiinae
|
--+ +----------------- Unioninae
| |
| | +-- Contradentini
+--+ +-----------+
| | +-- Rectidentini
| |
| | +-------- Pseudodontini
+--+ +--+
| | | +----- Gonideini
| | +--+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Lamprotulini
|
| +-- Amblemini **
| +--+
| | +-- Lampsilini **
+-----+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
0 |
4 |
116 |
116 |
3 |
20.0 |
+----------------- Parreysiinae
|
| +-------------- Pseudodontini
--+ |
| | +-- Rectidentini
| | +--------+
+--+ | +-- Contradentini
| |
| | +-------- Unioninae
| | |
| | | +-- Amblemini **
+--+ | |
| | +-- Ortmanniana **
| | |
| | +-- Lampsilis **
| | |
| | +-- Cambarunio **
| | |
+--+ +--+-- Elliptio **
| | |
| | +-- Eurynia **
| | |
| | +-- Pleurobema **
| | |
| | +-- Quadrula_quadrula **
| | |
| | +-- Quadrula_verrucosa **
| |
| | +-- Chamberlainia
+--+ |
| +-- Sinohyriopsis
| |
| +-- Lamprotula_caveata
| |
| +-- Lamprotula_leaii
| |
| +-- Potomida
| |
| +-- Pronodularia
| |
+--+-- Gonidea
|
+-- Leguminaia_wheatleyi
|
+-- Leguminaia
|
+-- Microcondylaea
|
+-- Sinosolenaia_carinata
|
+-- Sinosolenaia_oleivora
|
+-- Sinosolenaia_recognita
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
4 |
67 |
57 |
2 |
19.0 |
+-------------------- Pseudodontini
|
--+ +-- Rectidentini
| +--------------+
| | +-- Contradentini
+--+
| +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Unioninae
| |
+--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Pleurobemini **
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
9 |
2 |
2 |
1 |
1 |
19.0 |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
13 |
0 |
4 |
76 |
76 |
3 |
24.0 |
+-------------- Parreysiinae
|
| +----------- Pseudodontini
--+ |
| | +-- Rectidentini
| | +-----+
+--+ | +-- Contradentini
| |
| | +----- Unioninae
| | |
+--+ | +-- Lamprotula
| | |
| | +-- Potomida
| | |
| | +-- Pronodularia
+--+ |
| +-- Gonidea
| |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
+--+-- Amblemini **
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
+-- Cambarunio **
|
+-- Elliptio **
|
+-- Pleurobema **
|
+-- Quadrulini **
Serb & Barnhart, 2008 [Publ 2674], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
10 |
1 |
0 |
0 |
0 |
20.0 |
-- (Lampsilini
Serb, 2006 [Publ 2678], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
7 |
1 |
0 |
0 |
0 |
20.0 |
-- (Lampsilini
Buhay et al., 2002 [Publ 2661], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
3 |
1 |
0 |
0 |
0 |
22.0 |
-- (Lampsilini
Jones & Neves, 2010 [Publ 2646], fig. 16
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
23.0 |
-- (Epioblasma
Serb, 2006 [Publ 2678], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
8 |
1 |
0 |
0 |
0 |
20.0 |
-- (Lampsilini
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
0 |
4 |
29 |
25 |
2 |
18.0 |
+-- Amblemini **
|
+-- Ortmanniana **
|
+-- Lampsilis **
|
--+-- Cambarunio **
|
+-- Elliptio **
|
+-- Pleurobema **
|
+-- Quadrulini **
Serb & Barnhart, 2008 [Publ 2674], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
9 |
1 |
0 |
0 |
0 |
20.0 |
-- (Lampsilini
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
32 |
30 |
3 |
10.0 |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
16 |
14 |
2 |
27 |
26 |
2 |
27.0 |
+----- Prolasmidonta
+--+
| | +-- Alasmidonta
| +--+
+-----------------+ +-- Anemina
| |
| | +----- Pseudodontini
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Schistodesmus
| +-----+
| | | +-- Lepidodesmini
--+ | +--+
| | +-- Aculamprotula
| +-----------+
| | | +----- Platynaias
| | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Pyganodon
| | |
| | | +----- Sinanodonta
| | +--+
| | | +-- Cristaria
+--+ +--+
| +-- Anodontina
|
| +-- Utterbackia_peninsularis
| +-----+
| | +-- Potamilus_alatus **
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis
+--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosulinae
+--+
| +-------------- Quadrulini **
+--+
| +----------- Toxolasma **
+--+
| +-------- Lampsilis_ornata **
+--+
| +----- Potamilus_leptodon **
+--+
| +-- Lampsilis_siliquoidea **
+--+
+-- Lampsilis_powellii **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
29 |
27 |
3 |
10.0 |
+-- Quadrulini **
+-----+
| +-- Lampsilini **
--+
| +----- Gonideinae
+--+
| +-- Parreysiinae
+--+
+-- Unioninae
Jones & Neves, 2010 [Publ 2646], fig. 17
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
1 |
1 |
0 |
0 |
0 |
23.0 |
-- (Epioblasma
Buhay et al., 2002 [Publ 2661], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
22.0 |
-- (Lampsilini
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
14 |
13 |
2 |
22.0 |
-- (Quadrulina
Serb, 2006 [Publ 2678], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
8 |
1 |
0 |
0 |
0 |
20.0 |
-- (Lampsilini
Wu et al., 2021 [Publ 4366], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
85 |
68 |
3 |
17.0 |
+-------------------- Parreysiinae
|
--+ +----------------- Unioninae
| |
| | +-- Contradentini
+--+ +-----------+
| | +-- Rectidentini
| |
+--+ +----- Lampsilini **
| +-----+
| | | +-- Quadrulini **
| | +--+
+--+ +-- Pleurobemini **
|
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2023 [Publ 4339], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
13 |
11 |
1 |
17.0 |
+-------- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
11.0 |
+-- Quadrulini **
--+
+-- Lampsilini **
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
37 |
27 |
2 |
11.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
43 |
40 |
2 |
11.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
8 |
4 |
2 |
20.0 |
+-------- Gonideinae
--+
| +----- Unioninae
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
10 |
8 |
2 |
11.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
9 |
6 |
2 |
20.0 |
+-------------- Pseudodontini
--+
| +----------- Gonidea
+--+
| +-------- Inversidens
+--+
| +----- Unioninae
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Kandl et al., 2001 [Publ 3384], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
0 |
0 |
0 |
22.5 |
+-- Elliptio **
|
--+-- Quadrulini **
|
+-- Pleurobema **
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
26 |
26 |
2 |
11.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Keogh & Simons, 2019 [Publ 3935], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
6 |
1 |
0 |
0 |
0 |
21.0 |
-- (Lampsilis
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
8 |
7 |
3 |
10 |
7 |
2 |
19.0 |
+-------- Unioninae
|
--+ +-- Pleurobemini **
| +--+
| | +-- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
26 |
26 |
2 |
11.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
44 |
36 |
2 |
11.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
7 |
5 |
2 |
9 |
6 |
2 |
21.0 |
+-------------- Unioninae
|
--+ +----------- Lamprotulini
| |
+--+ +-- Gonideini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Margaritiferinae
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
39 |
36 |
2 |
11.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
2 |
30 |
4 |
2 |
20.0 |
+-------------- Gonideinae
|
--+ +----------- Cyrtonaias **
| |
+--+----------- Pleurobemini **
|
| +-------- Potamilus **
+--+
| +----- Lampsilis **
+--+
| +-- Ligumia **
+--+
+-- Ortmanniana **
Wu et al., 2021 [Publ 4366], fig. 2_I
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
85 |
68 |
3 |
15.0 |
+-------------------- Parreysiinae
|
--+ +----------------- Unioninae
| |
| | +-- Rectidentini
+--+ +-----------+
| | +-- Contradentini
| |
+--+ +----- Quadrulini **
| +-----+
| | | +-- Pleurobemini **
| | +--+
+--+ +-- Lampsilini **
|
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
3 |
2 |
1 |
20.0 |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
9 |
6 |
2 |
20.0 |
+-------------- Pseudodontini
--+
| +----------- Gonidea
+--+
| +-------- Inversidens
+--+
| +----- Unioninae
+--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
41 |
35 |
2 |
11.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
39 |
36 |
2 |
11.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
3 |
2 |
1 |
20.0 |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Krebs et al., 2013 [Publ 3515], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
21.0 |
-- (Lampsilini
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
3 |
13 |
12 |
2 |
16.0 |
+-- Quadrulini **
|
--+-- Pleurobemini **
|
+-- Lampsilini **
Manendo et al., 2008 [Publ 2669], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
8 |
8 |
1 |
19.0 |
+-------- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Amblemini **
+--+
+-- Pleurobemini **
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
34 |
31 |
2 |
12.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
4 |
29 |
26 |
3 |
15.0 |
+-------- Quadrulini **
--+
| +----- Pleurobemini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Buhay et al., 2002 [Publ 2661], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
21.0 |
-- (Lampsilini
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
35 |
25 |
3 |
14.0 |
+-------------- Parreysiinae
--+
| +----------- Unioninae
+--+
| +-------- Gonideinae
+--+
| +----- Quadrulini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Whelan et al., 2011 [Publ 3380], fig. 1-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
4 |
31 |
19 |
3 |
17.0 |
+----------------------- Parreysiinae
|
--+ +-- Pseudodontini
| +-----------------+
| | +-- Contradentini
+--+
| +----------------- Unioninae
+--+
| +-------------- Gonideini
+--+
| +----------- Lamprotulini
+--+
| +-------- Pleurobemini **
+--+
| +----- Quadrulini **
+--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Whelan et al., 2011 [Publ 3380], fig. 1-MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
4 |
31 |
19 |
3 |
16.0 |
+----------------------- Parreysiinae
|
--+ +-------------------- Contradentini
| |
+--+ +----------------- Pseudodontini
| |
+--+ +-------------- Unioninae
| |
+--+ +----------- Gonideini
| |
+--+ +-------- Lamprotulini
| |
+--+ +----- Quadrulini **
| |
+--+----- Pleurobemini **
|
| +-- Amblemini **
+--+
+-- Lampsilini **
Jones et al., 2006 [Publ 2681], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
4 |
1 |
0 |
0 |
0 |
21.0 |
-- (Epioblasma
Lane et al., 2016 [Publ 3735], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
1 |
1 |
0 |
0 |
0 |
20.0 |
-- (Lampsilini
Saxon et al., 2023 [Publ 4338], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
21.0 |
-- (Lampsilini
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
6 |
6 |
2 |
16.0 |
+-- Gonideinae
+-----+
| +-- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Quadrulini **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
28 |
26 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Patterson et al., 2021 [Publ 4222], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
0 |
0 |
0 |
18.0 |
+-- Lampsilini **
--+
+-- Amblemini **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
29 |
26 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Pfeiffer et al., 2018 [Publ 3835], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
18.0 |
-- (Quadrulini
Dai et al., 2024 [Publ 4386], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
36 |
36 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
21 |
15 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Johnson et al., 2023 [Publ 4323], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
20.0 |
-- (Pleurobemini
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
4 |
116 |
105 |
3 |
15.0 |
+----------------- Parreysiinae
|
| +-------------- Unioninae
| |
--+ | +-- Lens
| | |
| | +-- Ensidens
| | |
+--+ +--------+-- Physunio
| | |
| | +-- Rectidens
| | |
| | +-- Yaukthwa
| |
+--+ +-- Amblemini **
| |
| +-- Pleurobemini **
| +-----+
| | +-- Lampsilini **
| | |
| | +-- Quadrulini **
+--+
| +-- Chamberlainiini
| +--+
| | +-- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
16 |
15 |
2 |
13.0 |
+----- Gonideinae
--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
12 |
11 |
2 |
13.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
19 |
16 |
4 |
16 |
15 |
2 |
31.0 |
+-------- Amblemini **
|
+-------- Pleurobemini **
--+
+-------- Lampsilini **
|
| +----- Quadrulini **
+--+
| +-- Gonideinae
+--+
+-- Unioninae
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
21 |
15 |
2 |
13.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
34 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
14 |
8 |
2 |
18.0 |
+----------------- Pseudodontini
--+
| +-------------- Contradentini
+--+
| +----------- Unioninae
+--+
| +-------- Gonideini
+--+
| +----- Lampsilini **
+--+
| +-- Pleurobemini **
+--+
+-- Amblemini **
Gangloff et al., 2013 [Publ 3551], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
19.0 |
-- (Lampsilini
Zanatta & Harris, 2013 [Publ 3530], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
1 |
1 |
1 |
17.0 |
+-- Unioninae
--+
+-- Lampsilini **
Mulvey et al., 1997 [Publ 3390], fig. 1_3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
1 |
1 |
1 |
18.0 |
+----- Unioninae
--+
| +-- Quadrulini **
+--+
+-- Amblemini **
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
13.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
20 |
17 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
9 |
6 |
2 |
14.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
12 |
10 |
2 |
14.0 |
+-- Anodontina
+--------+
| +-- Utterbackia
|
--+ +----- Gonideinae
| +--+
| | | +-- Quadrulini **
| | +--+
+--+ +-- Lampsilini **
|
| +----- Unionini
+--+
| +-- Pyganodon
+--+
+-- Cristariina
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
8 |
6 |
2 |
14.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
1 |
1 |
1 |
18.0 |
+----- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Quadrulini **
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
2 |
9 |
6 |
2 |
16.0 |
+-------- Unioninae
--+
| +----- Lampsilini **
+--+
| +-- Quadrulini **
+--+
+-- Gonideinae
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
19 |
19 |
2 |
14.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Chen et al., 2012 [Publ 3624], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
4 |
4 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
14 |
14 |
2 |
14.0 |
+----------- Anodontini
|
--+ +-- Lanceolariini
| +-----+
| | +-- Unionini
+--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
19 |
14 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Wen et al., 2017 [Publ 3795], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
3 |
3 |
1 |
14.0 |
+----- Unioninae
--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
2 |
1 |
3 |
3 |
1 |
19.0 |
+-- Unioninae
--+
+-- Lampsilini **
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
19 |
14 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
20 |
19 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
17 |
15 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
7 |
5 |
2 |
14.0 |
+----- Gonideinae
--+
| +-- Lampsilini **
+--+
+-- Unioninae
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
14 |
12 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Smith & Johnson, 2020 [Publ 4229], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
19.0 |
-- (Potamilus
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
2 |
1 |
1 |
1 |
18.0 |
+----- Quadrulini **
--+
| +-- Unioninae
+--+
+-- Lampsilini **
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
18 |
18 |
2 |
14.0 |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Krebs, 2004 [Publ 2692], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
5 |
3 |
3 |
2 |
2 |
1 |
17.0 |
+-------- Quadrulini **
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Pleurobemini **
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
26 |
25 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
8 |
6 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
16 |
15 |
2 |
14.0 |
+-- Quadrulini **
+--+
| +-- Lampsilini **
--+
| +-- Unioninae
+--+
+-- Gonideinae
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
1 |
0 |
0 |
17.0 |
+-- Lampsilini **
--+
+-- Quadrulini **
Sheng et al., 2014 [Publ 3704], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
6 |
4 |
2 |
5 |
5 |
2 |
19.0 |
+-------------- Cristariina
--+
| +----------- Alasmidontina
+--+
| +-------- Lampsilis **
+--+
| +----- Gonideinae
+--+
| +-- Venustaconcha **
+--+
+-- Quadrulini **
Zieritz et al., 2024 [Publ 4381], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
44 |
42 |
3 |
-- |
+-- Amblemini **
+--------------+
| +-- Lampsilini **
|
| +-------- Parreysiinae
| +-----+
--+ | | +----- Unioninae
| | +--+
| | | +-- Chamberlainia
| | +--+
| | +-- Discomya
+--+
| +----- Ctenodesmini
| +-----+
| | | +-- Rectidentini
| | +--+
| | +-- Contradentini
+--+
| +-- Schepmaniini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Caudiculatus
+--+
| +-- Potomida
+--+
+-- Gonideini
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
11 |
2 |
-- |
+-- Amblemini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
7 |
0 |
2 |
14 |
8 |
3 |
20.0 |
+-------------------------- Lamellidens_mainwaringi
--+
| +----------------------- Lamprotula
+--+
| +-------------------- Potomida
+--+
| +----------------- Lampsilini **
+--+
| +-------------- Pleurobemini **
+--+
| +----------- Aculamprotula
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodontini
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus
Metzger et al., 2018 [Publ 3952], fig. 1C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
1 |
1 |
1 |
-- |
+----- Unioninae
--+
| +-- Pleurobemini **
+--+
+-- Lampsilini **
Giribet & Distel, 2003 [Publ 3670], fig. 3f5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
4 |
1 |
1 |
16.0 |
+-- Pleurobemini **
|
--+-- Lampsilini **
|
+-- Unioninae
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
9 |
0 |
2 |
13 |
7 |
3 |
22.0 |
+-- Aculamprotula
|
+-- Lamprotula
|
+-- Lampsilini **
|
+-- Potomida
|
--+-- Unio_pictorum
|
+-- Anodontini
|
+-- Unio_ravoisieri
|
+-- Parreysiinae
|
+-- Pleurobemini **
Xue et al., 2016 [Publ 3659], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
0 |
0 |
0 |
-- |
+-- Quadrulini **
--+
+-- Lampsilini **
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
11 |
3 |
-- |
+-------- Parreysiinae
|
--+ +-- Amblemini **
| +--+
| | +-- Lampsilini **
+--+
| +-- Unioninae
+--+
+-- Gonideinae
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
6 |
0 |
2 |
5 |
4 |
2 |
19.0 |
+-------- Quadrulini **
--+
| +----- Lampsilini **
+--+
| +-- Unioninae
+--+
+-- Gonideinae
Xue et al., 2016 [Publ 3659], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
14 |
14 |
2 |
-- |
+----- Unioninae
--+
| +-- Gonideinae
+--+
+-- Lampsilini **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
54 |
53 |
3 |
-- |
+----------- Parreysiinae
--+
| +-------- Unioninae
+--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Amblemini **
Wei et al., 2016 [Publ 3665], fig. 0
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
4 |
0 |
2 |
4 |
4 |
2 |
17.0 |
+----- Gonideinae
|
--+ +-- Cristariina
| |
| +-- Lampsilini **
+--+
+-- Alasmidontina
|
+-- Quadrulini **
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
6 |
0 |
2 |
5 |
4 |
2 |
19.0 |
+-------- Quadrulini **
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Unioninae
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+----- Gonideinae
|
--+----- Unioninae
|
| +-- Lampsilini **
+--+
+-- Quadrulini **
Krebs, 2004 [Publ 2692], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
2 |
2 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Pleurobemini **
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
30 |
28 |
1 |
-- |
+----- Gonideinae
--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
6 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
6 |
0 |
2 |
13 |
7 |
3 |
19.0 |
+-- Pleurobemini **
|
--+-- Lampsilini **
|
+-- Unioninae
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
10 |
2 |
-- |
+-- Amblemini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Burzynski et al., 2017 [Publ 3774], fig. 1-top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
11 |
11 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
4 |
2 |
-- |
+-------- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Quadrulini **
+--+
+-- Lampsilini **
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+----- Gonideinae
|
--+----- Unioninae
|
| +-- Lampsilini **
+--+
+-- Quadrulini **
Froufe et al., 2015 [Publ 3664], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
10 |
2 |
-- |
+-- Gonideini
+--------+
| +-- Lamprotulini
--+
| +-------- Chamberlainiini
+--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
7 |
2 |
-- |
+-- Lampsilini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
38 |
36 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
38 |
36 |
3 |
-- |
+-- Parreysiinae
+-----+
| +-- Unioninae
--+
| +----- Gonideinae
+--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
7 |
7 |
2 |
-- |
+-- Lampsilini **
+--+
| +-- Quadrulini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
3 |
0 |
2 |
15 |
4 |
2 |
17.0 |
+-------- Unionini
--+
| +----- Pleurobemini **
+--+
| +-- Anodontini
+--+
+-- Lampsilini **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
7 |
6 |
1 |
-- |
+-- Pleurobemini **
--+
+-- Unioninae
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
9 |
2 |
1 |
-- |
+----- Unioninae
--+
| +-- Amblemini **
+--+
+-- Lampsilini **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
1 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Quadrulini **
Campbell, 2000 [Publ 2724], fig. 1-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Unioninae
--+
+-- Pleurobemini **
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
-- Amblema_plicata
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Unioninae
|
--+-- Lampsilini **
|
+-- Gonideinae
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
Hoeh et al., 1996 [Publ 2694], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
1 |
-- |
+-- None__1
+--+
| +-- None__2
--+
| +-- Pleurobemini **
+--+
+-- Unioninae
Matos et al., 2024 [Publ 4384], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
18 |
1 |
-- |
+-- Amblemini **
--+
+-- Gonideinae
Wu et al., 2020 [Publ 4147], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
10 |
2 |
-- |
+----- Lampsilini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Gonideinae
--+
| +-- Lampsilini **
+--+
+-- Unioninae
Soroka, 2008 [Publ 3695], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-- Pleurobemini **
--+
+-- Unioninae
Huff et al., 2004 [Publ 2697], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
Takeuchi et al., 2015 [Publ 3661], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Lampsilis_cardium
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
2 |
1 |
-- |
+-- Unionini
|
--+-- Anodontini
|
+-- Lampsilini **
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
-- Amblema_plicata
Vikhrev et al., 2018 [Publ 3812], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Unioninae
--+
+-- Lampsilini **
Soroka, 2008 [Publ 3695], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+-- Pleurobemini **
--+
+-- Unioninae
Huff et al., 2004 [Publ 2697], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
2 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
Plazzi & Passamonti, 2010 [Publ 3354], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
1 |
1 |
1 |
-- |
+-- Gonideinae
--+
+-- Lampsilini **
Jiang et al., 2010 [Publ 3438], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Lampsilini **
|
--+-- Unioninae
|
+-- Gonideinae
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Gonideinae
--+
| +-- Lampsilini **
+--+
+-- Unioninae
Marshall et al., 2014 [Publ 3593], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
2 |
1 |
-- |
-- Amblema_plicata
Khalloufi et al., 2011 [Publ 2748], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Baker et al., 2004 [Publ 2685], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Epioblasma_triquetra
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unioninae
--+
| +-- Lampsilini **
+--+
+-- Parreysiinae
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
15 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Takeuchi et al., 2015 [Publ 3661], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Lampsilis_cardium
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unioninae
--+
| +-- Parreysiinae
+--+
+-- Lampsilini **
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
45 |
3 |
-- |
+-------------- Parreysiinae
--+
| +----------- Unioninae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lampsilini **
+--+
| +-- Lamprotulini
+--+
+-- Gonideini
Araujo et al., 2009 [Publ 2589], fig. 7A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
9 |
2 |
-- |
+----- Amblemini **
--+
| +-- Gonideinae
+--+
+-- Unioninae
Araujo et al., 2009 [Publ 2589], fig. 7C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
8 |
2 |
-- |
+----- Gonideinae
--+
| +-- Amblemini **
+--+
+-- Unioninae
Lemer et al., 2019 [Publ 4093], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Lampsilis_cardium
Cyr et al., 2007 [Publ 2745], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
4 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
Matos et al., 2024 [Publ 4383], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
18 |
1 |
-- |
+-- Amblemini **
--+
+-- Gonideinae
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilini **
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unioninae
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
10 |
3 |
-- |
+-- Lampsilini **
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unioninae
Huang et al., 2018 [Publ 3828], fig. 3_5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
5 |
3 |
-- |
+-------- Gonideinae
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Parreysiinae
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
45 |
3 |
-- |
+-------- Parreysiinae
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Gonideinae
Araujo et al., 2009 [Publ 2589], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
9 |
2 |
-- |
+----- Gonideinae
--+
| +-- Amblemini **
+--+
+-- Unioninae
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
3 |
1 |
-- |
+-- Lampsilini **
--+
+-- Unioninae
Baker et al., 2003 [Publ 2690], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Epioblasma_triquetra
Inoue et al., 2014 [Publ 3635], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
7 |
1 |
-- |
+-- Amblemini **
--+
+-- Unioninae
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
45 |
3 |
-- |
+-------- Parreysiinae
--+
| +----- Unioninae
+--+
| +-- Lampsilini **
+--+
+-- Gonideinae
| 
